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<document id="98A6B5252BCC71C407E486ABBAF84C14" ID-CLB-Dataset="54702" ID-DOI="10.11646/phytotaxa.600.3.4" ID-GBIF-Dataset="2a1cbd1e-993f-4c7c-b5ed-abe41c0fe91e" ID-ISSN="1179-3163" ID-Zenodo-Dep="8080860" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687759488761" checkinUser="plazi" docAuthor="Fischer, Eberhard &amp; Lobin, Wolfram" docDate="2023" docId="03C1726083427758D9A8FBDA4E1EFEDA" docLanguage="en" docName="phytotaxa.600.3.4.pdf" docOrigin="Phytotaxa 600 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.600.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Diplazium cyamudongoense Eb. Fisch. &amp; Lobin 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="175" masterDocId="FFF80A188343775ED920FFCA4C3CFF92" masterDocTitle="Synoptic Revision of Diplazium (Athyriaceae) in Rwanda with description of Diplazium cyamudongoense sp. nov." masterLastPageNumber="183" masterPageNumber="169" pageNumber="170" updateTime="1699530258385" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FA1B0050262A5589E598808CA9BDE7F9">Synoptic Revision of Diplazium (Athyriaceae) in Rwanda with description of Diplazium cyamudongoense sp. nov.</mods:title>
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<mods:role id="81564ABE9126E85152E38C370C3A81E7">
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<mods:namePart id="A5FFD5C5A66DDD2FC757C4695580FF6F">Fischer, Eberhard</mods:namePart>
<mods:nameIdentifier id="BA1D659C488BAC6A5140E0C6CCA5ACBB" type="ORCID">0000-0001-7455-9833</mods:nameIdentifier>
<mods:affiliation id="9060475F6E02DD11D1E66FC9E448F81C">Institute for Integrated Natural Sciences-Biology, University of Koblenz, Universitätsstrasse 1, 56070 Koblenz, Germany. efischer @ uni-koblenz. de; https: // orcid. org / 0000 - 0001 - 7455 - 9833</mods:affiliation>
<mods:nameIdentifier id="EF6C5362A5280518738187F25E809388" type="email">efischer@uni-koblenz.de</mods:nameIdentifier>
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<mods:namePart id="94B84FC453F6642C0467D86A37727C5E">Lobin, Wolfram</mods:namePart>
<mods:affiliation id="CC0864FA582218105F58EF32A67FD920">Botanical Gardens Bonn University, Meckenheimer Allee 171, D- 51115 Bonn, Germany.</mods:affiliation>
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<mods:title id="6FF0383D25BC4E180EBE253E24A01648">Phytotaxa</mods:title>
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<mods:part id="195AA13DD376A3B3FAE3DBB146FC9170">
<mods:date id="2CEEC89983B6F9BA3D7DE4D27A868D76">2023</mods:date>
<mods:detail id="AE0D107AF0F10A4409D05694D0D7ED75" type="pubDate">
<mods:number id="ADCF6F57A04EDBD96CC68872B4650E01">2023-06-26</mods:number>
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<mods:number id="A5C34C2527AE2C30C306A6B6D1861033">600</mods:number>
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<mods:number id="F516EC2C9FF36C478EE15A4B625B22B3">3</mods:number>
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<mods:url id="600694416F5569A611EBEFB383F92D51">http://dx.doi.org/10.11646/phytotaxa.600.3.4</mods:url>
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<mods:identifier id="C5DB204BDE1C0143F71C2F7FC5C791DD" type="DOI">10.11646/phytotaxa.600.3.4</mods:identifier>
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<treatment id="03C1726083427758D9A8FBDA4E1EFEDA" ID-DOI="http://doi.org/10.5281/zenodo.8083541" ID-GBIF-Taxon="210477575" ID-Zenodo-Dep="8083541" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA" httpUri="http://treatment.plazi.org/id/03C1726083427758D9A8FBDA4E1EFEDA" lastPageId="6" lastPageNumber="175" pageId="1" pageNumber="170">
<subSubSection id="C37290FD8342775FD9A8FBDA4804FBB8" box="[136,1080,1040,1067]" pageId="1" pageNumber="170" type="nomenclature">
<paragraph id="8BD7C3768342775FD9A8FBDA4804FBB8" blockId="1.[136,1080,1040,1067]" box="[136,1080,1040,1067]" pageId="1" pageNumber="170">
<heading id="D09F741A8342775FD9A8FBDA4804FBB8" box="[136,1080,1040,1067]" fontSize="11" level="2" pageId="1" pageNumber="170" reason="3">
