164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document id="AA3012647A1A9597FF24E50D7B1B38D3" ID-CLB-Dataset="4732" ID-DOI="10.1080/00222930110052463" ID-GBIF-Dataset="49960699-495a-4910-bdf5-ce616eea71bc" ID-ISSN="1464-5262" ID-Zenodo-Dep="5298706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629994576800" checkinUser="felipe" docAuthor="Cook, P. L. & Bock, P. E." docDate="2002" docId="03D1686A5A4FA056E3FBD0F2FE2890DF" docLanguage="en" docName="JNatHist.36.13.1601-1619.pdf" docOrigin="Journal of Natural History 36 (13)" docSource="http://www.tandfonline.com/doi/abs/10.1080/00222930110052463" docStyle="DocumentStyle:C540815C53297EEE0623138269730482.2:JNatHist.2000-2002.journal_article.1cover" docStyleId="C540815C53297EEE0623138269730482" docStyleName="JNatHist.2000-2002.journal_article.1cover" docStyleVersion="2" docTitle="Riscodopa paucipora Cook & Bock 2002, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1608" masterDocId="FFE810125A47A05CE238D271FFD59023" masterDocTitle="Notes on astogeny of some Petraliellidae (Bryozoa) from Australia" masterLastPageNumber="1619" masterPageNumber="1601" pageNumber="1608" updateTime="1699145546688" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A99E3152FBF61C6427040843582E6F06">Notes on astogeny of some Petraliellidae (Bryozoa) from Australia</mods:title>
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<mods:namePart id="046D2A73D1EE3CD284AF12A435265650">Cook, P. L.</mods:namePart>
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<mods:namePart id="8D6924344786181C0D028C2B5C7E4239">Bock, P. E.</mods:namePart>
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<mods:title id="C718C8EAC6839730F6BE9B4E500F7812">Journal of Natural History</mods:title>
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<mods:part id="011A4166839451A360A0F0E16E8CCD5A">
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<mods:date id="C815889ED059D40FCD940B1CF46E5462">2002</mods:date>
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<mods:number id="9BF5963A6872D39E06EA56C33A23C3AE">2002-09-30</mods:number>
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<mods:number id="51F757A8FCF1A72D09CD8D17B08B033D">36</mods:number>
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<mods:number id="D5BA2395C80816EFF2C0B2213A6DD9D7">13</mods:number>
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<mods:url id="C29418D335140C497ABCA706FA211EFB">http://www.tandfonline.com/doi/abs/10.1080/00222930110052463</mods:url>
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<treatment id="03D1686A5A4FA056E3FBD0F2FE2890DF" ID-DOI="http://doi.org/10.5281/zenodo.5305767" ID-GBIF-Taxon="185131624" ID-Zenodo-Dep="5305767" LSID="urn:lsid:plazi:treatment:03D1686A5A4FA056E3FBD0F2FE2890DF" httpUri="http://treatment.plazi.org/id/03D1686A5A4FA056E3FBD0F2FE2890DF" lastPageId="10" lastPageNumber="1608" pageId="8" pageNumber="1608">
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<subSubSection id="C3628AF75A4FA054E3FBD0F2FD629298" pageId="8" pageNumber="1608" type="nomenclature">
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<paragraph id="8BC7D97C5A4FA054E3FBD0F2FCDB92B8" blockId="8.[140,819,643,732]" box="[451,782,643,667]" pageId="8" pageNumber="1608">
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<heading id="D08F6E105A4FA054E3FBD0F2FCDB92B8" box="[451,782,643,667]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="1608" reason="3">
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<taxonomicName id="4C78A2FF5A4FA054E3FBD0F2FD7E92B8" ID-CoL="4T53F" authority="Cook & Bock, 2002" authorityName="Cook & Bock" authorityYear="2002" box="[451,683,643,667]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="1608" phylum="Bryozoa" rank="species" species="paucipora" status="sp. nov.">
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<emphasis id="B90C056E5A4FA054E3FBD0F2FD7E92B8" bold="true" box="[451,683,643,667]" italics="true" pageId="8" pageNumber="1608">Riscodopa paucipora</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23FB8155A4FA054E08CD0F5FCDB92B8" box="[692,782,644,667]" pageId="8" pageNumber="1608" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E023D0D2FD629298" blockId="8.[140,819,643,732]" box="[539,695,675,700]" pageId="8" pageNumber="1608">(gures 8–12)</paragraph>
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</subSubSection>
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<subSubSection id="C3628AF75A4FA056E2B4D0B7FE2890DF" lastPageId="10" lastPageNumber="1610" pageId="8" pageNumber="1608" type="materials_examined">
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<paragraph id="8BC7D97C5A4FA054E2B4D0B7FCE692F8" blockId="8.[140,819,643,732]" box="[140,819,710,732]" pageId="8" pageNumber="1608">
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<materialsCitation id="3B10D3215A4FA054E2B4D0B7FCE692F8" ID-GBIF-Occurrence="3339723304" box="[140,819,710,732]" collectionCode="MOV" location="Browns Creek" municipality="Late Eocene" pageId="8" pageNumber="1608" specimenCode="P128374" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
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<typeStatus id="54C367DE5A4FA054E2B4D0B7FF2092FF" box="[140,245,710,732]" pageId="8" pageNumber="1608" type="holotype">HOLOTYPE</typeStatus>
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.
