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<document id="43C49D473FC2B4CC8A67E2A654CC01B1" ID-DOI="10.5281/zenodo.183208" ID-GBIF-Dataset="0f46b88c-8545-4bc9-9053-62b52ac326b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="183208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459897967123" checkinUser="plazi" docAuthor="Zhu, Wei-Guang Lian Ming-Sheng &amp; Kury, Adriano B." docDate="2008" docId="03C30F644544285A9BE6BF5D5642FBE7" docLanguage="en" docName="zt01841p060.pdf" docOrigin="Zootaxa 1841" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tithaeus Thorell 1891" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FFFA771C4544285B9B71B821551CFFD3" masterDocTitle="A new species of the genus Tithaeus from China (Arachnida: Laniatores: Epedanidae)" masterLastPageNumber="60" masterPageNumber="53" pageNumber="53" updateTime="1698228490626" updateUser="plazi">
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<mods:title id="37DF5536B12E905D44A7C5134D85AD86">A new species of the genus Tithaeus from China (Arachnida: Laniatores: Epedanidae)</mods:title>
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<mods:namePart id="0B6FD311D820A274B5F33469DFB76BD3">Zhu, Wei-Guang Lian Ming-Sheng</mods:namePart>
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<mods:namePart id="E0521FA2FB92F680FCEC450934E3B6D1">Kury, Adriano B.</mods:namePart>
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<mods:date id="F90ECAEC0149D9410BF6D1708BB43318">2008</mods:date>
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<taxonomicName id="4C6AC5F14544285B9BE6BF5D54B1F845" authority="Thorell, 1891" authorityName="Thorell" authorityYear="1891" box="[151,429,1916,1942]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604544285B9BE6BF5D54B1F845" bold="true" box="[151,429,1916,1942]" pageId="0" pageNumber="53">
<emphasis id="B91E62604544285B9BE6BF5D55E3F845" bold="true" box="[151,255,1916,1942]" italics="true" pageId="0" pageNumber="53">Tithaeus</emphasis>
<bibRefCitation id="EFFBC3834544285B9A77BF5D54B1F845" author="Thorell" box="[262,429,1916,1942]" pageId="0" pageNumber="59" refString="Thorell, T. T. T. (1891) Aracnidi di Pinang raccolti nel 1889 dai Sig. ri L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 30, 269 - 383." type="journal article" year="1891">Thorell, 1891</bibRefCitation>
</emphasis>
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<paragraph id="8BD5BE724544285B9BE6BF9C502CF820" blockId="0.[151,1436,1980,2035]" pageId="0" pageNumber="53">
<treatmentCitationGroup id="AB7A995C4544285B9BE6BF9C502CF820" pageId="0" pageNumber="53">
<taxonomicName id="4C6AC5F14544285B9BE6BF9C54D8F800" authority="Thorell, 1891: 371" authorityName="Thorell" authorityPageNumber="371" authorityYear="1891" box="[151,452,1980,2004]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604544285B9BE6BF9C55EDF807" box="[151,241,1981,2004]" italics="true" pageId="0" pageNumber="53">Tithaeus</emphasis>
<treatmentCitation id="0ACB98634544285B9B88BF9D54D8F800" author="Thorell" box="[249,452,1980,2003]" page="371" pageId="0" pageNumber="53" year="1891">
<bibRefCitation id="EFFBC3834544285B9B88BF9D5493F800" author="Thorell" box="[249,399,1980,2003]" pageId="0" pageNumber="59" refString="Thorell, T. T. T. (1891) Aracnidi di Pinang raccolti nel 1889 dai Sig. ri L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 30, 269 - 383." type="journal article" year="1891">Thorell, 1891</bibRefCitation>
: 371
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</taxonomicName>
,
<typeStatus id="54D100D04544285B9AA1BF9D54E2F800" box="[464,510,1980,2003]" pageId="0" pageNumber="53">type</typeStatus>
species by original designation:
<taxonomicName id="4C6AC5F14544285B9813BF9C51D3F800" authority="Thorell, 1891" authorityName="Thorell" authorityYear="1891" box="[866,1231,1980,2004]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="B91E62604544285B9813BF9C512EF807" box="[866,1074,1981,2004]" italics="true" pageId="0" pageNumber="53">Tithaeus laevigatus</emphasis>
