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<document id="9BE333048F4DBA7182881F930921BB6A" ID-DOI="10.5281/zenodo.183208" ID-GBIF-Dataset="0f46b88c-8545-4bc9-9053-62b52ac326b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="183208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459897967123" checkinUser="plazi" docAuthor="Zhu, Wei-Guang Lian Ming-Sheng &amp; Kury, Adriano B." docDate="2008" docId="03C30F644545285F9BE6BCB3508AFE71" docLanguage="en" docName="zt01841p060.pdf" docOrigin="Zootaxa 1841" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tithaeus drac Zhu &amp; Kury, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FFFA771C4544285B9B71B821551CFFD3" masterDocTitle="A new species of the genus Tithaeus from China (Arachnida: Laniatores: Epedanidae)" masterLastPageNumber="60" masterPageNumber="53" pageNumber="54" updateTime="1698228490626" updateUser="plazi">
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<mods:title id="04060D5BEBDA2FEE62A71A9F534F402F">A new species of the genus Tithaeus from China (Arachnida: Laniatores: Epedanidae)</mods:title>
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<mods:namePart id="D49A113EC6B2E571E0A281211109A65B">Zhu, Wei-Guang Lian Ming-Sheng</mods:namePart>
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<mods:namePart id="79C82EA4AF208930800E7CD103872A70">Kury, Adriano B.</mods:namePart>
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<mods:date id="FE963AE338FD02D0247C2F9AFCBCC6DD">2008</mods:date>
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<treatment id="03C30F644545285F9BE6BCB3508AFE71" ID-DOI="http://doi.org/10.5281/zenodo.6235899" ID-GBIF-Taxon="119372556" ID-Zenodo-Dep="6235899" LSID="urn:lsid:plazi:treatment:03C30F644545285F9BE6BCB3508AFE71" httpUri="http://treatment.plazi.org/id/03C30F644545285F9BE6BCB3508AFE71" lastPageId="4" lastPageNumber="57" pageId="1" pageNumber="54">
<subSubSection id="C370EDF94545285A9BE6BCB355F3FB1C" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8BD5BE724545285A9BE6BCB354BDFB7F" blockId="1.[151,417,1170,1231]" box="[151,417,1170,1196]" pageId="1" pageNumber="54">
<heading id="D09D091E4545285A9BE6BCB354BDFB7F" bold="true" box="[151,417,1170,1196]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="B91E62604545285A9BE6BCB354BDFB7F" bold="true" box="[151,417,1170,1196]" pageId="1" pageNumber="54">
<taxonomicName id="4C6AC5F14545285A9BE6BCB35421FB7F" box="[151,317,1170,1196]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="drac" status="sp. nov.">
<emphasis id="B91E62604545285A9BE6BCB35421FB7F" bold="true" box="[151,317,1170,1196]" italics="true" pageId="1" pageNumber="54">Tithaeus drac</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22DDF1B4545285A9A35BCB354BDFB7F" box="[324,417,1170,1196]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD5BE724545285A9BE6BC9455F3FB1C" blockId="1.[151,417,1170,1231]" box="[151,239,1205,1231]" pageId="1" pageNumber="54">
(
<figureCitation id="1351A2F74545285A9BEEBC9455FFFB1C" box="[159,227,1205,1231]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285A9BE6BD2457C4FA6C" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="8BD5BE724545285A9BE6BD245036FA94" blockId="1.[151,1437,1285,1991]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BE6BD245456FACC" bold="true" box="[151,330,1285,1311]" pageId="1" pageNumber="54">
<typeStatus id="54D100D04545285A9BE6BD2455CEFACC" box="[151,210,1285,1311]" pageId="1" pageNumber="54">Type</typeStatus>
material.