<taxonomicName id="4C68B8F58342775FD9A8FBDA4E9AFBB8" ID-CoL="BLS58" authority="Eb.Fisch. &amp; Lobin" authorityName="Eb. Fisch. &amp; Lobin" authorityYear="2023" box="[136,678,1040,1067]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">
<emphasis id="B91C1F648342775FD9A8FBDA4DF2FBB9" bold="true" box="[136,462,1040,1067]" italics="true" pageId="1" pageNumber="170">Diplazium cyamudongoense</emphasis>
Eb.Fisch. &amp; Lobin
</taxonomicName>
<emphasis id="B91C1F648342775FDB8CFBDB4F3DFBB8" box="[684,769,1041,1066]" italics="true" pageId="1" pageNumber="170">
<taxonomicNameLabel id="A22FA21F8342775FDB8CFBDB4F3DFBB8" box="[684,769,1041,1066]" pageId="1" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1353DFF38342775FDA30FBDA4F47FBB8" box="[784,891,1040,1066]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,229,1953,1975]" captionStartId-1="3.[136,229,1953,1975]" captionStartId-2="4.[136,229,1953,1975]" captionTargetBox-0="[201,1386,187,1923]" captionTargetBox-1="[198,1387,192,1926]" captionTargetBox-2="[203,1385,196,1925]" captionTargetId-0="figure-27@2.[199,1388,187,1925]" captionTargetId-1="figure-23@3.[198,1390,190,1928]" captionTargetId-2="figure-27@4.[200,1388,190,1928]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Diplazium cyamudongoense Eb.Fisch. &amp; Lobin sp. nov.—A, B. Fronds.—C, D. Pinnules. Photographs: E.Fischer, Cyamudongo. Scale bars: A, B. 10 cm; C, D. 1 cm." captionText-1="FIGURE 2. Diplazium cyamudongoense Eb.Fisch.&amp; Lobin sp. nov.—A, B. Detail of fronds.—C.Base of stipes with scales.—D. Pinnules. Photographs: E.Fischer, Cyamudongo. Scale bars: A. 5 cm; B, D. 2 cm; C. 1 cm." captionText-2="FIGURE 3. Diplazium cyamudongoense Eb.Fisch. &amp; Lobin sp. nov.—A, B. Base of stipes with scales.—C–E. Fragments of brittle scales.—F, G. Detail of pinnules showing sori. Photographs: E.Fischer &amp; W.Lobin, Cyamudongo. Scale bars; A, B, G. 1 mm; C, D, E. 2 mm; F. 3 mm; G. 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8080862" figureDoi-1="http://doi.org/10.5281/zenodo.8080868" figureDoi-2="http://doi.org/10.5281/zenodo.8080872" httpUri-0="https://zenodo.org/record/8080862/files/figure.png" httpUri-1="https://zenodo.org/record/8080868/files/figure.png" httpUri-2="https://zenodo.org/record/8080872/files/figure.png" pageId="1" pageNumber="170">Figs. 1–3</figureCitation>
,
<figureCitation id="1353DFF38342775FDAA7FBDA4FEDFBB9" box="[903,977,1040,1067]" captionStart="FIGURE 4" captionStartId="5.[136,229,1733,1755]" captionTargetBox="[151,1426,190,1702]" captionTargetId="figure-23@5.[151,1435,190,1709]" captionTargetPageId="5" captionText="FIGURE 4. Diplazium cyamudongoense Eb.Fisch. &amp; Lobin sp. nov.—A–D. Spores.— Diplazium humbertii —E–F. Spores. Photographs: E.Fischer, W.Lobin &amp; Y.Malkowsky, A–D. Cyamudongo; E–F. Cyamudongo. Scale bars: A, B, C, E, F. 10 µm; D. 8 µm." figureDoi="http://doi.org/10.5281/zenodo.8080874" httpUri="https://zenodo.org/record/8080874/files/figure.png" pageId="1" pageNumber="170">4 A–D</figureCitation>
,
<figureCitation id="1353DFF38342775FDAFFFBDA4817FBB8" box="[991,1067,1040,1067]" captionStart="FIGURE 5" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[201,1389,190,1926]" captionTargetId="figure-23@7.[199,1389,190,1928]" captionTargetPageId="7" captionText="FIGURE 5. Diplazium nemorale.—A. Pinnules.—Diplazium cyamudongoense Eb.Fisch.&amp;Lobin sp.nov.—B–C.Pinnules.Photographs: E.Fischer, A. Pócs &amp; Chambuko 6223; B–C. E.Fischer s.n. Scale bars: A, C. 3 mm; B. 5 mm.A reproduced with permission from BR." figureDoi="http://doi.org/10.5281/zenodo.8080876" httpUri="https://zenodo.org/record/8080876/files/figure.png" pageId="1" pageNumber="170">5 B–C</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8342775FD9A8FB964F17FB2B" pageId="1" pageNumber="170" type="materials_examined">
<paragraph id="8BD7C3768342775FD9A8FB964F17FB2B" blockId="1.[136,1452,1116,1210]" pageId="1" pageNumber="170">
<emphasis id="B91C1F648342775FD9A8FB964CFEFBE3" bold="true" box="[136,194,1116,1137]" pageId="1" pageNumber="170">Type:</emphasis>
—
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<collectingCountry id="F37F83E68342775FD9FBFB964D7BFBE3" box="[219,327,1116,1137]" name="Rwanda" pageId="1" pageNumber="170">RWANDA</collectingCountry>
.
