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<document id="1AA711B057F4C35154EFF8AC8E640640" ID-CLB-Dataset="6390" ID-DOI="10.5252/g2014n4a4" ID-GBIF-Dataset="8bc26961-9853-4113-a63d-4cf691aa7100" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538625" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150850454" checkinUser="felipe" docAuthor="Vianey-Liaud, Monique, Comte, Bernard, Marandat, Bernard, Peigné, Stéphane, Rage, Jean-Claude &amp; Sudre, Jean" docDate="2014" docId="03D6987B4477B12BFEBF15E3FDFEE720" docLanguage="en" docName="Geodiversitas.36.4.565-622.pdf" docOrigin="Geodiversitas 36 (4)" docSource="http://dx.doi.org/10.5252/g2014n4a4" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Krokolithes Hirsch 1985" docType="treatment" docVersion="7" lastPageNumber="572" masterDocId="FFEFE0034471B12CFF8F106FFF81E50F" masterDocTitle="A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)" masterLastPageNumber="622" masterPageNumber="565" pageNumber="571" updateTime="1698929460103" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="67971D1237C4CD6C11A8FBE238897E75">A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)</mods:title>
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<mods:namePart id="E3CFBC7BE3B1D1C6D388531A3FE804DF">Vianey-Liaud, Monique</mods:namePart>
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<mods:namePart id="7CBD8D31868F539F3AAB71A4F3F69B9F">Comte, Bernard</mods:namePart>
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<mods:namePart id="D54DF872E9414B37E553BD5D6A7D2BF7">Marandat, Bernard</mods:namePart>
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<mods:namePart id="552AC597721CC1EDEFE7486594366621">Peigné, Stéphane</mods:namePart>
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<mods:namePart id="AA5F2FF6246DFF2A4E46D57B8A295DA8">Sudre, Jean</mods:namePart>
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<subSubSection id="C3657AE64477B12AFEBF15E3FE49E0A9" box="[304,456,1420,1446]" pageId="6" pageNumber="571" type="nomenclature">
<paragraph id="8BC0296D4477B12AFEBF15E3FE49E0A9" blockId="6.[304,456,1420,1446]" box="[304,456,1420,1446]" pageId="6" pageNumber="571">
<heading id="D0889E014477B12AFEBF15E3FE49E0A9" box="[304,456,1420,1446]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="571" reason="2">
<taxonomicName id="4C7F52EE4477B12AFEBF15E3FE22E0A9" authorityName="Hirsch" authorityYear="1985" box="[304,419,1420,1446]" class="Reptilia" family="Krokolithidae" genus="Krokolithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="571" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4477B12AFEBF15E3FE22E0A9" box="[304,419,1420,1446]" italics="true" pageId="6" pageNumber="571">Krokolithes</emphasis>
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sp.
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<paragraph id="8BC0296D4477B12AFFFE15A8FEEBE31C" blockId="6.[113,647,1478,1555]" pageId="6" pageNumber="571">
MATERIAL EXAMINED. — Seven small fragments (from
<quantity id="4C8784884477B12AFFFE158EFF5AE0F7" box="[113,219,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="571" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 mm</quantity>
2); thickness
<emphasis id="B90BF57F4477B12AFED6158EFEE6E0F7" box="[345,359,1505,1528]" italics="true" pageId="6" pageNumber="571">=</emphasis>
0.33, 0.35, 0.38, 0.39, 0.40, 0.42,
<quantity id="4C8784884477B12AFF251594FE86E31C" box="[170,263,1531,1555]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="571" unit="mm" value="0.45">0.45 mm</quantity>
;
<collectionCode id="ED6EB1A84477B12AFE9D1594FEC1E31C" box="[274,320,1531,1555]" pageId="6" pageNumber="571">SPV</collectionCode>
750.