</emphasis>
Female
<typeStatus id="54D100D04545285A9AC1BD245705FACC" box="[432,537,1285,1311]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
,
<collectingCountry id="F37DFEE24545285A9954BD24579AFACC" box="[549,646,1285,1311]" name="China" pageId="1" pageNumber="54">CHINA</collectingCountry>
: Guangxi Province, Longsheng County, Hongtan village, about
<quantity id="4C9213974545285A9E01BD2455B2FA94" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="54" unit="m" value="700.0">700 m</quantity>
alt.,
<geoCoordinate id="EE5ED8B54545285A9B99BD0C544DFA94" box="[232,337,1325,1351]" direction="north" orientation="latitude" pageId="1" pageNumber="54" precision="925" value="25.383333">24°83N</geoCoordinate>
,
<geoCoordinate id="EE5ED8B54545285A9A2DBD0C54D2FA94" box="[348,462,1325,1351]" direction="east" orientation="longitude" pageId="1" pageNumber="54" precision="925" value="110.433334">110°26E</geoCoordinate>
,
<date id="FFD498B24545285A9AABBD0C576BFA94" box="[474,631,1325,1351]" pageId="1" pageNumber="54" value="2005-09-12">12 Sep. 2005</date>
, Gao Chao leg.;
<specimenCount id="9D6C75FB4545285A984ABD0C5696FA94" box="[827,906,1325,1351]" pageId="1" pageNumber="54" type="male">1 male</specimenCount>
<typeStatus id="54D100D04545285A98E0BD0C56E5FA94" box="[913,1017,1325,1351]" pageId="1" pageNumber="54" type="paratype">paratype</typeStatus>
, same details as
<typeStatus id="54D100D04545285A9FCABD0C5039FA94" box="[1211,1317,1325,1351]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BD5BE724545285A9BB7BD7457C4FA6C" blockId="1.[151,1437,1285,1991]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BB7BD7454D9FABC" bold="true" box="[198,453,1365,1391]" pageId="1" pageNumber="54">Additional material.</emphasis>
Two females,
<collectingCountry id="F37DFEE24545285A9902BD7457C8FABC" box="[627,724,1365,1391]" name="China" pageId="1" pageNumber="54">CHINA</collectingCountry>
: Guangxi Province, Tianlin County, Langping village, about
<quantity id="4C9213974545285A9BE6BD5C55F2FA44" box="[151,238,1405,1431]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="1" pageNumber="54" unit="m" value="1125.0">1125 m</quantity>
alt.,
<geoCoordinate id="EE5ED8B54545285A9A59BD5C548CFA44" box="[296,400,1405,1431]" direction="north" orientation="latitude" pageId="1" pageNumber="54" precision="925" value="24.483334">24°29N</geoCoordinate>
, 106°216E,
<date id="FFD498B24545285A9958BD5C5798FA44" box="[553,644,1405,1431]" pageId="1" pageNumber="54" value="1927-05">27 May</date>
, 2006, Ming-Sheng Zhu and Wei-Guang Lian leg. WWF Ecoregion IM0118 (Jian Nan subtropical evergreen forests).
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285A9BB4BDEC517DF9E4" pageId="1" pageNumber="54" type="etymology">
<paragraph id="8BD5BE724545285A9BB4BDEC517DF9E4" blockId="1.[151,1437,1285,1991]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BB4BDEC544EFA34" bold="true" box="[197,338,1485,1511]" pageId="1" pageNumber="54">Etymology.</emphasis>
Species name refers to the fictitious reptilian species called &quot;
<taxonomicName id="4C6AC5F14545285A9F6EBDEC514BFA34" box="[1055,1111,1485,1511]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="drac">drac</taxonomicName>
&quot; from the 1985 science fiction film &quot;Enemy mine&quot;, produced by Twentieth Century Fox and directed by Wolfgang Petersen. The shape of the drac's head is strongly reminiscent of the cheliceral bulla of
<taxonomicName id="4C6AC5F14545285A98D2BE3C56EEF9E5" box="[931,1010,1565,1590]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="drac" status="sp. nov.">
<emphasis id="B91E62604545285A98D2BE3C56EEF9E5" box="[931,1010,1565,1590]" italics="true" pageId="1" pageNumber="54">T. drac</emphasis>
</taxonomicName>
<emphasis id="B91E62604545285A988DBE3C5146F9E4" bold="true" box="[1020,1114,1565,1591]" pageId="1" pageNumber="54">
<taxonomicNameLabel id="A22DDF1B4545285A988DBE3C5146F9E4" box="[1020,1114,1565,1591]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285A9BB4BE64578CF8F5" pageId="1" pageNumber="54" type="diagnosis">