<collectingRegion id="49AC0D948342775FD874FB964E3EFBE0" box="[340,514,1116,1138]" pageId="1" pageNumber="170">Western Province</collectingRegion>
:
<location id="8EB795AD8342775FDB2CFB964EADFBE0" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8342775FDB2CFB964EADFBE0" box="[524,657,1116,1138]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.5408027" longLatPrecision="1" longitude="28.989017" municipality="Nyungwe National Park" name="Lacs Edouard" pageId="1" pageNumber="170" stateProvince="Western Province">Lacs Edouard</location>
et
<location id="8EB795AD8342775FDB91FB964EDFFBE0" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8342775FDB91FB964EDFFBE0" box="[689,739,1116,1138]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.5408027" longLatPrecision="1" longitude="28.989017" municipality="Nyungwe National Park" name="Kivu" pageId="1" pageNumber="170" stateProvince="Western Province">Kivu</location>
,
<location id="8EB795AD8342775FDBCEFB964FE6FBE0" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8342775FDBCEFB964FE6FBE0" box="[750,986,1116,1138]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.5408027" longLatPrecision="1" longitude="28.989017" municipality="Nyungwe National Park" name="Nyungwe National Park" pageId="1" pageNumber="170" stateProvince="Western Province">Nyungwe National Park</location>
,
<location id="8EB795AD8342775FDAC5FB964897FBE3" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8342775FDAC5FB964897FBE3" box="[997,1195,1116,1138]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.5408027" longLatPrecision="1" longitude="28.989017" municipality="Nyungwe National Park" name="Cyamudongo Forest" pageId="1" pageNumber="170" stateProvince="Western Province">Cyamudongo Forest</location>
, forest floor along stream,
<geoCoordinate id="EE5CA5B18342775FD99DFB4A4D7AFB04" box="[189,326,1152,1174]" degrees="2" direction="south" minutes="32" orientation="latitude" pageId="1" pageNumber="170" precision="1" seconds="26.89" value="-2.5408027">S2°32`26.89``</geoCoordinate>
<geoCoordinate id="EE5CA5B18342775FD86AFB4A4DDFFB04" box="[330,483,1152,1174]" degrees="28" direction="east" minutes="59" orientation="longitude" pageId="1" pageNumber="170" precision="1" seconds="20.46" value="28.989017">E28°59`20.46``</geoCoordinate>
,
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,
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,
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<collectorName id="269DA6A08342775FDBDCFB4A4F61FB07" box="[764,861,1152,1173]" pageId="1" pageNumber="170">E. Fischer</collectorName>
s.n.
</emphasis>
(
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!;
</materialsCitation>
<materialsCitation id="3B00C92B8342775FDDE7FB4A4F1AFB2B" ID-GBIF-Occurrence="4135850304" collectingDate="2021-09-18" collectionCode="KOBL" collectorName="E. Fischer" country="Rwanda" county="Cyamudongo Forest" elevation="1941" latitude="-2.5408027" location="Lacs Edouard" longLatPrecision="1" longitude="28.989017" municipality="Nyungwe National Park" pageId="1" pageNumber="170" specimenCode="B 20 0220597!, B 20 0220598!, B 20 0220599!" specimenCount="2" stateProvince="Western Province" typeStatus="isotype">
<typeStatus id="54D37DD48342775FDDE7FB4A492AFB04" box="[1223,1302,1152,1174]" pageId="1" pageNumber="170" type="isotype">isotypes</typeStatus>
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,
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,
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[frond on three sheets],
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!)
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.
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8342775FD9A8FB224834FA24" pageId="1" pageNumber="170" type="diagnosis">
<paragraph id="8BD7C3768342775FD9A8FB224834FA24" blockId="1.[136,1452,1256,2002]" pageId="1" pageNumber="170">
<emphasis id="B91C1F648342775FD9A8FB224D39FA90" bold="true" box="[136,261,1256,1282]" pageId="1" pageNumber="170">Diagnosis:</emphasis>
—The new species is morphologically similar to
<taxonomicName id="4C68B8F58342775FDA38FB224FC7FA90" authorityName="Schelpe" authorityYear="1967" baseAuthorityName="Baker" baseAuthorityYear="1876" box="[792,1019,1256,1282]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="nemorale">
<emphasis id="B91C1F648342775FDA38FB224FC7FA90" box="[792,1019,1256,1282]" italics="true" pageId="1" pageNumber="170">Diplazium nemorale</emphasis>
</taxonomicName>
(
<figureCitation id="1353DFF38342775FDD27FB224861FA90" box="[1031,1117,1256,1283]" captionStart="FIGURE 5" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[201,1389,190,1926]" captionTargetId="figure-23@7.[199,1389,190,1928]" captionTargetPageId="7" captionText="FIGURE 5. Diplazium nemorale.—A. Pinnules.—Diplazium cyamudongoense Eb.Fisch.&amp;Lobin sp.nov.—B–C.Pinnules.Photographs: E.Fischer, A. Pócs &amp; Chambuko 6223; B–C. E.Fischer s.n. Scale bars: A, C. 3 mm; B. 5 mm.A reproduced with permission from BR." figureDoi="http://doi.org/10.5281/zenodo.8080876" httpUri="https://zenodo.org/record/8080876/files/figure.png" pageId="1" pageNumber="170">Fig.5A</figureCitation>
).
<emphasis id="B91C1F648342775FDD52FB224CDCFAB4" italics="true" pageId="1" pageNumber="170">
<taxonomicName id="4C68B8F58342775FDD52FB224990FA90" authority="Fischer &amp; Lobin, 2023" authorityName="Fischer &amp; Lobin" authorityYear="2023" box="[1138,1452,1256,1282]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">Diplazium cyamudongoense</taxonomicName>
<taxonomicNameLabel id="A22FA21F8342775FD9A8FAC74CDCFAB4" box="[136,224,1293,1318]" pageId="1" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has deeply incised to almost pinnate pinnules, all pinnae up to the terminal pinna are pinnatifid, there are 6–8(11) sori per pinnule lobe that are elongate,
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long, and the base of stipe bears numerous large scales, which are more than
<quantity id="4C906E938342775FD85BFA9E4DF7FAFD" box="[379,459,1364,1391]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="170" unit="mm" value="18.0">18 mm</quantity>
long. In
<taxonomicName id="4C68B8F58342775FDB14FA9F4EFEFAFC" authorityName="Schelpe" authorityYear="1967" baseAuthorityName="Baker" baseAuthorityYear="1876" box="[564,706,1364,1390]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="nemorale">
<emphasis id="B91C1F648342775FDB14FA9F4EFEFAFC" box="[564,706,1364,1390]" italics="true" pageId="1" pageNumber="170">D. nemorale</emphasis>
</taxonomicName>
the pinnules are shallowly to deeply cut, but the lamina does not appear tripinnate and is almost undivided towards pinna apex and on upper pinnae, the sori are 1–6 per pinnule lobe and 3–4(5) mm long, the scales at stipes base are not exceeding
<quantity id="4C906E938342775FDA76FA564FABFA25" box="[854,919,1436,1463]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="170" unit="mm" value="9.0">9 mm</quantity>
in length.