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DESCRIPTION (
<figureCitation id="134435E84477B12AFE9C1624FEE2E36A" box="[275,355,1611,1637]" captionStart="FIG" captionStartId="14.[113,124,1459,1477]" captionTargetBox="[116,1197,215,1426]" captionTargetId="figure-140@14.[478,661,773,867]" captionTargetPageId="14" captionText="FIG. 3. — Saint-Privat-des-Vieux (Gard, Lower Chattian, MP 26): A, SEM view of a fragment of Krokolithes sp. SPV750; B-H teeth of mammals: B, cf. Geotrypus sp. SPV800, left m1, occlusal view; C-I, Cuvierimops sp.: C, SPV810, left p4-m1, labial view; D, SPV814, left m3: D1, occlusal view, D2, distal view; E, SPV812, left m2: E1, occlusal view, drawing, E2, occlusal view SEM, E3, distal view; F, SPV815, right upper canine: F1, occlusal view, F2, lingual view, F3, labial view; G, SPV820, right M3, occlusal view; H, SPV816, left M1: H1, occlusal view, H2, lingual view; I, SPV817, right M2: I1, occlusal view, I2, lingual view. Scale bars:A, 167 µm; B-I2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4538633" httpUri="https://zenodo.org/record/4538633/files/figure.png" pageId="6" pageNumber="571">Fig. 3A</figureCitation>
)
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<paragraph id="8BC0296D4477B12AFFFE1605FDF5E3AC" blockId="6.[113,646,1610,1700]" pageId="6" pageNumber="571">
Two fragments of eggshells from this locality are figured in
<bibRefCitation id="EFEE549C4477B12AFF6C16E5FE39E3AC" author="BONNET A. &amp; COMTE B. &amp; LARDET H. &amp; MARANDAT B. &amp; REMY J. A. &amp; SUDRE J. &amp; LIAUD M." box="[227,440,1673,1700]" pageId="6" pageNumber="571" pagination="9 - 18" refId="ref31791" refString="BONNET A., COMTE B., LARDET H., MARANDAT B., REMY J. A., SUDRE J. &amp; VIANEY- LIAUD M. 2011. - Le gisement a mammiferes de Saint-Privat-des-Vieux (Gard), un nouveau repere chronologique dans la sedimentation du fosse oligocene d'Ales. Bulletin de la Societe d'Etudes en Sciences naturelles 68: 9 - 18." type="journal article" year="2011">
Bonnet
<emphasis id="B90BF57F4477B12AFEB616E5FEECE3AC" box="[313,365,1673,1699]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
(2011
</bibRefCitation>
: pl. II, figs B, C).
</paragraph>
<paragraph id="8BC0296D4477B12AFD341481FBF0E3AC" blockId="6.[674,1212,1261,1699]" pageId="6" pageNumber="571">
Ŋe size of fragments does not allow to infer the shape of the egg.Like in
<taxonomicName id="4C7F52EE4477B12AFCD91562FB12E027" authority="Hirsch, 1985" authorityName="Hirsch" authorityYear="1985" box="[854,1171,1293,1320]" class="Reptilia" family="Krokolithidae" genus="Krokolithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="wilsoni">
<emphasis id="B90BF57F4477B12AFCD91562FB8BE028" box="[854,1034,1293,1319]" italics="true" pageId="6" pageNumber="571">Krokolithes wilsoni</emphasis>
<bibRefCitation id="EFEE549C4477B12AFB811562FB12E027" author="HIRSCH K. F." box="[1038,1171,1293,1320]" pageId="6" pageNumber="571" pagination="531 - 542" refId="ref33052" refString="HIRSCH K. F. 1985. - Fossil crocodylian eggs from the Eocene of Colorado. Journal of Palaeontology 59: 531 - 542." type="journal article" year="1985">Hirsch, 1985</bibRefCitation>
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the outer surface is rough, and shows erosion craters and circular pore openings(diameter = 0.09 and
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) with stepped erosion rings;therefore there are low blunt anastomosed and irregular ridges.Basal knobs are well distinct on two fragments and worn on the others. Ŋus, the radiating wedges are visible. Ŋe units are relatively wide (as wide as high, or wider than high) and display the typical wedge pattern of crocodylian eggshells.Ŋere are deep insterstices between the units. Ŋe fragments are thinner than in the Eocene
<taxonomicName id="4C7F52EE4477B12AFBFA1624FCD6E38B" authority="Hirsch, 1985" authorityName="Hirsch" authorityYear="1985" class="Reptilia" family="Krokolithidae" genus="Krokolithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="wilsoni">