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<collectionCode id="ED6941B95A4FA054E33BD0B7FEEB92FF" box="[259,318,710,732]" pageId="8" pageNumber="1608">MOV</collectionCode>
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<specimenCode id="DBDE71075A4FA054E37ED0B7FE4E92FF" box="[326,411,710,732]" collectionCode="MOV" pageId="8" pageNumber="1608">P128374</specimenCode>
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,
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<collectingMunicipality id="6BA343065A4FA054E393D0B7FDFE92F8" box="[427,555,710,731]" pageId="8" pageNumber="1608">Late Eocene</collectingMunicipality>
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,
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<location id="8EA78FA75A4FA054E002D0B7FD1E92FF" LSID="urn:lsid:plazi:treatment:03D1686A5A4FA056E3FBD0F2FE2890DF:8EA78FA75A4FA054E002D0B7FD1E92FF" box="[570,715,710,732]" municipality="Late Eocene" name="Browns Creek" pageId="8" pageNumber="1608" stateProvince="Victoria">Browns Creek</location>
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,
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<collectingRegion id="49BC179E5A4FA054E0E2D0B7FCF892FF" box="[730,813,710,732]" country="Australia" name="Victoria" pageId="8" pageNumber="1608">Victoria</collectingRegion>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E2B4D172FED69338" blockId="8.[140,1094,771,859]" box="[140,259,771,795]" pageId="8" pageNumber="1608">
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<emphasis id="B90C056E5A4FA054E2B4D172FED69338" box="[140,259,771,795]" italics="true" pageId="8" pageNumber="1608">Etymology</emphasis>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E28BD152FEA29379" blockId="8.[140,1094,771,859]" pageId="8" pageNumber="1608">
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From
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<emphasis id="B90C056E5A4FA054E338D155FE9C9318" box="[256,329,804,827]" italics="true" pageId="8" pageNumber="1608">paucus</emphasis>
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(Latin)— few, and
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<emphasis id="B90C056E5A4FA054E000D155FDA69318" box="[568,627,804,827]" italics="true" pageId="8" pageNumber="1608">porus</emphasis>
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(Latin)—hole, referring to the lack of frontal pseudopores.
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E2B4D1F5FEDF93B8" blockId="8.[140,1095,900,1273]" box="[140,266,900,923]" pageId="8" pageNumber="1608">
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<emphasis id="B90C056E5A4FA054E2B4D1F5FEDF93B8" box="[140,266,900,923]" italics="true" pageId="8" pageNumber="1608">Description</emphasis>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E28BD1D3FB9394DA" blockId="8.[140,1095,900,1273]" pageId="8" pageNumber="1608">
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<taxonomicName id="4C78A2FF5A4FA054E28BD1D3FEF19399" authorityName="Gordon" authorityYear="1989" box="[179,292,930,954]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="1608" phylum="Bryozoa" rank="genus">
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<emphasis id="B90C056E5A4FA054E28BD1D3FEF19399" box="[179,292,930,954]" italics="true" pageId="8" pageNumber="1608">Riscodopa</emphasis>
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</taxonomicName>
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with non-tatiform ancestrula. Autozooids almost vertical, ori ces occupying the major part of the frontal shield. Frontal shield imperforate except for a single series of marginal or submarginal pores, most of which appear to be frontal septular pores. Primary ori ce large, subterminal, with paired lateral denticles, and a narrow, hammer-shaped lyrula, which together de ne a pair of proximal sinuses. Five to six oral spines, proximal mucro small, with a terminal avicularium with a subtriangular rostrum, orientated proximally and frontally, and a complete bar. Paired disto-lateral avicularia on swollen subrostral chambers, orientated laterally. Basal surface of each zooid separated by a groove from its neighbours, with a curved lamina raised above its proximal end. Each wall with a crescent or small group of three to ve septular pores arranged transversely in the mid-line. Ovicell not found.