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8342775FD99DFA0A4ECFF840" pageId="1" pageNumber="170" type="description">
<paragraph id="8BD7C3768342775FD99DFA0A4ECFF840" blockId="1.[136,1452,1256,2002]" pageId="1" pageNumber="170">
<emphasis id="B91C1F648342775FD99DFA0A4D6DFA48" bold="true" box="[189,337,1472,1498]" pageId="1" pageNumber="170">Description:</emphasis>
—Terrestrial fern.
<emphasis id="B91C1F648342775FDB04FA0A4EB8FA48" box="[548,644,1472,1498]" italics="true" pageId="1" pageNumber="170">Rhizome</emphasis>
erect, scaly; scales very large, dark brown, of
<specimenCount id="9D6E08FF8342775FDDB7FA0B493BFA49" box="[1175,1287,1473,1499]" pageId="1" pageNumber="170" type="generic" typeStatus="types">two types</specimenCount>
, long filiform, extremely fragile, thus almost all broken, larger scales with broad base, more than
<quantity id="4C906E938342775FDD0FFA2E489EFA6D" box="[1071,1186,1508,1535]" metricMagnitude="-2" metricUnit="m" metricValue="1.8379999999999999" pageId="1" pageNumber="170" unit="mm" value="18.38">18.38 mm</quantity>
long, at base
<quantity id="4C906E938342775FDC60FA2E499BFA6D" box="[1344,1447,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="170" unit="mm" value="1.95">1.95 mm</quantity>
, further upwards
<quantity id="4C906E938342775FD863F9C24D95F9B1" box="[323,425,1544,1571]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="170" unit="mm" value="0.97">0.97 mm</quantity>
, at the tip
<quantity id="4C906E938342775FDB01F9C24EBAF9B1" box="[545,646,1544,1571]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="170" unit="mm" value="0.54">0.54 mm</quantity>
wide, smaller scales linear, more than
<quantity id="4C906E938342775FDD1EF9C2489FF9B1" box="[1086,1187,1544,1571]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="1" pageNumber="170" unit="mm" value="15.3">15.3 mm</quantity>
long, at base
<quantity id="4C906E938342775FDC60F9C2499BF9B1" box="[1344,1447,1544,1571]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="170" unit="mm" value="0.74">0.74 mm</quantity>
, further upwards
<quantity id="4C906E938342775FD861F9E64D98F9D5" box="[321,420,1580,1607]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="170" unit="mm" value="0.38">0.38 mm</quantity>
and at tip
<quantity id="4C906E938342775FDB39F9E64E41F9D5" box="[537,637,1580,1607]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="170" unit="mm" value="0.08">0.08 mm</quantity>
wide.
<emphasis id="B91C1F648342775FDBE8F9E64F24F9D4" box="[712,792,1580,1606]" italics="true" pageId="1" pageNumber="170">Fronds</emphasis>
tufted.
<emphasis id="B91C1F648342775FDA4DF9E64F8CF9D4" box="[877,944,1580,1606]" italics="true" pageId="1" pageNumber="170">Stipes</emphasis>
light green, at base dark brown, canaliculate, 40–47(–108) ×
<quantity id="4C906E938342775FD814F99A4DBEF9F9" box="[308,386,1616,1643]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="170" unit="cm" value="0.7">0.7 cm</quantity>
, base with scales similar to those from rhizome.
<emphasis id="B91C1F648342775FDAB9F99B4FD3F9F8" box="[921,1007,1617,1642]" italics="true" pageId="1" pageNumber="170">Lamina</emphasis>
59–69(–105) ×
<quantity id="4C906E938342775FDD81F99A491AF9F9" box="[1185,1318,1616,1643]" metricMagnitude="-1" metricUnit="m" metricValue="5.575" metricValueMax="6.4" metricValueMin="4.75" pageId="1" pageNumber="170" unit="cm" value="55.75" valueMax="64.0" valueMin="47.5">47.5–64 cm</quantity>
, triangular– ovate, green, shiny on upper surface, 2–pinnate–pinnatifid to nearly 3–pinnate.
<taxonomicName id="4C68B8F58342775FDD3CF9BE4854F91C" authorityName="Albers" authorityYear="1850" box="[1052,1128,1652,1678]" class="Gastropoda" family="Cerastidae" genus="Rachis" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="170" phylum="Mollusca" rank="genus">
<emphasis id="B91C1F648342775FDD3CF9BE4854F91C" box="[1052,1128,1652,1678]" italics="true" pageId="1" pageNumber="170">Rachis</emphasis>
</taxonomicName>
greenish–brown to middle– brown, with few scales.