<paragraph id="8BD5BE724545285A9BB4BE64578CF8F5" blockId="1.[151,1437,1285,1991]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BB4BE645459F98C" bold="true" box="[197,325,1605,1631]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="4C6AC5F14545285A99BEBE6456F2F98C" authority="Suzuki, 1985" authorityName="Suzuki" authorityYear="1985" box="[719,1006,1605,1631]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91E62604545285A99BEBE645620F98D" box="[719,828,1605,1630]" italics="true" pageId="1" pageNumber="54">T. similis</emphasis>
<bibRefCitation id="EFFBC3834545285A983DBE6456F2F98C" author="Suzuki" box="[844,1006,1605,1631]" pageId="1" pageNumber="59" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11, 69 - 110." type="journal article" year="1985">Suzuki, 1985</bibRefCitation>
</taxonomicName>
, described from northern
<collectingCountry id="F37DFEE24545285A9E45BE645080F98C" box="[1332,1436,1605,1631]" name="Thailand" pageId="1" pageNumber="54">Thailand</collectingCountry>
(
<bibRefCitation id="EFFBC3834545285A9BEEBE4C54C5F954" author="Suzuki" box="[159,473,1645,1671]" pageId="1" pageNumber="59" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11, 69 - 110." type="journal article" year="1985">Suzuki, 1985: 7879, fig 5</bibRefCitation>
) in general appearance, but differs distinctly from the latter by the following characters: 1) Female materials with larger body in size, chelicera and palpus strong and completely similar to male; 2) cheliceral bulla of male with notable posterior granulous elevation, absent on
<taxonomicName id="4C6AC5F14545285A9FE7BE9C51E2F905" box="[1174,1278,1725,1750]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91E62604545285A9FE7BE9C51E2F905" box="[1174,1278,1725,1750]" italics="true" pageId="1" pageNumber="54">T. similis</emphasis>
</taxonomicName>
; 3) the distal margin of ventral plate of penis has a small shallow cleft compared to the deeper cleft of
<taxonomicName id="4C6AC5F14545285A9FBEBEC45024F92D" box="[1231,1336,1765,1790]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91E62604545285A9FBEBEC45024F92D" box="[1231,1336,1765,1790]" italics="true" pageId="1" pageNumber="54">T. similis</emphasis>
</taxonomicName>
and the setae of the ovipositor are distinctly longer.
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285F9BB4BF1456D8FEF9" lastPageId="4" lastPageNumber="57" pageId="1" pageNumber="54" type="description">
<paragraph id="8BD5BE724545285A9BB4BF14515CF814" blockId="1.[151,1437,1285,1991]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BB4BF145441F89C" bold="true" box="[197,349,1845,1871]" pageId="1" pageNumber="54">Description.</emphasis>
Female (
<typeStatus id="54D100D04545285A9ABDBF14572AF89C" box="[460,566,1845,1871]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
) habitus as in
<figureCitation id="1351A2F74545285A9993BF14563BF89C" box="[738,807,1845,1871]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="1" pageNumber="54">Figs 1</figureCitation>
AB. Coloration: body and appendages entirely dark brown; carapace and eye tubercle with yellow-brown reticulation; lateral margins and opisthosomal areas of scutum, and free tergites banded with black brown; chelicerae and palpus dark brown, with yellow dots above; legs dark brown, with yellow dots on the dorsal surface from trochanter to tibia.
</paragraph>
<caption id="DF15EEFA454628599BE6BEE25745F8B6" httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="2" pageNumber="55" targetBox="[189,1421,194,1689]" targetPageId="2">
<paragraph id="8BD5BE72454628599BE6BEE25745F8B6" blockId="2.[151,1436,1731,1893]" pageId="2" pageNumber="55">
<emphasis id="B91E6260454628599BE6BEE25405F908" bold="true" box="[151,281,1731,1755]" pageId="2" pageNumber="55">FIGURE 1.</emphasis>
<taxonomicName id="4C6AC5F1454628599A51BEE254AFF909" box="[288,435,1731,1754]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="drac" status="sp. nov.">
<emphasis id="B91E6260454628599A51BEE254AFF909" box="[288,435,1731,1754]" italics="true" pageId="2" pageNumber="55">Tithaeus drac</emphasis>
</taxonomicName>
<emphasis id="B91E6260454628599ACBBEE25713F908" bold="true" box="[442,527,1731,1755]" pageId="2" pageNumber="55">
<taxonomicNameLabel id="A22DDF1B454628599ACBBEE25713F908" box="[442,527,1731,1755]" pageId="2" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AH female holotype, IL male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: AB, 1.0 mm; CG, 0.5 mm; H, L, 0.2 mm; IK, 0.1mm.