<bibRefCitation id="EFFBC3834544285B9F4BBF9D51D3F800" author="Thorell" box="[1082,1231,1980,2003]" pageId="0" pageNumber="59" refString="Thorell, T. T. T. (1891) Aracnidi di Pinang raccolti nel 1889 dai Sig. ri L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 30, 269 - 383." type="journal article" year="1891">Thorell, 1891</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="0ACB98634544285B9FAABF9D508BF800" author="Loman" box="[1243,1431,1980,2003]" page="33" pageId="0" pageNumber="53" year="1905">
<bibRefCitation id="EFFBC3834544285B9FAABF9D5073F800" author="Loman" box="[1243,1391,1980,2003]" pageId="0" pageNumber="59" refString="Loman, J. C. C. (1905) Opilioniden aus Java. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, Hamburg, 22 (2) (1904 - 1905), 29 - 34." type="journal article" year="1905">
<collectingCountry id="F37DFEE24544285B9FAABF9D5035F800" box="[1243,1321,1980,2003]" name="Oman" pageId="0" pageNumber="53">Loman</collectingCountry>
, 1905
</bibRefCitation>
: 33
</treatmentCitation>
;
<treatmentCitation id="0ACB98634544285B9BB6BFFD548DF820" author="Roewer" box="[199,401,2012,2035]" page="120" pageId="0" pageNumber="53" year="1912">
<bibRefCitation id="EFFBC3834544285B9BB6BFFD5443F820" author="Roewer" box="[199,351,2012,2035]" pageId="0" pageNumber="59" refString="Roewer, C. - Fr. (1912) Die Familien der Assamiiden und Phalangodiden der Opiliones-Laniatores (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv fur Naturgeschichte, (A), 78, 1 - 242." type="journal article" year="1912">Roewer, 1912</bibRefCitation>
: 120
</treatmentCitation>
(in part); 1923: 79; 1927: 279; 1949: 44; Banks, 1931: 67;
<treatmentCitation id="0ACB98634544285B9F7BBFFD51D3F820" author="Suzuki" box="[1034,1231,2012,2035]" page="24" pageId="0" pageNumber="53" year="1969">
<bibRefCitation id="EFFBC3834544285B9F7BBFFD51BAF820" author="Suzuki" box="[1034,1190,2012,2035]" pageId="0" pageNumber="59" refString="Suzuki, S. (1969 a) On a collection of Opilionids from Southeast Asia. Journal of Science of the Hiroshima University (B 1), 22, 11 - 77." type="journal article" year="1969" yearSuffix="a">Suzuki, 1969a</bibRefCitation>
: 24
</treatmentCitation>
; 1972: 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD5BE724545285A9BE6B8B956F8FF1C" blockId="1.[151,1341,152,207]" pageId="1" pageNumber="54">
<treatmentCitationGroup id="AB7A995C4545285A9BE6B8B956F8FF1C" pageId="1" pageNumber="54">
<taxonomicName id="4C6AC5F14545285A9BE6B8B95494FF7C" ID-CoL="8KYX8" authority="Loman, 1892: 12" authorityName="Loman" authorityPageNumber="12" authorityYear="1892" box="[151,392,152,175]" class="Insecta" family="Carabidae" genus="Sinis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A9BE6B8B955D7FF7C" box="[151,203,152,175]" italics="true" pageId="1" pageNumber="54">Sinis</emphasis>
<treatmentCitation id="0ACB98634545285A9BA0B8B95494FF7C" author="Loman" box="[209,392,152,175]" page="12" pageId="1" pageNumber="54" year="1892">
<bibRefCitation id="EFFBC3834545285A9BA0B8B9547FFF7C" author="Loman" box="[209,355,152,175]" pageId="1" pageNumber="59" refString="Loman, J. C. C. (1892) Opilioniden von Sumatra, Java und Flores. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, vol. 3, pp. 1 - 27, table 1. E. J. Brill, Leiden (1894). 476 pp., 22 plates." type="book chapter" year="1892">
<collectingCountry id="F37DFEE24545285A9BA0B8B95403FF7C" box="[209,287,152,175]" name="Oman" pageId="1" pageNumber="54">Loman</collectingCountry>
, 1892
</bibRefCitation>
: 12
</treatmentCitation>
</taxonomicName>
[junior homonym of
<taxonomicName id="4C6AC5F14545285A991FB8B956B9FF7C" authority="Heer 1862 (Coleoptera)" authorityName="Heer 1862 (Coleoptera" box="[622,933,152,175]" class="Insecta" family="Carabidae" genus="Sinis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A991FB8B957BDFF7C" box="[622,673,152,175]" italics="true" pageId="1" pageNumber="54">Sinis</emphasis>
Heer 1862 (Coleoptera)
</taxonomicName>
and of
<taxonomicName id="4C6AC5F14545285A9887B8B9502FFF7C" authority="Thorell 1878" authorityName="Thorell" authorityYear="1878" box="[1014,1331,152,175]" class="Insecta" family="Carabidae" genus="Sinis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A9887B8B95135FF7C" box="[1014,1065,152,175]" italics="true" pageId="1" pageNumber="54">Sinis</emphasis>
Thorell 1878 (Araneae)
</taxonomicName>
].