<emphasis id="B91C1F648342775FD8B5F9534DD8F920" box="[405,484,1689,1714]" italics="true" pageId="1" pageNumber="170">Pinnae</emphasis>
in ca. 15–18 pairs, alternate, arranged at an angle of ca. 45°, (15.3–)23.7–34 ×
<quantity id="4C906E938342775FDC70F9524C92F945" metricMagnitude="-1" metricUnit="m" metricValue="1.575" metricValueMax="1.7" metricValueMin="1.45" pageId="1" pageNumber="170" unit="cm" value="15.75" valueMax="17.0" valueMin="14.5">14.5–17 cm</quantity>
; pinnae rachis light–brown to greenish, base often darker, with few scales, sometimes minutely hairy.
<emphasis id="B91C1F648342775FDC0CF97649B2F944" box="[1324,1422,1724,1750]" italics="true" pageId="1" pageNumber="170">Pinnules</emphasis>
in 12–14(–19) pairs in each pinna, opposite, (4.5–)5.1–6.6(–9.2) × 1.6–2.3(–4.5) cm, oblong–lanceolate.
<emphasis id="B91C1F648342775FDC03F92A4CF9F88C" italics="true" pageId="1" pageNumber="170">
Pinnules 3
<superScript id="7C1D6E3E8342775FDCBDF92A4997F97C" attach="none" box="[1437,1451,1760,1774]" fontSize="6" pageId="1" pageNumber="170">rd</superScript>
order
</emphasis>
in 6–10(–12) pairs tapering into the caudate apex, opposite, 0.9–1.1 ×
<quantity id="4C906E938342775FDAD7F8CE4845F88D" box="[1015,1145,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="170" unit="cm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 cm</quantity>
, decurrent, deeply incised; racheolule mostly winged.
<emphasis id="B91C1F648342775FD89AF8E24DD5F8D0" box="[442,489,1832,1858]" italics="true" pageId="1" pageNumber="170">Sori</emphasis>
elongate, 1.98–2.95 ×
<quantity id="4C906E938342775FDBCEF8E24FA8F8D1" box="[750,916,1832,1859]" metricMagnitude="-4" metricUnit="m" metricValue="8.45" metricValueMax="9.8" metricValueMin="7.1" pageId="1" pageNumber="170" unit="mm" value="0.845" valueMax="0.98" valueMin="0.71">0.71–0.98 mm</quantity>
, in basic pinnule lobes up to 6 sori, in all other sori usually in two rows along midrib, 6–8(11), paraphyses present, 237–350 µm long, sometimes with short branches.
<emphasis id="B91C1F648342775FD9A8F8BA4CCCF818" box="[136,240,1904,1930]" italics="true" pageId="1" pageNumber="170">Indusium</emphasis>
elongate, pale, glabrous, same size as sori attached to one side of sorus (asplenioid).
<emphasis id="B91C1F648342775FDD91F8BA48DCF818" box="[1201,1248,1904,1930]" italics="true" pageId="1" pageNumber="170">Sori</emphasis>
6–11 per pinnule.
<emphasis id="B91C1F648342775FD9A8F85E4CEFF83C" box="[136,211,1940,1966]" italics="true" pageId="1" pageNumber="170">Spores</emphasis>
with alate perispore, 42–50.5 × 25.7–31.46 µm, alae curved and narrow, 4.29–7.15 µm high and 1.14–1.43 µm wide, the remaining perispore smooth with small folds.
</paragraph>
</subSubSection>
<caption id="DF1793FE8341775CD9A8F86B4E48F848" ID-DOI="http://doi.org/10.5281/zenodo.8080862" ID-Zenodo-Dep="8080862" httpUri="https://zenodo.org/record/8080862/files/figure.png" pageId="2" pageNumber="171" startId="2.[136,229,1953,1975]" targetBox="[201,1386,187,1923]" targetPageId="2" targetType="figure">
<paragraph id="8BD7C3768341775CD9A8F86B4E48F848" blockId="2.[136,1452,1953,2011]" pageId="2" pageNumber="171">
<emphasis id="B91C1F648341775CD9A8F86B4D39F824" bold="true" box="[136,261,1953,1975]" pageId="2" pageNumber="171">FIGURE 1.</emphasis>
<taxonomicName id="4C68B8F58341775CD831F86B4EC9F825" authority="Eb.Fisch. &amp; Lobin" authorityName="Eb. Fisch. &amp; Lobin" authorityYear="2023" box="[273,757,1953,1975]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">
<emphasis id="B91C1F648341775CD831F86B4E1BF824" box="[273,551,1953,1974]" italics="true" pageId="2" pageNumber="171">Diplazium cyamudongoense</emphasis>
Eb.Fisch. &amp; Lobin
</taxonomicName>
<taxonomicNameLabel id="A22FA21F8341775CDA22F8684F6DF824" box="[770,849,1953,1975]" pageId="2" pageNumber="171" rank="species">
<emphasis id="B91C1F648341775CDA22F8684F71F825" box="[770,845,1954,1975]" italics="true" pageId="2" pageNumber="171">sp. nov</emphasis>
.
</taxonomicNameLabel>
—A, B. Fronds.—C, D. Pinnules. Photographs: E.Fischer, Cyamudongo. Scale bars: A, B. 10 cm; C, D. 1 cm.