</paragraph>
</caption>
<paragraph id="8BD5BE72454628589BB4BFB85752FF09" blockId="2.[151,1436,1945,2011]" lastBlockId="3.[151,1436,152,218]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">Dorsal scutum trapezoid in shape; abdomen blunt behind. Eye tubercle broad oval, removed from the anterior border of carapace, without a median spine. Opisthosomal region of scutum with five areas. A row of rounded tubercles along the lateral margins of the scutum. Free tergites each with a transverse row of conical tubercles spread over its entire width.</paragraph>
<caption id="DF15EEFA454728589BE6BBA15420FC2C" httpUri="https://zenodo.org/record/183210/files/figure.png" pageId="3" pageNumber="56" targetBox="[167,1432,283,847]" targetPageId="3">
<paragraph id="8BD5BE72454728589BE6BBA15420FC2C" blockId="3.[151,1436,896,1023]" pageId="3" pageNumber="56">
<emphasis id="B91E6260454728589BE6BBA15403FC4B" bold="true" box="[151,287,896,920]" pageId="3" pageNumber="56">FIGURE 2.</emphasis>
A,
<taxonomicName id="4C6AC5F1454728589A23BBA057D5FC44" authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" box="[338,713,896,920]" class="Arachnida" family="Epedanidae" genus="Kilungius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B91E6260454728589A23BBA05730FC4B" box="[338,556,897,920]" italics="true" pageId="3" pageNumber="56">Kilungius insulanus</emphasis>
(Hirst, 1911)
</taxonomicName>
(redrawn from Suzuki 1973). B,
<taxonomicName id="4C6AC5F1454728589F21BBA05080FC44" authority="Suzuki, 1985" authorityName="Suzuki" authorityYear="1985" box="[1104,1436,896,920]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91E6260454728589F21BBA051E1FC4B" box="[1104,1277,897,920]" italics="true" pageId="3" pageNumber="56">Tithaeus similis</emphasis>
Suzuki, 1985
</taxonomicName>
(redrawn from Suzuki 1985). C,
<taxonomicName id="4C6AC5F1454728589A84BB825689FC69" authority="Ubick &amp; Briggs, 1992" authorityName="Ubick &amp; Briggs" authorityYear="1992" box="[501,917,931,954]" class="Arachnida" family="Phalangodidae" genus="Texella" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="bilobata">
<emphasis id="B91E6260454728589A84BB8257BDFC69" box="[501,673,931,954]" italics="true" pageId="3" pageNumber="56">Texella bilobata</emphasis>
Ubick &amp; Briggs, 1992
</taxonomicName>
(redrawn from Ubick &amp; Briggs 1992). Abbreviations: S = stylus, SL = stylar lobe, BS = basal sac, BK = basal knob, ML = middle lobe, SA = stypar apophysis, PSL = parastylar lobe.
</paragraph>
</caption>
<paragraph id="8BD5BE72454728589BB4BC125148FB16" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">Venter. Coxa I armed with two transverse rows of hair-tipped granules, these distinctly larger than the remaining granules scattered on coxae IIIV. Dorsal surface of coxa IV with several slightly larger hair-tipped tubercles. Coxa III with a row of low humps along the frontal and rear margins. Tracheal stigma clearly visible. A transverse row of very small, hair-tipped granules across each free sternite.</paragraph>
<paragraph id="8BD5BE72454728589BB4BCF2506BFAEE" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">
Chelicera (
<figureCitation id="1351A2F7454728589A35BCF25495FB3E" box="[324,393,1235,1261]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">Figs 1</figureCitation>
CE). Proximal segment relatively long, visibly swollen disto-dorsally, with notable posterior granulous elevation. Second segment widened, with two slightly larger tubercles in the middle of the prodorsal surface (
<figureCitation id="1351A2F7454728589A00BD0254AFFAEE" box="[369,435,1315,1341]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
E). Fingers relatively short but strong; inner edges toothed as illustrated (