<emphasis id="B90BF57F4477B12AFBFA1624FD4AE38C" italics="true" pageId="6" pageNumber="571">K. wilsoni</emphasis>
<bibRefCitation id="EFEE549C4477B12AFD5E1606FCD6E38B" author="HIRSCH K. F." box="[721,855,1641,1668]" pageId="6" pageNumber="571" pagination="531 - 542" refId="ref33052" refString="HIRSCH K. F. 1985. - Fossil crocodylian eggs from the Eocene of Colorado. Journal of Palaeontology 59: 531 - 542." type="journal article" year="1985">Hirsch, 1985</bibRefCitation>
</taxonomicName>
and much more than the Miocene crocodylian indet. (
<bibRefCitation id="EFEE549C4477B12AFCE716E6FBE4E3AC" author="HIRSCH K. F. &amp; KOHRING R." box="[872,1125,1673,1699]" pageId="6" pageNumber="571" pagination="59 - 65" refId="ref33078" refString="HIRSCH K. F. &amp; KOHRING R. 1992. - Crocodilian eggs from the Middle Eocene Bridger Formation, Wyoming. Journal of Vertebrate Paleontology 12: 59 - 65." type="journal article" year="1992">Hirsch &amp; Kohring 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64476B12BFFFE10B7FDFEE720" pageId="7" pageNumber="572" type="discussion">
<paragraph id="8BC0296D4476B12BFFFE10B7FF72E5FF" blockId="7.[111,649,216,559]" box="[113,243,216,242]" pageId="7" pageNumber="572">DISCUSSION</paragraph>
<paragraph id="8BC0296D4476B12BFFFE1097FDFEE720" blockId="7.[111,649,216,559]" pageId="7" pageNumber="572">
Ŋe small size of our fragments does not allow to make a precise identification. Ŋe general ultrastructure coarse wedges and tabular crystalline structure , the figures of erosion, the arrangement of units and their width allow an assignment to
<taxonomicName id="4C7F52EE4476B12BFE521119FDCCE49F" authorityName="Hirsch" authorityYear="1985" box="[477,589,374,400]" class="Reptilia" family="Krokolithidae" genus="Krokolithes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4476B12BFE521119FDCCE49F" box="[477,589,374,400]" italics="true" pageId="7" pageNumber="572">Krokolithes</emphasis>
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. Ŋis ootaxon was formerly recovered from the Eocene of North America (
<bibRefCitation id="EFEE549C4476B12BFE9311DAFE23E4DF" author="HIRSCH K. F." box="[284,418,437,464]" pageId="7" pageNumber="572" pagination="531 - 542" refId="ref33052" refString="HIRSCH K. F. 1985. - Fossil crocodylian eggs from the Eocene of Colorado. Journal of Palaeontology 59: 531 - 542." type="journal article" year="1985">Hirsch 1985</bibRefCitation>
) and from the Eocene and Miocene of
<collectingCountry id="F36869FD4476B12BFE9C11BAFEF7E4E0" box="[275,374,469,495]" name="Germany" pageId="7" pageNumber="572">Germany</collectingCountry>
.
<bibRefCitation id="EFEE549C4476B12BFEF211BAFD06E4FF" author="KOHRING R. &amp; HIRSCH K. F." box="[381,647,469,496]" pageId="7" pageNumber="572" pagination="67 - 80" refId="ref33522" refString="KOHRING R. &amp; HIRSCH K. F. 1996. - Crocodilian and avian eggshells from the middle Eocene of the Geiseltal, Eastern Germany. Journal of Vertebrate Paleontology 16: 67 - 80." type="journal article" year="1996">Kohring &amp; Hirsch (1996)</bibRefCitation>
suggested that at least part of the material assigned to
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<emphasis id="B90BF57F4476B12BFF01127AFF7FE720" box="[142,254,533,559]" italics="true" pageId="7" pageNumber="572">Krokolithes</emphasis>
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may represent eggs of
<taxonomicName id="4C7F52EE4476B12BFE62127AFDF8E720" authorityName="Pomel" authorityYear="1847" box="[493,633,533,559]" class="Mammalia" family="Docodontidae" genus="Diplocynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Docodonta" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4476B12BFE62127AFDF8E720" box="[493,633,533,559]" italics="true" pageId="7" pageNumber="572">Diplocynodon</emphasis>
</taxonomicName>
.
</paragraph>
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