</paragraph>
</caption>
<caption id="DF1793FE8340775DD9A8F86B4FB3F848" ID-DOI="http://doi.org/10.5281/zenodo.8080868" ID-Zenodo-Dep="8080868" httpUri="https://zenodo.org/record/8080868/files/figure.png" pageId="3" pageNumber="172" startId="3.[136,229,1953,1975]" targetBox="[198,1387,192,1926]" targetPageId="3" targetType="figure">
<paragraph id="8BD7C3768340775DD9A8F86B4FB3F848" blockId="3.[136,1453,1953,2011]" pageId="3" pageNumber="172">
<emphasis id="B91C1F648340775DD9A8F86B4CC0F824" bold="true" box="[136,252,1953,1975]" pageId="3" pageNumber="172">FIGURE 2.</emphasis>
<taxonomicName id="4C68B8F58340775DD820F86B4EFDF825" authority="Eb.Fisch. &amp; Lobin" authorityName="Eb. Fisch. &amp; Lobin" authorityYear="2023" box="[256,705,1953,1975]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">
<emphasis id="B91C1F648340775DD820F86B4E32F824" box="[256,526,1953,1974]" italics="true" pageId="3" pageNumber="172">Diplazium cyamudongoense</emphasis>
Eb.Fisch. &amp; Lobin
</taxonomicName>
<taxonomicNameLabel id="A22FA21F8340775DDBE6F8684F31F824" box="[710,781,1953,1975]" pageId="3" pageNumber="172" rank="species">
<emphasis id="B91C1F648340775DDBE6F8684F35F825" box="[710,777,1954,1975]" italics="true" pageId="3" pageNumber="172">sp. nov</emphasis>
.
</taxonomicNameLabel>
—A, B. Detail of fronds.—C. Base of stipes with scales.—D. Pinnules. Photographs: E.Fischer, Cyamudongo. Scale bars: A. 5 cm; B, D. 2 cm; C. 1 cm.
</paragraph>
</caption>
<caption id="DF1793FE8347775AD9A8F86B4D4DF86C" ID-DOI="http://doi.org/10.5281/zenodo.8080872" ID-Zenodo-Dep="8080872" httpUri="https://zenodo.org/record/8080872/files/figure.png" pageId="4" pageNumber="173" startId="4.[136,229,1953,1975]" targetBox="[203,1385,196,1925]" targetPageId="4" targetType="figure">
<paragraph id="8BD7C3768347775AD9A8F86B4D4DF86C" blockId="4.[136,1451,1953,2046]" pageId="4" pageNumber="173">
<emphasis id="B91C1F648347775AD9A8F86B4D3DF824" bold="true" box="[136,257,1953,1975]" pageId="4" pageNumber="173">FIGURE 3.</emphasis>
<taxonomicName id="4C68B8F58347775AD82AF86B4EE3F825" authority="Eb.Fisch. &amp; Lobin" authorityName="Eb. Fisch. &amp; Lobin" authorityYear="2023" box="[266,735,1953,1975]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">
<emphasis id="B91C1F648347775AD82AF86B4E20F824" box="[266,540,1953,1974]" italics="true" pageId="4" pageNumber="173">Diplazium cyamudongoense</emphasis>
Eb.Fisch. &amp; Lobin
</taxonomicName>
<emphasis id="B91C1F648347775ADBC8F8684F09F825" box="[744,821,1954,1975]" italics="true" pageId="4" pageNumber="173">
<taxonomicNameLabel id="A22FA21F8347775ADBC8F8684F09F825" box="[744,821,1954,1975]" pageId="4" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—A, B. Base of stipes with scales.—C–E. Fragments of brittle scales.—F, G. Detail of pinnules showing sori. Photographs: E.Fischer &amp; W.Lobin, Cyamudongo. Scale bars; A, B, G. 1 mm; C, D, E. 2 mm; F. 3 mm; G. 1 mm.
</paragraph>
</caption>
<caption id="DF1793FE8346775BD9A8F90F4931F96D" ID-DOI="http://doi.org/10.5281/zenodo.8080874" ID-Zenodo-Dep="8080874" httpUri="https://zenodo.org/record/8080874/files/figure.png" pageId="5" pageNumber="174" startId="5.[136,229,1733,1755]" targetBox="[151,1426,190,1702]" targetPageId="5" targetType="figure">
<paragraph id="8BD7C3768346775BD9A8F90F4931F96D" blockId="5.[136,1453,1733,1791]" pageId="5" pageNumber="174">
<emphasis id="B91C1F648346775BD9A8F90F4CC0F948" bold="true" box="[136,252,1733,1755]" pageId="5" pageNumber="174">FIGURE 4.</emphasis>
<taxonomicName id="4C68B8F58346775BD825F90F4EFBF949" authority="Eb.Fisch. &amp; Lobin" authorityName="Eb. Fisch. &amp; Lobin" authorityYear="2023" box="[261,711,1733,1755]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="174" phylum="Tracheophyta" rank="species" species="cyamudongoense" status="sp. nov.">
<emphasis id="B91C1F648346775BD825F90F4E2FF948" box="[261,531,1733,1754]" italics="true" pageId="5" pageNumber="174">Diplazium cyamudongoense</emphasis>
Eb.Fisch. &amp; Lobin
</taxonomicName>
<taxonomicNameLabel id="A22FA21F8346775BDBECF90C4F2EF949" box="[716,786,1734,1755]" pageId="5" pageNumber="174" rank="species">
<emphasis id="B91C1F648346775BDBECF90C4F33F949" box="[716,783,1734,1755]" italics="true" pageId="5" pageNumber="174">sp. nov</emphasis>
.
</taxonomicNameLabel>
—A–D. Spores. —
<taxonomicName id="4C68B8F58346775BDAEEF90F48AFF948" authorityName="Pichi Sermolli" authorityYear="1973" baseAuthorityName="Christensen" baseAuthorityYear="1937" box="[974,1171,1733,1754]" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="174" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis id="B91C1F648346775BDAEEF90F48AFF948" box="[974,1171,1733,1754]" italics="true" pageId="5" pageNumber="174">Diplazium humbertii</emphasis>
</taxonomicName>
—E–F. Spores. Photographs: E.Fischer, W.Lobin &amp; Y.Malkowsky, A–D. Cyamudongo; E–F. Cyamudongo. Scale bars: A, B, C, E, F. 10 µm; D. 8 µm.