<figureCitation id="1351A2F7454728589E66BD025040FAEE" box="[1303,1372,1315,1341]" captionStart="FIGURE 2. A" captionStartId="3.[151,255,896,920]" captionTargetBox="[167,1432,283,847]" captionTargetId="figure@3.[151,1436,264,869]" captionTargetPageId="3" captionText="FIGURE 2. A, Kilungius insulanus (Hirst, 1911) (redrawn from Suzuki 1973). B, Tithaeus similis Suzuki, 1985 (redrawn from Suzuki 1985). C, Texella bilobata Ubick &amp; Briggs, 1992 (redrawn from Ubick &amp; Briggs 1992). Abbreviations: S = stylus, SL = stylar lobe, BS = basal sac, BK = basal knob, ML = middle lobe, SA = stypar apophysis, PSL = parastylar lobe." httpUri="https://zenodo.org/record/183210/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
E)
</paragraph>
<paragraph id="8BD5BE72454728589BB4BD6A55F2F9FE" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">
Palpus (
<figureCitation id="1351A2F7454728589A56BD6A5473FAB6" box="[295,367,1355,1381]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">Figs 1</figureCitation>
FG) short and robust, femur especially so. Trochanter ventrally with a short setiferous tubercle followed transversely by a small hair-tipped tubercle. Femur ventrally with three setiferous tubercles, distally on prolateral side with a short setiferous tubercle as illustrated. Patella medio-distally with a setiferous tubercle. The ventral surface of tibia and tarsus with three prolateral and retrolateral setiferous tubercles respectively arranged as illustrated (
<figureCitation id="1351A2F7454728589939BDCA5793F9D6" box="[584,655,1515,1541]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">Figs 1</figureCitation>
FG). Tarsal claw longer than one half length of tarsus, strongly curved.
</paragraph>
<paragraph id="8BD5BE72454728589BB4BE1A5710F976" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">
Legs slender and relatively elongated. Femora IIV straight. All leg segments unarmed, with very short hairs. Tarsi IIIIV with simple double claws, no scopulae. Tarsal formula:
<date id="FFD498B2454728589F73BE425151F9AE" box="[1026,1101,1635,1661]" pageId="3" pageNumber="56" value="1911-05-05">5/11/5</date>
/6. Distitarsi of first and second tarsi each with two tarsalia.
</paragraph>
<paragraph id="8BD5BE72454728589BB4BE92517CF91E" blockId="3.[151,1436,1075,1981]" box="[197,1120,1715,1741]" pageId="3" pageNumber="56">
Ovipositor as illustrated (
<figureCitation id="1351A2F7454728589A9EBE92572DF91E" box="[495,561,1715,1741]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
H). Each lobe with 2 ventral and 2 dorsal setae.
</paragraph>
<paragraph id="8BD5BE72454728589BB4BEFA55F5F8CE" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">
Male. Shape, coloration and markings similar to the female, but smaller in size. Tarsal formula (IIV):
<date id="FFD498B2454728589EF4BEFA55D0F8CE" pageId="3" pageNumber="56" value="1910-05-05">5/ 10/5</date>
/6.
</paragraph>
<paragraph id="8BD5BE72454728589BB4BF0A5675F86E" blockId="3.[151,1436,1075,1981]" pageId="3" pageNumber="56">
Penis (
<figureCitation id="1351A2F7454728589A64BF0A5446F896" box="[277,346,1835,1861]" captionStart="FIGURE 1" captionStartId="2.[151,255,1731,1755]" captionTargetBox="[189,1421,194,1689]" captionTargetId="figure@2.[151,1436,182,1712]" captionTargetPageId="2" captionText="FIGURE 1. Tithaeus drac sp. nov. (A H female holotype, I L male paratype). A. Female body, dorsal view. B, Female body, lateral view. C, Left chelicera, female, anterior view. D, Left chelicera, female, posterior view. E, Distal segment of the left chelicera, female, above view. F, Left palpus, female, anterior view. G, Left palpus, female, posterior view. H, Ovipositor. I, Penis, lateral view. J, Penis, ventral view. K, Penis, dorsal view. L, Whole penis. Scale bars: A B, 1.0 mm; C G, 0.5 mm; H, L, 0.2 mm; I K, 0.1 mm." httpUri="https://zenodo.org/record/183209/files/figure.png" pageId="3" pageNumber="56">