<taxonomicName id="4C6AC5F14545285A9BE6B89954CCFF1C" authority="Loman, 1902: 198" authorityName="Loman" authorityPageNumber="198" authorityYear="1902" box="[151,464,184,207]" class="Arachnida" family="Epedanidae" genus="Sinniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A9BE6B899541AFF1C" box="[151,262,184,207]" italics="true" pageId="1" pageNumber="54">Sinniculus</emphasis>
<treatmentCitation id="0ACB98634545285A9A7DB89954CCFF1C" author="Loman" box="[268,464,184,207]" page="198" pageId="1" pageNumber="54" year="1902">
<bibRefCitation id="EFFBC3834545285A9A7DB8995482FF1C" author="Loman" box="[268,414,184,207]" pageId="1" pageNumber="59" refString="Loman, J. C. C. (1902) Neue aussereuropaische Opilioniden. Zoologische Jahrb u cher, Jena, Abteilung f u r Systematik, O kologie und Geographie der Tiere, 16 (2), 163 - 216." type="journal article" year="1902">
<collectingCountry id="F37DFEE24545285A9A7DB8995446FF1C" box="[268,346,184,207]" name="Oman" pageId="1" pageNumber="54">Loman</collectingCountry>
, 1902
</bibRefCitation>
: 198
</treatmentCitation>
</taxonomicName>
[valid replacement name for
<taxonomicName id="4C6AC5F14545285A987AB89956C6FF1C" authority="Loman, 1892" authorityName="Loman" authorityYear="1892" box="[779,986,184,207]" class="Insecta" family="Carabidae" genus="Sinis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A987AB8995623FF1C" box="[779,831,184,207]" italics="true" pageId="1" pageNumber="54">Sinis</emphasis>
<bibRefCitation id="EFFBC3834545285A9837B89956C6FF1C" author="Loman" box="[838,986,184,207]" pageId="1" pageNumber="59" refString="Loman, J. C. C. (1892) Opilioniden von Sumatra, Java und Flores. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, vol. 3, pp. 1 - 27, table 1. E. J. Brill, Leiden (1894). 476 pp., 22 plates." type="book chapter" year="1892">
<collectingCountry id="F37DFEE24545285A9837B8995688FF1C" box="[838,916,184,207]" name="Oman" pageId="1" pageNumber="54">Loman</collectingCountry>
, 1892
</bibRefCitation>
</taxonomicName>
].
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</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285A9BE6B8DB5170FDFF" pageId="1" pageNumber="54" type="diagnosis">
<paragraph id="8BD5BE724545285A9BE6B8DB5170FDFF" blockId="1.[151,1437,250,1076]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BE6B8DB540BFEC7" bold="true" box="[151,279,250,276]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
Medium-sized epedanids with a low or moderate common eye tubercle removed from the anterior margin of carapace, without a median spine. Few with a slight hump situated between the eye tubercle and the anterior margin, lower than eye tubercle. Scutal region divided into five areas. Palpus relatively short and thickened, its femur and patella each usually provided with a setiferous tubercle medially-distally. Tarsal formula (I¦IV): 5: more than 5: 5: 6. Distitarsi of first and second tarsus usually with two tarsalia each. Tarsi III IV with simple and smooth double claws, no scopulae. Distal margin of ventral plate of penis usually with deep cleft, glans with simple membranous lobe to protect the stylus, stylar tip ending with a bifurcate lobe or slightly inflatable. Each lobe of ovipositor usually with 2 ventral and 2 dorsal setae.
</paragraph>
</subSubSection>
<subSubSection id="C370EDF94545285A9BB7BA1B5642FBE7" pageId="1" pageNumber="54" type="distribution">
<paragraph id="8BD5BE724545285A9BB7BA1B5642FBE7" blockId="1.[151,1437,250,1076]" pageId="1" pageNumber="54">
<emphasis id="B91E62604545285A9BB7BA1B547BFD87" bold="true" box="[198,359,570,596]" pageId="1" pageNumber="54">Distribution.</emphasis>
Indo-Malaysian Region:
<collectingCountry id="F37DFEE24545285A99D5BA1B57E8FD87" box="[676,756,570,596]" name="Myanmar" pageId="1" pageNumber="54">Burma</collectingCountry>
(
<collectingCountry id="F37DFEE24545285A9878BA1B569DFD87" box="[777,897,570,596]" name="Myanmar" pageId="1" pageNumber="54">Myanmar</collectingCountry>
),
<collectingCountry id="F37DFEE24545285A98EABA1B510CFD87" box="[923,1040,570,596]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
,
<collectingCountry id="F37DFEE24545285A9F50BA1B518CFD87" box="[1057,1168,570,596]" name="Malaysia" pageId="1" pageNumber="54">Malaysia</collectingCountry>
,
<collectingCountry id="F37DFEE24545285A9FD0BA1B5007FD87" box="[1185,1307,570,596]" name="Singapore" pageId="1" pageNumber="54">Singapore</collectingCountry>
,
<collectingCountry id="F37DFEE24545285A9E5DBA1B508BFD87" box="[1324,1431,570,596]" name="Thailand" pageId="1" pageNumber="54">Thailand</collectingCountry>
, (
<bibRefCitation id="EFFBC3834545285A9BEEBA43545CFDAF" author="Roewer" box="[159,320,610,636]" pageId="1" pageNumber="59" refString="Roewer, C. - Fr. (1912) Die Familien der Assamiiden und Phalangodiden der Opiliones-Laniatores (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv fur Naturgeschichte, (A), 78, 1 - 242." type="journal article" year="1912">Roewer 1912</bibRefCitation>
,
<bibRefCitation id="EFFBC3834545285A9A3ABA435494FDAF" author="Roewer" box="[331,392,610,636]" pageId="1" pageNumber="59" refString="Roewer, C. - Fr. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1116 pp." type="book" year="1923">1923</bibRefCitation>
,
<bibRefCitation id="EFFBC3834545285A9AE5BA4354CEFDAF" author="Roewer" box="[404,466,610,636]" pageId="1" pageNumber="59" refString="Roewer, C. - Fr. (1949) &quot; ber Phalangodiden I. (Subfam. Phalangodinae, Tricommatinae, Samoinae.) Weitere Weberknechte XIII. Senckenbergiana, 30, 11 - 61." type="journal article" year="1949">1949</bibRefCitation>
;
<bibRefCitation id="EFFBC3834545285A9AAFBA43579CFDAF" author="Suzuki" box="[478,640,610,636]" pageId="1" pageNumber="59" refString="Suzuki, S. (1969 a) On a collection of Opilionids from Southeast Asia. Journal of Science of the Hiroshima University (B 1), 22, 11 - 77." type="journal article" year="1969" yearSuffix="a">Suzuki 1969a</bibRefCitation>
,
<bibRefCitation id="EFFBC3834545285A99FABA4357CBFDAF" author="Suzuki" box="[651,727,610,636]" pageId="1" pageNumber="59" refString="Suzuki, S. (1969 b) Some phalangids from Thailand. Journal of Science of the Hiroshima University (B 1), 22, 79 - 101." type="journal article" year="1969" yearSuffix="b">1969b</bibRefCitation>
,
<bibRefCitation id="EFFBC3834545285A9992BA43563CFDAF" author="Suzuki" box="[739,800,610,636]" pageId="1" pageNumber="59" refString="Suzuki, S. (1972) Opiliones of Semangkok Forest Reserve, Malaysia. Journal of Science of the Hiroshima University (B 1), 24, 1 - 29." type="journal article" year="1972">1972</bibRefCitation>
,
<bibRefCitation id="EFFBC3834545285A985DBA435674FDAF" author="Suzuki" box="[812,872,610,636]" pageId="1" pageNumber="59" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11, 69 - 110." type="journal article" year="1985">1985</bibRefCitation>
) and
<collectingCountry id="F37DFEE24545285A98D9BA4356F2FDAF" box="[936,1006,610,636]" name="China" pageId="1" pageNumber="54">China</collectingCountry>
(new record). There is a record from Timor in the Australasian Region, but this is highly doubtful because it does not fit with the known distribution of the genus, otherwise confined to the Indo-Malaysian side of Wallaces Line (
<figureCitation id="1351A2F74545285A9F0CBA9351D9FD1F" box="[1149,1221,690,716]" captionStart="FIGURE 3" captionStartId="4.[151,255,1906,1930]" captionTargetBox="[151,1436,775,1879]" captionTargetId="figure@4.[151,1436,775,1879]" captionTargetPageId="4" captionText="FIGURE 3. Southeast Asia, showing the distribution of species of Tithaeus from literature records (open circles), the doubtful record for Timor (black circle) and the records for the new species described here: Tithaeus drac (black triangle)." httpUri="https://zenodo.org/record/183211/files/figure.png" pageId="1" pageNumber="54">Fig. 3</figureCitation>
). Timor is a common word in Bahasa
<collectingCountry id="F37DFEE24545285A9AECBAFB5713FD27" box="[413,527,730,756]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
and applies to many different places. In the absence of further records of
<taxonomicName id="4C6AC5F14545285A9BE6BB2355E6FCC8" box="[151,250,770,795]" class="Arachnida" family="Epedanidae" genus="Tithaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A9BE6BB2355E6FCC8" box="[151,250,770,795]" italics="true" pageId="1" pageNumber="54">Tithaeus</emphasis>
</taxonomicName>
from Australasia, and in view of the notorious cases of mislabelled localities in the Roewer collection (e.g.,
<bibRefCitation id="EFFBC3834545285A9A7FBB0B5752FC97" author="Helversen" box="[270,590,810,836]" pageId="1" pageNumber="59" refString="Helversen, O. von &amp; Martens, J. (1972) Unrichtige Fundort-Angaben in der Arachniden-Sammlung Roewer. Senckenbergiana Biologica, 53, 109 - 123." type="journal article" year="1972">Helversen &amp; Martens 1972</bibRefCitation>
;
<bibRefCitation id="EFFBC3834545285A9928BB0B57C6FC97" author="Kury" box="[601,730,810,836]" pageId="1" pageNumber="59" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologi a, vol. especial monogr. fico, n ° 1, 1 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
), it is reasonable to assume a mislabeling by Roewer. On the other hand, significance of the Wallaces Line and other related lines in the Wallacea is evident for mammals, birds, some reptiles, and freshwater fish. However, biogeographical boundaries between Oriental and Australasia are rather obscure for insects and arachnids in the Wallacea. In
<taxonomicName id="4C6AC5F14545285A9897BB835144FC6F" box="[998,1112,930,956]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
there are many representatives that are distributed in both western and eastern part of
<collectingCountry id="F37DFEE24545285A98E4BBEB511BFC37" box="[917,1031,970,996]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
beyond the Wallaces Line, like laniatorids Dampetrinae, podoctids such as
<taxonomicName id="4C6AC5F14545285A99E4BBD356BBFBDF" authority="Karsch, 1880" authorityName="Karsch" authorityYear="1880" box="[661,935,1010,1036]" class="Arachnida" family="Podoctidae" genus="Ibalonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A99E4BBD35618FBD8" box="[661,772,1010,1035]" italics="true" pageId="1" pageNumber="54">Ibalonius</emphasis>
Karsch, 1880
</taxonomicName>
and epedanids such as
<taxonomicName id="4C6AC5F14545285A9FC9BBD355CEFBE7" authority="Thorell, 1891" authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Epedanidae" genus="Beloniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91E62604545285A9FC9BBD3502BFBD8" box="[1208,1335,1010,1035]" italics="true" pageId="1" pageNumber="54">Beloniscus</emphasis>
<bibRefCitation id="EFFBC3834545285A9E4FBBD355CEFBE7" author="Thorell" pageId="1" pageNumber="59" refString="Thorell, T. T. T. (1891) Aracnidi di Pinang raccolti nel 1889 dai Sig. ri L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 30, 269 - 383." type="journal article" year="1891">Thorell, 1891</bibRefCitation>
</taxonomicName>
(assuming these taxa do represent monophyletic units).
</paragraph>
</subSubSection>
</treatment>
</document>