</paragraph>
</caption>
<subSubSection id="C37290FD8346775BD99DF8E74F6CF802" pageId="5" pageNumber="174" type="materials_examined">
<paragraph id="8BD7C3768346775BD99DF8E74F6CF802" blockId="5.[136,1451,1837,2080]" pageId="5" pageNumber="174">
<materialsCitation id="3B00C92B8346775BD99DF8E74F6CF802" ID-GBIF-Occurrence="4135850302" collectingDate="2022-03-16" collectionCode="KOBL" collectorName="E. Fischer &amp; P. Ballings &amp; B. Wursten" country="Rwanda" county="Cyamudongo Forest" elevation="1877" latitude="-2.554839" location="Lacs Edouard" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" pageId="5" pageNumber="174" specimenCount="1" stateProvince="Western Province" typeStatus="paratype">
<emphasis id="B91C1F648346775BD99DF8E74EB3F8D5" bold="true" box="[189,655,1837,1864]" pageId="5" pageNumber="174">
Additional specimens seen (
<typeStatus id="54D37DD48346775BDB37F8E74EBBF8D5" box="[535,647,1837,1863]" pageId="5" pageNumber="174" type="paratype">Paratype</typeStatus>
)
</emphasis>
:—
<collectingCountry id="F37F83E68346775BDB93F8E44F0DF8DA" box="[691,817,1838,1864]" name="Rwanda" pageId="5" pageNumber="174">RWANDA</collectingCountry>
.
<collectingRegion id="49AC0D948346775BDA65F8E44829F8D5" box="[837,1045,1837,1864]" pageId="5" pageNumber="174">Western Province</collectingRegion>
:
<location id="8EB795AD8346775BDD05F8E448F9F8D5" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BDD05F8E448F9F8D5" box="[1061,1221,1837,1864]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Lacs Edouard" pageId="5" pageNumber="174" stateProvince="Western Province">Lacs Edouard</location>
et
<location id="8EB795AD8346775BDDD3F8E74912F8D5" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BDDD3F8E74912F8D5" box="[1267,1326,1837,1863]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Kivu" pageId="5" pageNumber="174" stateProvince="Western Province">Kivu</location>
,
<location id="8EB795AD8346775BDC1EF8E44D19F8F9" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BDC1EF8E44D19F8F9" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Nyungwe National Park" pageId="5" pageNumber="174" stateProvince="Western Province">Nyungwe National Park</location>
,
<location id="8EB795AD8346775BD80FF89B4E29F8FE" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BD80FF89B4E29F8FE" box="[303,533,1873,1900]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Cyamudongo Forest" pageId="5" pageNumber="174" stateProvince="Western Province">Cyamudongo Forest</location>
, forest floor along stream
<location id="8EB795AD8346775BDA17F89B4FFDF8F9" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BDA17F89B4FFDF8F9" box="[823,961,1873,1899]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Nyamabuye" pageId="5" pageNumber="174" stateProvince="Western Province">Nyamabuye</location>
,
<quantity id="4C906E938346775BDAECF898481EF8FE" box="[972,1058,1874,1900]" metricMagnitude="3" metricUnit="m" metricValue="1.877" pageId="5" pageNumber="174" unit="m" value="1877.0">
<elevation id="004524458346775BDAECF898481EF8FE" box="[972,1058,1874,1900]" metricMagnitude="3" metricUnit="m" metricValue="1.877" pageId="5" pageNumber="174" unit="m" value="1877.0">1877 m</elevation>
</quantity>
,
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<geoCoordinate id="EE5CA5B18346775BDDF3F89B49B9F8FE" box="[1235,1413,1873,1900]" degrees="28" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="174" precision="1" seconds="03.56" value="28.984322">E28°59`03.56``</geoCoordinate>
,
<date id="FFD6E5B68346775BDCAFF89B4D2FF802" pageId="5" pageNumber="174" value="2022-03-16">
<collectingDate id="EF921C5E8346775BDCAFF89B4D2FF802" pageId="5" pageNumber="174" value="2022-03-16">16 March 2022</collectingDate>
</date>
,
<emphasis id="B91C1F648346775BD83EF8BC4E2AF81D" box="[286,534,1909,1935]" italics="true" pageId="5" pageNumber="174">
<collectorName id="269DA6A08346775BD83EF8BC4DAEF81D" box="[286,402,1909,1935]" pageId="5" pageNumber="174">E. Fischer</collectorName>
,
<collectorName id="269DA6A08346775BD8BEF8BC4E2AF81D" box="[414,534,1909,1935]" pageId="5" pageNumber="174">P. Ballings</collectorName>
</emphasis>
&amp;
<emphasis id="B91C1F648346775BDB1AF8BC4EDFF81D" box="[570,739,1910,1935]" italics="true" pageId="5" pageNumber="174">
<collectorName id="269DA6A08346775BDB1AF8BC4E89F81D" box="[570,693,1910,1935]" pageId="5" pageNumber="174">B. Wursten</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED795BB38346775BDBD2F8BC4F7CF802" box="[754,832,1910,1936]" pageId="5" pageNumber="174">KOBL</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8346775BD99DF8534EEAF826" box="[189,726,1945,1972]" pageId="5" pageNumber="174" type="distribution">
<paragraph id="8BD7C3768346775BD99DF8534EEAF826" blockId="5.[136,1451,1837,2080]" box="[189,726,1945,1972]" pageId="5" pageNumber="174">
<emphasis id="B91C1F648346775BD99DF8534D65F821" bold="true" box="[189,345,1945,1971]" pageId="5" pageNumber="174">
<collectorName id="269DA6A08346775BD99DF8534D68F821" box="[189,340,1945,1971]" pageId="5" pageNumber="174">Distribution</collectorName>
:
</emphasis>
—
<collectorName id="269DA6A08346775BD855F8534DEDF821" box="[373,465,1945,1971]" pageId="5" pageNumber="174">Rwanda</collectorName>
(
<location id="8EB795AD8346775BD8FFF8534EF6F826" LSID="urn:lsid:plazi:treatment:03C1726083427758D9A8FBDA4E1EFEDA:8EB795AD8346775BD8FFF8534EF6F826" box="[479,714,1945,1972]" country="Rwanda" county="Cyamudongo Forest" latitude="-2.554839" longLatPrecision="1" longitude="28.984322" municipality="Montane forest" name="Cyamudongo Forest" pageId="5" pageNumber="174">Cyamudongo Forest</location>
).