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E2B4D753FEFE951A" blockId="8.[140,1095,1314,1401]" box="[140,299,1314,1337]" pageId="8" pageNumber="1608">
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<emphasis id="B90C056E5A4FA054E2B4D753FEFE951A" box="[140,299,1314,1337]" italics="true" pageId="8" pageNumber="1608">Measurements</emphasis>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E28BD730FEAA955B" blockId="8.[140,1095,1314,1401]" pageId="8" pageNumber="1608">
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Length of ancestrula
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<quantity id="4C8074995A4FA054E39BD730FDD4957B" box="[419,513,1345,1369]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="1608" unit="mm" value="0.28">0.28 mm</quantity>
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. Length of zooids
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<quantity id="4C8074995A4FA054E0ECD730FCA4957B" box="[724,881,1345,1369]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.8" metricValueMin="2.3" pageId="8" pageNumber="1608" unit="mm" value="0.305" valueMax="0.38" valueMin="0.23">0.23–0.38 mm</quantity>
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. Width of primary ori ce
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<quantity id="4C8074995A4FA054E2E2D710FEA2955B" box="[218,375,1377,1401]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="8" pageNumber="1608" unit="mm" value="0.18" valueMax="0.19" valueMin="0.17">0.17–0.19 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BC7D97C5A4FA054E2B4D7D0FF3A959A" blockId="8.[140,1095,1441,1687]" box="[140,239,1441,1465]" pageId="8" pageNumber="1608">
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<emphasis id="B90C056E5A4FA054E2B4D7D0FF3A959A" box="[140,239,1441,1465]" italics="true" pageId="8" pageNumber="1608">Remarks</emphasis>
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</paragraph>
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<paragraph id="8BC7D97C5A4FA056E28BD7B1FE2890DF" blockId="8.[140,1095,1441,1687]" lastBlockId="10.[140,1094,133,253]" lastPageId="10" lastPageNumber="1610" pageId="8" pageNumber="1608">
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Only one colony has been found. The ancestrula is sunken; its basal wall is missing. The primary ori ce is so large that the ancestrula almost appears to be tatiform. Although slightly damaged, the ori ce shows one lateral denticle, which resembles a condyle, and a proximal avicularian mucro, but no trace of oral spines. The ancestrula is surrounded by a distal triad of primary zooids, and a proximolateral pair budded later in astogeny, one of which has an ori ce occluded by calci cation, which includes the spine bases. The zooid ori ces are so large, and the frontal wall so restricted, that the network of pseudopores typical of other species of
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<taxonomicName id="4C78A2FF5A4EA055E297D7D0FEF5959A" authorityName="Gordon" authorityYear="1989" box="[175,288,1441,1465]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="genus">
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<emphasis id="B90C056E5A4EA055E297D7D0FEF5959A" box="[175,288,1441,1465]" italics="true" pageId="9" pageNumber="1609">Riscodopa</emphasis>
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</taxonomicName>
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is absent. The marginal series appear to be all septular pores. The paired avicularia are not raised on columnar processes as in
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<emphasis id="B90C056E5A4EA055E10AD7B0FC7895FB" box="[818,941,1472,1496]" italics="true" pageId="9" pageNumber="1609">
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<taxonomicName id="4C78A2FF5A4EA055E10AD7B0FC7095FB" baseAuthorityName="Waters" baseAuthorityYear="1882" box="[818,933,1472,1496]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="species" species="biincisa">R. biincisa</taxonomicName>
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.
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</emphasis>
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The presence of an avicularium on the ancestrula is unusual but has been reported in other species (
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<bibRefCitation id="EFE9A48D5A4EA055E2AFD471FE27963B" author="HAYWARD, P. J. & COOK, P. L." box="[151,498,1536,1560]" pageId="9" pageNumber="1609" pagination="43 - 130" refId="ref8308" refString="HAYWARD, P. J. and COOK, P. L., 1979, The South African Museum's Meiring Naude cruises, Part 9, Bryozoa, Annals of the South African Museum, 79, 43 - 130." type="journal article" year="1979">Hayward and Cook, 1979: 77</bibRefCitation>
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). The basal septular pores diVer from those of
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<taxonomicName id="4C78A2FF5A4EA055E614D470FF0B961B" baseAuthorityName="Waters" baseAuthorityYear="1882" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="species" species="biincisa">
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<emphasis id="B90C056E5A4EA055E614D470FF0B961B" italics="true" pageId="9" pageNumber="1609">R. biincisa</emphasis>
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</taxonomicName>
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in being more numerous and arranged in a crescent across the mid-line of the zooid. The curved proximal lamina is less robust, but similar to that occurring in some specimens of
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<emphasis id="B90C056E5A4EA055E3BCD410FDD4965B" box="[388,513,1632,1656]" italics="true" pageId="9" pageNumber="1609">
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<taxonomicName id="4C78A2FF5A4EA055E3BCD410FE2C965B" baseAuthorityName="Waters" baseAuthorityYear="1882" box="[388,505,1632,1656]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="species" species="biincisa">R. biincisa</taxonomicName>
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.