Figs
<date id="FFD498B2454728589A21BF0A5446F896" box="[336,346,1835,1861]" pageId="3" pageNumber="56">1</date>
</figureCitation>
IL
<emphasis id="B91E6260454728589AF8BF0A548FF896" bold="true" box="[393,403,1835,1861]" pageId="3" pageNumber="56">)</emphasis>
. Ventral plate with a moderate cleft in middle of distal margin; setae arranged as shown in
<figureCitation id="1351A2F7454728589BC7BF7255E0F8BE" box="[182,252,1875,1901]" captionStart="FIGURE 2. A" captionStartId="3.[151,255,896,920]" captionTargetBox="[167,1432,283,847]" captionTargetId="figure@3.[151,1436,264,869]" captionTargetPageId="3" captionText="FIGURE 2. A, Kilungius insulanus (Hirst, 1911) (redrawn from Suzuki 1973). B, Tithaeus similis Suzuki, 1985 (redrawn from Suzuki 1985). C, Texella bilobata Ubick &amp; Briggs, 1992 (redrawn from Ubick &amp; Briggs 1992). Abbreviations: S = stylus, SL = stylar lobe, BS = basal sac, BK = basal knob, ML = middle lobe, SA = stypar apophysis, PSL = parastylar lobe." httpUri="https://zenodo.org/record/183210/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
J. Basal sac inflatable, developed immovable and partly sunken into truncus, lacking complex introverting structures, stylus smooth and arising straight from the glans, stylar tip ended sharply and unexpanded, stylar lobe entire and warped upwards surrounding the stylus.
</paragraph>
<paragraph id="8BD5BE724540285F9BB4B8B65720FED1" blockId="4.[151,1436,151,338]" pageId="4" pageNumber="57">
<emphasis id="B91E62604540285F9BB4B8B65460FF62" bold="true" box="[197,380,151,177]" pageId="4" pageNumber="57">Measurements</emphasis>
:
<typeStatus id="54D100D04540285F9AFDB8B954E5FF61" box="[396,505,152,178]" pageId="4" pageNumber="57" type="holotype">Holotype</typeStatus>
female (male
<typeStatus id="54D100D04540285F99D1B8B95619FF61" box="[672,773,152,178]" pageId="4" pageNumber="57" type="paratype">paratype</typeStatus>
in parentheses): Body 3.97 (3.89) long, 2.38 (2.38) wide at the widest portion, scutum 3.10 (3.02) long; eye tubercle 0.68 (0.65) long; 0.35 (0.35) wide. Measurements of left palpus and legs as in Table 1.
</paragraph>
<paragraph id="8BD5BE724540285F9BB7B92E56D8FEF9" blockId="4.[151,1436,151,338]" box="[198,964,271,298]" pageId="4" pageNumber="57">
<emphasis id="B91E62604540285F9BB7B92E5432FEFA" bold="true" box="[198,302,271,297]" pageId="4" pageNumber="57">Habitat.</emphasis>
Collected under humid fallen logs on a hill near village.
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94540285F9BB7B916508AFE71" pageId="4" pageNumber="57" type="distribution">
<paragraph id="8BD5BE724540285F9BB7B9165783FE81" blockId="4.[151,1436,151,338]" box="[198,671,311,338]" pageId="4" pageNumber="57">
<emphasis id="B91E62604540285F9BB7B9165443FE82" bold="true" box="[198,351,311,337]" pageId="4" pageNumber="57">Distribution</emphasis>
: Guangxi Province,
<collectingCountry id="F37DFEE24540285F9920B9195787FE81" box="[593,667,312,338]" name="China" pageId="4" pageNumber="57">China</collectingCountry>
.
</paragraph>
<paragraph id="8BD5BE724540285F9BE6B9A6508AFE71" blockId="4.[151,1430,391,418]" box="[151,1430,391,418]" pageId="4" pageNumber="57">
<emphasis id="B91E62604540285F9BE6B9A65409FE72" bold="true" box="[151,277,391,417]" pageId="4" pageNumber="57">TABLE 1.</emphasis>
Measurements (in mm) of palpus and legs of female
<typeStatus id="54D100D04540285F98F6B9A956F2FE71" box="[903,1006,392,418]" pageId="4" pageNumber="57" type="holotype">holotype</typeStatus>
and male
<typeStatus id="54D100D04540285F9F19B9A951D1FE71" box="[1128,1229,392,418]" pageId="4" pageNumber="57" type="paratype">paratype</typeStatus>
(in parentheses).
</paragraph>
</subSubSection>
</treatment>
</document>