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8346775BD99DF8774F49F84A" box="[189,885,1981,2008]" pageId="5" pageNumber="174" type="biology_ecology">
<paragraph id="8BD7C3768346775BD99DF8774F49F84A" blockId="5.[136,1451,1837,2080]" box="[189,885,1981,2008]" pageId="5" pageNumber="174">
<emphasis id="B91C1F648346775BD99DF8774D1FF845" bold="true" box="[189,291,1981,2007]" pageId="5" pageNumber="174">
<collectorName id="269DA6A08346775BD99DF8774D21F845" box="[189,285,1981,2007]" pageId="5" pageNumber="174">Habitat</collectorName>
:
</emphasis>
—
<collectingMunicipality id="6BB3590C8346775BD81FF8744DD5F845" box="[319,489,1981,2008]" pageId="5" pageNumber="174">Montane forest</collectingMunicipality>
along streams, elev.
<quantity id="4C906E938346775BDBF3F8744F4DF84A" box="[723,881,1982,2008]" metricMagnitude="3" metricUnit="m" metricValue="1.909" metricValueMax="1.941" metricValueMin="1.877" pageId="5" pageNumber="174" unit="m" value="1909.0" valueMax="1941.0" valueMin="1877.0">
<elevation id="004524458346775BDBF3F8744F4DF84A" box="[723,881,1982,2008]" metricMagnitude="3" metricUnit="m" metricValue="1.909" metricValueMax="1.941" metricValueMin="1.877" pageId="5" pageNumber="174" unit="m" value="1909.0" valueMax="1941.0" valueMin="1877.0">1877–1941 m</elevation>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37290FD8346775BD99DF828489BF7B2" pageId="5" pageNumber="174" type="etymology">
<paragraph id="8BD7C3768346775BD99DF828489BF7B2" blockId="5.[136,1451,1837,2080]" pageId="5" pageNumber="174">
<emphasis id="B91C1F648346775BD99DF8284D74F86E" bold="true" box="[189,328,2018,2044]" pageId="5" pageNumber="174">
<collectorName id="269DA6A08346775BD99DF8284D7EF86E" box="[189,322,2018,2044]" pageId="5" pageNumber="174">Etymology</collectorName>
:
</emphasis>
—
<collectorName id="269DA6A08346775BD844F82B4DB3F869" box="[356,399,2017,2043]" pageId="5" pageNumber="174">The</collectorName>
species name refers to the
<collectingCounty id="62B6BBFA8346775BDBE5F82B4F91F86E" box="[709,941,2017,2044]" pageId="5" pageNumber="174">Cyamudongo Forest</collectingCounty>
, a place of extreme high diversity in animals and plants.
<collectorName id="269DA6A08346775BD828F7CF4D7EF78D" box="[264,322,2053,2079]" pageId="5" pageNumber="174">Until</collectorName>
now, the new species is only known from two localities in this exciting area.
</paragraph>
</subSubSection>
<subSubSection id="C37290FD83457758D99DFF544E1EFEDA" pageId="6" pageNumber="175" type="conservation">
<paragraph id="8BD7C37683457758D99DFF544E1EFEDA" blockId="6.[136,1452,158,329]" pageId="6" pageNumber="175">
<emphasis id="B91C1F6483457758D99DFF544D8AFF2A" bold="true" box="[189,438,158,184]" pageId="6" pageNumber="175">Conservation status:</emphasis>
—With only two collecting sites, it is not possible to calculate an Extent of Occurrence (EOO).
<emphasis id="B91C1F6483457758D9C3FF084EB1FF42" bold="true" box="[227,653,194,220]" pageId="6" pageNumber="175">
EOO: not applicable; AOO:
<quantity id="4C906E9383457758DB1BFF084EB9FF4E" box="[571,645,194,220]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="175" unit="km" value="18.0">18 km</quantity>
<superScript id="7C1D6E3E83457758DBA5FF084EB1FF42" attach="left" box="[645,653,194,208]" fontSize="6" pageId="6" pageNumber="175">2</superScript>
</emphasis>
. The area of occupancy (AOO) falls within the limits for the Critically Endangered category. However, as the species occurs within a well-protected area, only in protected areas and therefore to experience no threat or decline, it cannot be assessed in any threat category; Therefore, it should be assessed as Least Concern (Protected area dependent).
</paragraph>
</subSubSection>
</treatment>
</document>
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