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</emphasis>
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It suggests that while the rest of the basal calci ed wall is an interior wall, the rhizoids of
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<taxonomicName id="4C78A2FF5A4EA055E06CD4F1FD1E96B4" baseAuthorityName="Waters" baseAuthorityYear="1882" box="[596,715,1663,1687]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="species" species="biincisa">
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<emphasis id="B90C056E5A4EA055E06CD4F1FD1E96B4" box="[596,715,1663,1687]" italics="true" pageId="9" pageNumber="1609">R. biincisa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C78A2FF5A4EA055E132D4F1FC4F96B4" authorityName="Cook & Bock" authorityYear="2002" box="[778,922,1664,1687]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1609" phylum="Bryozoa" rank="species" species="paucipora">
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<emphasis id="B90C056E5A4EA055E132D4F1FC4F96B4" box="[778,922,1664,1687]" italics="true" pageId="9" pageNumber="1609">R. paucipora</emphasis>
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</taxonomicName>
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may have had a swollen origin surrounded by a rim of exterior calci ed wall, which included several septular pores, like those of
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<taxonomicName id="4C78A2FF5A4DA056E015D2D4FBE8909E" authority="(Cook and Chimonides 1981 a)" baseAuthorityName="Cook and Chimonides" baseAuthorityYear="1981" box="[557,1085,165,189]" class="Gymnolaemata" genus="Parastichopora" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1610" phylum="Bryozoa" rank="genus">
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<emphasis id="B90C056E5A4DA056E015D2D4FD01909E" box="[557,724,165,189]" italics="true" pageId="10" pageNumber="1610">Parastichopora</emphasis>
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(
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<bibRefCitation id="EFE9A48D5A4DA056E0DDD2D4FBE0909E" author="COOK, P. L. & CHIMONIDES, P. J." box="[741,1077,165,189]" pageId="10" pageNumber="1610" pagination="97 - 134" refId="ref8006" refString="COOK, P. L. and CHIMONIDES, P. J., 1981 a, Morphology and systematics of some rooted cheilostome Bryozoa, Journal of Natural History, 15, 97 - 134." type="journal article" year="1981">Cook and Chimonides 1981a</bibRefCitation>
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)
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</taxonomicName>
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. The rhizoids of
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<taxonomicName id="4C78A2FF5A4DA056E371D2B7FE7F90FE" authorityName="Gordon" authorityYear="1989" box="[329,426,198,221]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1610" phylum="Bryozoa" rank="species" species="parva">
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<emphasis id="B90C056E5A4DA056E371D2B7FE7F90FE" box="[329,426,198,221]" italics="true" pageId="10" pageNumber="1610">R. parva</emphasis>
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||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C78A2FF5A4DA056E385D2B7FDF090FE" baseAuthorityName="Cook and Chimonides" baseAuthorityYear="1981" box="[445,549,197,221]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1610" phylum="Bryozoa" rank="species" species="cotyla">
|
||
<emphasis id="B90C056E5A4DA056E385D2B7FDF090FE" box="[445,549,197,221]" italics="true" pageId="10" pageNumber="1610">R. cotyla</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C78A2FF5A4DA056E05DD2B7FD0F90FE" authorityName="Cook & Bock" authorityYear="2002" box="[613,730,197,221]" class="Gymnolaemata" family="Petraliidae" genus="Riscodopa" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1610" phylum="Bryozoa" rank="species" species="hyalina">
|
||
<emphasis id="B90C056E5A4DA056E05DD2B7FD0F90FE" box="[613,730,197,221]" italics="true" pageId="10" pageNumber="1610">R. hyalina</emphasis>
|
||
</taxonomicName>
|
||
are numerous, but only one is produced by each septular pore.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |