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<document id="92DE40F1BFEA504BBEA2A76BE1ABDCEC" ID-CLB-Dataset="21909" ID-DOI="10.5852/ejt.2020.650" ID-GBIF-Dataset="04bc4f07-1283-4058-9641-2fce5ee10aca" ID-ISSN="2118-9773" ID-Zenodo-Dep="3839797" ID-ZooBank="3EAB8173-2208-43AC-84E1-AFCE8B9376AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590155299496" checkinUser="valdenar" docAuthor="Phan, Quoc Toan &amp; Ngo, Quoc Phu" docDate="2020" docId="03BBEF6A2264CA73CA04FED262C4FC8B" docLanguage="en" docName="ejt-650_phan_ngo.pdf.imf" docOrigin="European Journal of Taxonomy 650" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Prodasineura kong Phan &amp; Ngo 2020, sp. nov." docType="treatment" docUuid="6BEEA763-7AFB-4426-B919-024387DEC48E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="17" masterDocId="FF8297122268CA63C871FFD86161FFDF" masterDocTitle="A revision of the systematics and distribution of the damselfly genus Prodasineura Cowley, 1934 (Odonata: Zygoptera: Platycnemididae) in Vietnam with description of two new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="13" updateTime="1698840923651" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0CA6C86FA1787FE8CB7E854D47947D5B">A revision of the systematics and distribution of the damselfly genus Prodasineura Cowley, 1934 (Odonata: Zygoptera: Platycnemididae) in Vietnam with description of two new species</mods:title>
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<mods:name id="A361F30CE35DF9C7F59D52B45836B04F" type="personal">
<mods:role id="D242CDE21092A6D2A5BFB9D96ED8FC36">
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<mods:namePart id="C5C7131F689068976BC0A79C7C9E6776">Phan, Quoc Toan</mods:namePart>
<mods:nameIdentifier id="6B5E1852886FFD8F1689617101AD7E65" type="LSID">9DBB241C-A070-45DF-A111-B7BDDB56EEEE</mods:nameIdentifier>
<mods:affiliation id="D5E9B22E3CBD6D58215BB537C4880C20">The Center for Entomology &amp; Parasitology Researches, Institute of Researches and Training of Medicine, Biology &amp; Pharmacy, Duy Tan University, 3 Quang Trung, Da Nang, Vietnam. &amp; urn: lsid: zoobank. org: author: 9 DBB 241 C-A 070 - 45 DF-A 111 - B 7 BDDB 56 EEEE &amp; Corresponding author: pqtoan 84 @ gmail. com; phanquoctoan 1 @ duytan. edu. vn</mods:affiliation>
<mods:nameIdentifier id="12F7A973D9A0B321B7DDB8326CB68BFC" type="email">pqtoan84@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E857DCEA2E8C090989D25B85B114B304">Ngo, Quoc Phu</mods:namePart>
<mods:nameIdentifier id="903CAA1FE7BDFE21D278E28CB74A52F4" type="LSID">34581D77-1365-4D24-A1BF-EB791D08ABD5</mods:nameIdentifier>
<mods:affiliation id="471F17DA816F3CD43E402463FB9D586C">The Center for Entomology &amp; Parasitology Researches, Institute of Researches and Training of Medicine, Biology &amp; Pharmacy, Duy Tan University, 3 Quang Trung, Da Nang, Vietnam. &amp; Email: ngoquocphu @ gmail. com; ngoquocphu 1 @ duytan. edu. vn &amp; urn: lsid: zoobank. org: author: 34581 D 77 - 1365 - 4 D 24 - A 1 BF-EB 791 D 08 ABD 5</mods:affiliation>
<mods:nameIdentifier id="ECD1A965277EBB7B60C7B65AF0CF3745" type="email">ngoquocphu@gmail.com</mods:nameIdentifier>
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<mods:title id="B4823BB1F71A39A6519DBE42078CE7C6">European Journal of Taxonomy</mods:title>
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<mods:date id="A721721BDFB7D031005587247FE17236">2020</mods:date>
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<mods:number id="3747CA77734F39107D009749F4930976">2020-05-22</mods:number>
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<mods:number id="FAE6F854C4EC7E28BDC2F35BF723C1FC">650</mods:number>
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<treatment id="03BBEF6A2264CA73CA04FED262C4FC8B" ID-DOI="http://doi.org/10.5281/zenodo.3853282" ID-GBIF-Taxon="164249246" ID-Zenodo-Dep="3853282" LSID="urn:lsid:zoobank.org:act:6BEEA763-7AFB-4426-B919-024387DEC48E" httpUri="http://treatment.plazi.org/id/03BBEF6A2264CA73CA04FED262C4FC8B" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
<subSubSection id="C3080DF72264CA6FCA04FED265A4FE97" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BAD5E7C2264CA6FCA04FED262DEFEFA" blockId="12.[361,1226,266,364]" box="[629,959,266,293]" pageId="12" pageNumber="13">
<heading id="D0E5E9102264CA6FCA04FED262DEFEFA" box="[629,959,266,293]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C1225FF2264CA6FCA04FED2623CFEFA" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[629,861,266,293]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2264CA6FCA04FED2623CFEFA" bold="true" box="[629,861,266,293]" italics="true" pageId="12" pageNumber="13">Prodasineura kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FCB15FED362DEFEFA" box="[868,959,267,293]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
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<subSubSection id="C3080DF72264CA6FC918FE8965ABFEB4" box="[361,1226,337,364]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BAD5E7C2264CA6FC918FE8965ABFEB4" blockId="12.[361,1226,266,364]" box="[361,1226,337,364]" pageId="12" pageNumber="13">
<figureCitation id="132942F92264CA6FC918FE8960ABFEB4" box="[361,458,337,363]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13">Figs 9A</figureCitation>
, C–D,
<figureCitation id="132942F92264CA6FCA6DFE896359FEB4" box="[540,568,337,363]" captionStart="Fig" captionStartId="13.[189,232,1932,1958]" captionTargetBox="[214,1348,1250,1882]" captionTargetId="figure@13.[189,1398,1241,1895]" captionTargetPageId="13" captionText="Fig. 10. Wings base and genital ligula of Prodasineura Cowley, 1934 spp., males, holotypes A–B. P. kong sp. nov., ZCDTU 2019071811-ODO. C–D. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916144" httpUri="https://zenodo.org/record/5916144/files/figure.png" pageId="12" pageNumber="13">10</figureCitation>
A–B,
<figureCitation id="132942F92264CA6FCA0EFE8963FAFEB4" box="[639,667,337,363]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
A, C, E,
<figureCitation id="132942F92264CA6FCB73FE89627FFEB4" box="[770,798,337,363]" captionStart="Fig" captionStartId="15.[189,232,1896,1922]" captionTargetBox="[196,1391,1266,1855]" captionTargetId="figure@15.[189,1398,1263,1858]" captionTargetPageId="15" captionText="Fig. 12. Head of Prodasineura Cowley, 1934 spp., frontal view. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov. C. holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916152" httpUri="https://zenodo.org/record/5916152/files/figure.png" pageId="12" pageNumber="13">12</figureCitation>
A–B,
<figureCitation id="132942F92264CA6FCB14FE8962E1FEB4" box="[869,896,337,363]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="12" pageNumber="13">13</figureCitation>
C–D,
<figureCitation id="132942F92264CA6FCBB8FE896284FEB4" box="[969,997,337,363]" captionStart="Fig" captionStartId="17.[189,232,1894,1920]" captionTargetBox="[222,1369,1178,1818]" captionTargetId="figure@17.[189,1398,1165,1857]" captionTargetPageId="17" captionText="Fig. 14. Habitus of Prodasineura Cowley, 1934 spp. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov., C. ♂, holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5916162" httpUri="https://zenodo.org/record/5916162/files/figure.png" pageId="12" pageNumber="13">14</figureCitation>
A–B,
<figureCitation id="132942F92264CA6FCC5DFE896528FEB3" box="[1068,1097,337,364]" captionStart="Fig" captionStartId="18.[189,232,2003,2029]" captionTargetBox="[189,1398,334,1962]" captionTargetId="figure@18.[186,1401,331,1964]" captionTargetPageId="18" captionText="Fig. 15. Prodasineura Cowley, 1934 spp., males, in nature. A. P. kong sp. nov. B. P. lancastrei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5916166" httpUri="https://zenodo.org/record/5916166/files/figure.png" pageId="12" pageNumber="13">15</figureCitation>
A,
<figureCitation id="132942F92264CA6FCC1CFE8965EAFEB4" box="[1133,1163,337,363]" captionStart="Fig" captionStartId="19.[189,232,1790,1816]" captionTargetBox="[227,1357,268,1747]" captionTargetId="figure@19.[208,1376,262,1753]" captionTargetPageId="19" captionText="Fig. 16. Habitat of Prodasineura Cowley, 1934 spp. in Vietnam. A. Gia Trang stream in Chu Mom Ray National Park, home of P. doisuthepensis Hoess, 2007 and P. verticalis Selys, 1860. B. Typical stream where P. lancastrei sp. nov., P. autumnalis (Fraser, 1922) and P. croconota Ris, 1916 occur, in Pu Mat National Park. C. Open limestone stream in Tan Hoa Commune, Minh Hoa District of Quang Binh Province with three species: P. autumnalis, P. kong sp. nov. and P. coerulescens (Fraser, 1932) coexisting. D. Habitat of P. hoffmanni Kosterin, 2015 and P. autumnalis in Mang Canh, Kon Plong District of Kon Tum Province." figureDoi="http://doi.org/10.5281/zenodo.5916170" httpUri="https://zenodo.org/record/5916170/files/figure.png" pageId="12" pageNumber="13">16</figureCitation>
C,
<figureCitation id="132942F92264CA6FCCDCFE8965ABFEB4" box="[1197,1226,337,363]" captionStart="Fig" captionStartId="21.[189,232,1932,1958]" captionTargetBox="[357,1189,265,1880]" captionTargetId="figure@21.[357,1216,261,1894]" captionTargetPageId="21" captionText="Fig. 18. Distribution map of Prodasineura Cowley, 1934 spp. in Vietnam (orange group species). (●) P. croconota Ris, 1916. (●) P. verticalis Selys, 1860. (●) Prodasineura kong sp. nov. (●) P. lancastrei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5916174" httpUri="https://zenodo.org/record/5916174/files/figure.png" pageId="12" pageNumber="13">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3080DF72264CA6FC8CCFE4A655FFC80" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BAD5E7C2264CA6FC8CCFE4A6057FE73" blockId="12.[189,310,402,428]" box="[189,310,402,428]" pageId="12" pageNumber="13">
<heading id="D0E5E9102264CA6FC8CCFE4A6057FE73" bold="true" box="[189,310,402,428]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B966826E2264CA6FC8CCFE4A6057FE73" bold="true" box="[189,310,402,428]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD5E7C2264CA6FC8CCFE19655FFC80" blockId="12.[189,1399,449,863]" pageId="12" pageNumber="13">
Male very similar to those of
<taxonomicName id="4C1225FF2264CA6FCA6FFE1A63D0FE04" authorityName="Ris" authorityYear="1916" box="[542,689,450,475]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croconota">
<emphasis id="B966826E2264CA6FCA6FFE1A63D0FE04" box="[542,689,450,475]" italics="true" pageId="12" pageNumber="13">P. croconota</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1225FF2264CA6FCA9DFE1A6216FE04" authorityName="Selys" authorityYear="1860" box="[748,887,449,475]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966826E2264CA6FCA9DFE1A6216FE04" box="[748,887,449,475]" italics="true" pageId="12" pageNumber="13">P. verticalis</emphasis>
</taxonomicName>
but differing in following characters: large transversal stripe on dorsal surface of head (
<figureCitation id="132942F92264CA6FCAA4FE3C6257FE20" box="[725,822,484,511]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13">Fig. 9A</figureCitation>
), interrupted centrally, not connecting, covering anterior ocellus as in
<taxonomicName id="4C1225FF2264CA6FC9B3FDD0632FFDFD" authorityName="Selys" authorityYear="1860" box="[450,590,520,546]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966826E2264CA6FC9B3FDD0632FFDFD" box="[450,590,520,546]" italics="true" pageId="12" pageNumber="13">P. verticalis</emphasis>
</taxonomicName>
(
<figureCitation id="132942F92264CA6FCA10FDD063DDFDFD" box="[609,700,519,546]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[201,1386,670,1878]" captionTargetId="figure@9.[189,1398,658,1890]" captionTargetPageId="9" captionText="Fig. 6. Prodasineura Cowley, 1934 spp. A–C. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. D–F. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. A, E = ♂, head and thorax, lateral view; C–D = ♂, synthorax, dorsal view; B, F = ♀, head and thorax, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916136" httpUri="https://zenodo.org/record/5916136/files/figure.png" pageId="12" pageNumber="13">Fig. 6E</figureCitation>
) (with small spots in
<taxonomicName id="4C1225FF2264CA6FCBB6FDD0653DFDFE" authorityName="Ris" authorityYear="1916" box="[967,1116,520,545]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croconota">
<emphasis id="B966826E2264CA6FCBB6FDD0653DFDFE" box="[967,1116,520,545]" italics="true" pageId="12" pageNumber="13">P. croconota</emphasis>
</taxonomicName>
,
<figureCitation id="132942F92264CA6FCC1CFDD065ACFDFD" box="[1133,1229,519,546]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[201,1386,670,1878]" captionTargetId="figure@9.[189,1398,658,1890]" captionTargetPageId="9" captionText="Fig. 6. Prodasineura Cowley, 1934 spp. A–C. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. D–F. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. A, E = ♂, head and thorax, lateral view; C–D = ♂, synthorax, dorsal view; B, F = ♀, head and thorax, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916136" httpUri="https://zenodo.org/record/5916136/files/figure.png" pageId="12" pageNumber="13">Fig. 6A</figureCitation>
); antehumeral stripe narrow (
<figureCitation id="132942F92264CA6FC916FDF360A5FD9A" box="[359,452,555,581]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13">Fig. 9A</figureCitation>
), not large shield-shaped as in
<taxonomicName id="4C1225FF2264CA6FCB5BFDF462DDFD9A" authorityName="Ris" authorityYear="1916" box="[810,956,556,581]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croconota">
<emphasis id="B966826E2264CA6FCB5BFDF462DDFD9A" box="[810,956,556,581]" italics="true" pageId="12" pageNumber="13">P. croconota</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1225FF2264CA6FCB85FDF4651CFD9A" authorityName="Selys" authorityYear="1860" box="[1012,1149,555,581]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966826E2264CA6FCB85FDF4651CFD9A" box="[1012,1149,555,581]" italics="true" pageId="12" pageNumber="13">P. verticalis</emphasis>
</taxonomicName>
(
<figureCitation id="132942F92264CA6FCCFDFDF365B1FD99" box="[1164,1232,555,582]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[201,1386,670,1878]" captionTargetId="figure@9.[189,1398,658,1890]" captionTargetPageId="9" captionText="Fig. 6. Prodasineura Cowley, 1934 spp. A–C. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. D–F. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. A, E = ♂, head and thorax, lateral view; C–D = ♂, synthorax, dorsal view; B, F = ♀, head and thorax, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916136" httpUri="https://zenodo.org/record/5916136/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
C–D). In fully mature specimens, paraprocts of
<taxonomicName id="4C1225FF2264CA6FCA4DFD9763A4FDB7" authorityName="Selys" authorityYear="1860" box="[572,709,590,616]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966826E2264CA6FCA4DFD9763A4FDB7" box="[572,709,590,616]" italics="true" pageId="12" pageNumber="13">P. verticalis</emphasis>
</taxonomicName>
entirely black (
<figureCitation id="132942F92264CA6FCB08FD9662DDFDB6" box="[889,956,590,617]" captionStart="Fig" captionStartId="10.[189,232,1932,1958]" captionTargetBox="[205,1369,1189,1890]" captionTargetId="figure@10.[189,1398,1186,1893]" captionTargetPageId="10" captionText="Fig. 7. Prodasineura Cowley, 1934 spp., males. A–C. P. croconota, ZCDTU 2017060802-ODO. D–F. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. A, D, G = tip of abdomen, dorsal view; B, E = tip of abdomen, lateral view; C, F = genital ligula, oblique-ventral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916138" httpUri="https://zenodo.org/record/5916138/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
D–E) while yellow apically in
<taxonomicName id="4C1225FF2264CA6FCD53FD976417FDB7" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1314,1398,590,616]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2264CA6FCD53FD976417FDB7" box="[1314,1398,590,616]" italics="true" pageId="12" pageNumber="13">P. kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FC8CCFDAA607BFD53" box="[189,282,626,652]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="132942F92264CA6FC95DFDA960EAFD54" box="[300,395,625,651]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13">Fig. 9C</figureCitation>
). Cerci distinctly different in shape (
<figureCitation id="132942F92264CA6FCB37FDA962A9FD53" box="[838,968,625,652]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13">Fig. 9C, E</figureCitation>
). Cerci, paraprocts yellow or bluish (except for black ventral side of cerci and base of paraprocts) in
<taxonomicName id="4C1225FF2264CA6FCBBBFD4D653EFD71" authorityName="Ris" authorityYear="1916" box="[970,1119,661,686]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croconota">
<emphasis id="B966826E2264CA6FCBBBFD4D653EFD71" box="[970,1119,661,686]" italics="true" pageId="12" pageNumber="13">P. croconota</emphasis>
</taxonomicName>
(
<figureCitation id="132942F92264CA6FCC00FD4C65D9FD70" box="[1137,1208,660,687]" captionStart="Fig" captionStartId="10.[189,232,1932,1958]" captionTargetBox="[205,1369,1189,1890]" captionTargetId="figure@10.[189,1398,1186,1893]" captionTargetPageId="10" captionText="Fig. 7. Prodasineura Cowley, 1934 spp., males. A–C. P. croconota, ZCDTU 2017060802-ODO. D–F. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. A, D, G = tip of abdomen, dorsal view; B, E = tip of abdomen, lateral view; C, F = genital ligula, oblique-ventral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916138" httpUri="https://zenodo.org/record/5916138/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
A–B). Structure of posterior pronotal lobe of female prothorax somewhat similar to that in
<taxonomicName id="4C1225FF2264CA6FCC46FD6065AAFD0E" authorityName="Ris" authorityYear="1916" box="[1079,1227,696,721]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croconota">
<emphasis id="B966826E2264CA6FCC46FD6065AAFD0E" box="[1079,1227,696,721]" italics="true" pageId="12" pageNumber="13">P. croconota</emphasis>
</taxonomicName>
(
<figureCitation id="132942F92264CA6FCCACFD60640CFD0D" box="[1245,1389,696,722]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="12" pageNumber="13">Fig. 13A, D</figureCitation>
) with two short, robust horns, unlike two long horns of
<taxonomicName id="4C1225FF2264CA6FCB3FFD0462B8FD2A" authorityName="Selys" authorityYear="1860" box="[846,985,731,757]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966826E2264CA6FCB3FFD0462B8FD2A" box="[846,985,731,757]" italics="true" pageId="12" pageNumber="13">P. verticalis</emphasis>
</taxonomicName>
(
<figureCitation id="132942F92264CA6FCB9AFD036521FD2A" box="[1003,1088,731,757]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="12" pageNumber="13">Fig. 13</figureCitation>
G–H).
<taxonomicName id="4C1225FF2264CA6FCCE5FD036417FD2A" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1172,1398,731,757]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2264CA6FCCE5FD036417FD2A" box="[1172,1398,731,757]" italics="true" pageId="12" pageNumber="13">Prodasineura kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FC8CCFD276076FCC6" box="[189,279,767,793]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
similar to
<taxonomicName id="4C1225FF2264CA6FC9E2FD276342FCC7" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[403,547,766,792]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2264CA6FC9E2FD276342FCC7" box="[403,547,766,792]" italics="true" pageId="12" pageNumber="13">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FCA5BFD2763E5FCC6" box="[554,644,767,793]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
by sharing same character of narrow antehumeral stripe on male synthorax, but males differ very much in structure of appendages, wing colouration. Females differ in structure of posterior pronotal lobe (see Diagnosis for
<taxonomicName id="4C1225FF2264CA6FCB4BFC9D62AAFC81" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[826,971,836,862]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2264CA6FCB4BFC9D62AAFC81" box="[826,971,836,862]" italics="true" pageId="12" pageNumber="13">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FCBA3FC9D6550FC80" box="[978,1073,837,863]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C3080DF72264CA6FC8CCFC5E634BFBCA" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8BAD5E7C2264CA6FC8CCFC5E6025FC7F" blockId="12.[189,324,902,928]" box="[189,324,902,928]" pageId="12" pageNumber="13">
<heading id="D0E5E9102264CA6FC8CCFC5E6025FC7F" bold="true" box="[189,324,902,928]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B966826E2264CA6FC8CCFC5E6025FC7F" bold="true" box="[189,324,902,928]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD5E7C2264CA6FC8CCFC6C634BFBCA" blockId="12.[189,1399,948,1045]" pageId="12" pageNumber="13">
The new species is named after the giant ape Kong since it was discovered at the filming location of the American monster film “Kong: Skull Island”, in Tu Lan Caves System of
<collectingRegion id="49D6909E2264CA6FCC6AFC00647AFC2D" box="[1051,1307,984,1010]" country="Vietnam" name="Quang Binh" pageId="12" pageNumber="13">Quang Binh Province</collectingRegion>
, central
<collectingCountry id="F3051EEC2264CA6FC8CCFC236045FBCA" box="[189,292,1019,1045]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
. A noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF6D0EF42264CA6FC8CCF8B0607CF833" ID-DOI="http://doi.org/10.5281/zenodo.5916142" ID-Zenodo-Dep="5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1896,1922]" targetBox="[203,1384,1207,1855]" targetPageId="12">
<paragraph id="8BAD5E7C2264CA6FC8CCF8B0607CF833" blockId="12.[189,1399,1896,2028]" pageId="12" pageNumber="13">
<emphasis id="B966826E2264CA6FC8CCF8B0606FF85D" bold="true" box="[189,270,1896,1922]" pageId="12" pageNumber="13">Fig. 9.</emphasis>
Structures of
<taxonomicName id="4C1225FF2264CA6FC9CEF8B06270F85D" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[447,785,1896,1922]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2264CA6FC9CEF8B0633EF85D" box="[447,607,1896,1922]" italics="true" pageId="12" pageNumber="13">Prodasineura</emphasis>
<bibRefCitation id="EF83238D2264CA6FCA1BF8B06270F85D" author="Cowley J." box="[618,785,1896,1922]" pageId="12" pageNumber="13" pagination="200 - 205" refId="ref10504" refString="Cowley J. 1934. Changes in the generic names of the Odonata. Entomologist 67: 200 - 205." type="journal article" year="1934">Cowley, 1934</bibRefCitation>
</taxonomicName>
spp., males, holotypes.
<emphasis id="B966826E2264CA6FCC48F8B065FBF85D" bold="true" box="[1081,1178,1896,1922]" pageId="12" pageNumber="13">A, C–D</emphasis>
.
<taxonomicName id="4C1225FF2264CA6FCCDCF8B16467F85D" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1197,1286,1896,1922]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2264CA6FCCDCF8B16467F85D" box="[1197,1286,1896,1922]" italics="true" pageId="12" pageNumber="13">P. kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FCD60F8B16413F85C" box="[1297,1394,1897,1923]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019071811-ODO.
<emphasis id="B966826E2264CA6FCA64F854630FF879" bold="true" box="[533,622,1932,1958]" pageId="12" pageNumber="13">B, E–F</emphasis>
.
<taxonomicName id="4C1225FF2264CA6FCA0EF8546272F879" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[639,787,1932,1958]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2264CA6FCA0EF8546272F879" box="[639,787,1932,1958]" italics="true" pageId="12" pageNumber="13">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152264CA6FCB6CF854621CF879" box="[797,893,1932,1958]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale.
</paragraph>
</caption>
<subSubSection id="C3080DF72265CA6EC8CCFED362CDFD5B" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BAD5E7C2265CA6EC8CCFED360CBFEFA" blockId="13.[189,426,267,293]" box="[189,426,267,293]" pageId="13" pageNumber="14">
<heading id="D0E5E9102265CA6EC8CCFED360CBFEFA" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B966826E2265CA6EC8CCFED360CBFEFA" bold="true" box="[189,426,267,293]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7A54212265CA6EC89BFEE265E9FE45" ID-GBIF-Occurrence="2625217341" collectingDate="2019-07-18" collectionCode="T" collectorName="Q. T. Phan &amp; Q. P. Ngo" country="Vietnam" elevation="144" latitude="17.776" location="Tan Hoa Commune" longLatPrecision="55" longitude="106.0701" municipality="Minh Hoa District" pageId="13" pageNumber="14" specimenCode="ZCDTU 2019071811-ODO" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
<paragraph id="8BAD5E7C2265CA6EC89BFEE2603DFE8B" blockId="13.[189,1398,314,410]" box="[234,348,314,340]" pageId="13" pageNumber="14">
<emphasis id="B966826E2265CA6EC89BFEE2603DFE8B" bold="true" box="[234,348,314,340]" pageId="13" pageNumber="14">
<typeStatus id="54A9E0DE2265CA6EC89BFEE2603DFE8B" box="[234,348,314,340]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFE8565E9FE45" blockId="13.[189,1398,314,410]" pageId="13" pageNumber="14">
<collectingCountry id="F3051EEC2265CA6EC8CCFE856025FEA8" box="[189,324,349,375]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
•
<specimenCount id="9D1495F52265CA6EC92BFE856013FEA8" box="[346,370,349,375]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
;
<collectingRegion id="49D6909E2265CA6EC90FFE85631DFEA9" box="[382,636,348,375]" country="Vietnam" name="Quang Binh" pageId="13" pageNumber="14">Quang Binh Province</collectingRegion>
,
<collectingMunicipality id="6BC9C4062265CA6ECAF6FE84623FFEA9" box="[647,862,348,375]" pageId="13" pageNumber="14">Minh Hoa District</collectingMunicipality>
,
<location id="8ECD08A72265CA6ECB19FE85652FFEA8" LSID="urn:lsid:plazi:treatment:03BBEF6A2264CA73CA04FED262C4FC8B:8ECD08A72265CA6ECB19FE85652FFEA8" box="[872,1102,349,375]" country="Vietnam" latitude="17.776" longLatPrecision="55" longitude="106.0701" municipality="Minh Hoa District" name="Tan Hoa Commune" pageId="13" pageNumber="14" stateProvince="Quang Binh Province">Tan Hoa Commune</location>
;
<geoCoordinate id="EE2638BB2265CA6ECC29FE8465B3FEA8" box="[1112,1234,348,375]" degrees="17.776" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="17.776">17.776° N</geoCoordinate>
,
<geoCoordinate id="EE2638BB2265CA6ECCACFE846410FEA8" box="[1245,1393,348,375]" degrees="106.0701" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="106.0701">106.0701° E</geoCoordinate>
;
<elevation id="003FB94F2265CA6EC8CCFE586026FE45" box="[189,327,384,410]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="13" pageNumber="14" unit="m" value="144.0">
<quantity id="4CEAF3992265CA6EC8CCFE586066FE45" box="[189,263,384,410]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="13" pageNumber="14" unit="m" value="144.0">144 m</quantity>
a.s.l.
</elevation>
;
<date id="FFAC78BC2265CA6EC924FE586089FE45" box="[341,488,384,410]" pageId="13" pageNumber="14" value="2019-07-18">
<collectingDate id="EFE881542265CA6EC924FE586089FE45" box="[341,488,384,410]" pageId="13" pageNumber="14" value="2019-07-18">18 Jul. 2019</collectingDate>
</date>
;
<collectorName id="26E73BAA2265CA6EC985FE58630BFE45" box="[500,618,384,410]" pageId="13" pageNumber="14">Q.T. Phan</collectorName>
&amp;
<collectorName id="26E73BAA2265CA6ECAFEFE58639AFE45" box="[655,763,384,410]" pageId="13" pageNumber="14">Q.P. Ngo</collectorName>
leg.;
<specimenCode id="DBB4F6072265CA6ECB4DFE5865E5FE45" box="[828,1156,384,410]" pageId="13" pageNumber="14">ZCDTU 2019071811-ODO</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BAD5E7C2265CA6EC89BFE626377FE0B" blockId="13.[189,1399,442,644]" box="[234,534,442,468]" pageId="13" pageNumber="14">
<emphasis id="B966826E2265CA6EC89BFE626006FE0B" bold="true" box="[234,359,442,468]" pageId="13" pageNumber="14">
<typeStatus id="54A9E0DE2265CA6EC89BFE626006FE0B" box="[234,359,442,468]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
</emphasis>
(9 ♂♂,
<specimenCount id="9D1495F52265CA6EC9BBFE62636BFE0B" box="[458,522,442,468]" pageId="13" pageNumber="14" type="female">2 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFE0562CDFD5B" blockId="13.[189,1399,442,644]" pageId="13" pageNumber="14">
<materialsCitation id="3B7A54212265CA6EC8CCFE0563D5FDC5" ID-GBIF-Occurrence="2625217314" collectingDate="2016-04-14" collectionCode="T" collectorName="Q. T. Phan" country="Vietnam" elevation="144" latitude="17.776" location="Quang Binh Province" longLatPrecision="55" longitude="106.0701" municipality="Minh Hoa District" pageId="13" pageNumber="14" specimenCode="ZCDTU 2016041412-ODO" specimenCount="1" specimenCount-male="1" stateProvince="Quang Binh Province">
<collectingCountry id="F3051EEC2265CA6EC8CCFE056025FE28" box="[189,324,477,503]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
–
<emphasis id="B966826E2265CA6EC91CFE0563E7FE28" bold="true" box="[365,646,477,503]" pageId="13" pageNumber="14">
<collectingRegion id="49D6909E2265CA6EC91CFE0563E7FE28" box="[365,646,477,503]" country="Vietnam" name="Quang Binh" pageId="13" pageNumber="14">Quang Binh Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1495F52265CA6ECADAFE0563BEFE28" box="[683,735,477,503]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<date id="FFAC78BC2265CA6ECCBBFE056413FE28" box="[1226,1394,477,503]" pageId="13" pageNumber="14" value="2016-04-14">
<collectingDate id="EFE881542265CA6ECCBBFE056413FE28" box="[1226,1394,477,503]" pageId="13" pageNumber="14" value="2016-04-14">14 Apr. 2016</collectingDate>
</date>
;
<collectorName id="26E73BAA2265CA6EC8CCFDD86050FDC5" box="[189,305,512,538]" pageId="13" pageNumber="14">Q.T. Phan</collectorName>
leg.;
<specimenCode id="DBB4F6072265CA6EC901FDD863D5FDC5" box="[368,692,512,538]" pageId="13" pageNumber="14">ZCDTU 2016041412-ODO</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7A54212265CA6ECABBFDD860A5FDE2" ID-GBIF-Occurrence="2625217332" collectingDate="2019-07-18" collectionCode="ZCDTU" collectorName="Phan &amp; Q. P. Ngo" country="Vietnam" elevation="144" latitude="17.776" location="Tan Hoa Commune" longLatPrecision="55" longitude="106.0701" municipality="Minh Hoa District" pageId="13" pageNumber="14" specimenCode="ZCDTU 2019071812-18-ODO" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Quang Binh">
<specimenCount id="9D1495F52265CA6ECABBFDD8626CFDC4" box="[714,781,512,539]" pageId="13" pageNumber="14" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D1495F52265CA6ECB69FDD8623BFDC4" box="[792,858,512,539]" pageId="13" pageNumber="14" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBB4F6072265CA6ECD64FDD860A5FDE2" pageId="13" pageNumber="14">ZCDTU 2019071812-18-ODO</specimenCode>
</materialsCitation>
. –
<materialsCitation id="3B7A54212265CA6EC99BFDFB6256FDBE" ID-GBIF-Occurrence="2625217313" collectingDate="2014-06-25" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="22.444" location="Cho Ra Commune" longLatPrecision="75" longitude="105.846" municipality="Ba Be District" pageId="13" pageNumber="14" specimenCount="2" specimenCount-male="2" stateProvince="Bac Kan Province">
<emphasis id="B966826E2265CA6EC99BFDFB63B1FDE2" bold="true" box="[490,720,547,573]" pageId="13" pageNumber="14">
<collectingRegion id="49D6909E2265CA6EC99BFDFB63B1FDE2" box="[490,720,547,573]" country="Vietnam" name="Bac Kan" pageId="13" pageNumber="14">Bac Kan Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1495F52265CA6ECA9FFDFB6254FDE1" box="[750,821,547,574]" pageId="13" pageNumber="14" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="6BC9C4062265CA6ECB34FDFC6295FDE2" box="[837,1012,547,574]" pageId="13" pageNumber="14">Ba Be District</collectingMunicipality>
,
<location id="8ECD08A72265CA6ECC72FDFB6587FDE2" LSID="urn:lsid:plazi:treatment:03BBEF6A2264CA73CA04FED262C4FC8B:8ECD08A72265CA6ECC72FDFB6587FDE2" box="[1027,1254,547,574]" country="Vietnam" latitude="22.444" longLatPrecision="75" longitude="105.846" municipality="Ba Be District" name="Cho Ra Commune" pageId="13" pageNumber="14" stateProvince="Bac Kan Province">Cho Ra Commune</location>
;
<geoCoordinate id="EE2638BB2265CA6ECC85FDFB6413FDE1" box="[1268,1394,547,574]" degrees="22.444" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="22.444">22.444° N</geoCoordinate>
,
<geoCoordinate id="EE2638BB2265CA6EC8CCFD9E6029FDBE" box="[189,328,582,609]" degrees="105.846" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="105.846">105.846° E</geoCoordinate>
;
<date id="FFAC78BC2265CA6EC929FD9E6098FDBE" box="[344,505,582,609]" pageId="13" pageNumber="14" value="2014-06-25">
<collectingDate id="EFE881542265CA6EC929FD9E6098FDBE" box="[344,505,582,609]" pageId="13" pageNumber="14" value="2014-06-25">25 Jun. 2014</collectingDate>
</date>
;
<collectorName id="26E73BAA2265CA6ECA79FD9F63F3FDBF" box="[520,658,582,609]" pageId="13" pageNumber="14">T. Kompier</collectorName>
leg.;
<collectionCode id="ED03C6B92265CA6ECAAAFD9F6256FDBE" box="[731,823,583,609]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B7A54212265CA6ECB2EFD9E62C9FD5B" ID-GBIF-Occurrence="2625217310" collectingDate="2015-05-20" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="22.681" location="City" longLatPrecision="75" longitude="106.273" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang Province">
<emphasis id="B966826E2265CA6ECB2EFD9E6537FDBF" bold="true" box="[863,1110,582,609]" pageId="13" pageNumber="14">
<collectingRegion id="49D6909E2265CA6ECB2EFD9E6537FDBF" box="[863,1110,582,609]" country="Vietnam" name="Cao Bang" pageId="13" pageNumber="14">Cao Bang Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1495F52265CA6ECC07FD9F65C9FDBE" box="[1142,1192,583,609]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D6909E2265CA6ECCC8FD9F6450FDBE" box="[1209,1329,583,609]" country="Vietnam" name="Cao Bang" pageId="13" pageNumber="14">Cao Bang</collectingRegion>
<location id="8ECD08A72265CA6ECD4DFD9E6413FDBF" LSID="urn:lsid:plazi:treatment:03BBEF6A2264CA73CA04FED262C4FC8B:8ECD08A72265CA6ECD4DFD9E6413FDBF" box="[1340,1394,582,608]" country="Vietnam" latitude="22.681" longLatPrecision="75" longitude="106.273" name="City" pageId="13" pageNumber="14" stateProvince="Cao Bang Province">City</location>
;
<geoCoordinate id="EE2638BB2265CA6EC8CCFDB26058FD5B" box="[189,313,618,644]" degrees="22.681" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="22.681">22.681° N</geoCoordinate>
,
<geoCoordinate id="EE2638BB2265CA6EC934FDB260ADFD5B" box="[325,460,618,644]" degrees="106.273" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="106.273">106.273° E</geoCoordinate>
;
<date id="FFAC78BC2265CA6EC9A9FDB26319FD5B" box="[472,632,617,644]" pageId="13" pageNumber="14" value="2015-05-20">
<collectingDate id="EFE881542265CA6EC9A9FDB26319FD5B" box="[472,632,617,644]" pageId="13" pageNumber="14" value="2015-05-20">20 May 2015</collectingDate>
</date>
;
<collectorName id="26E73BAA2265CA6ECAF5FDB2626BFD5B" box="[644,778,618,644]" pageId="13" pageNumber="14">T. Kompier</collectorName>
leg.;
<collectionCode id="ED03C6B92265CA6ECB3DFDB262C9FD5B" box="[844,936,618,644]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3080DF72265CA73C8CCFD7B65FEFE07" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="description">
<paragraph id="8BAD5E7C2265CA6EC8CCFD7B602FFD62" blockId="13.[189,334,675,701]" box="[189,334,675,701]" pageId="13" pageNumber="14">
<heading id="D0E5E9102265CA6EC8CCFD7B602FFD62" bold="true" box="[189,334,675,701]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B966826E2265CA6EC8CCFD7B602FFD62" bold="true" box="[189,334,675,701]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC89BFD0A638DFD32" blockId="13.[189,1095,722,784]" box="[234,748,722,749]" pageId="13" pageNumber="14">
<emphasis id="B966826E2265CA6EC89BFD0A604BFD33" bold="true" box="[234,298,722,748]" pageId="13" pageNumber="14">Male</emphasis>
(
<figureCitation id="132942F92265CA6EC948FD0A60E2FD33" box="[313,387,722,748]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="13" pageNumber="14">Figs 9</figureCitation>
A–C, 10A–B, 12A, 14A, 15A)
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFD2E6526FCCF" blockId="13.[189,1095,722,784]" box="[189,1095,757,784]" pageId="13" pageNumber="14">
MEASUREMENTS. Hind wing
<quantity id="4CEAF3992265CA6EC98EFD2D6334FCCF" box="[511,597,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="20.0">20 mm</quantity>
, abdomen (including appendages)
<quantity id="4CEAF3992265CA6ECB9CFD2D6522FCCF" box="[1005,1091,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="14" unit="mm" value="31.0">31 mm</quantity>
.
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFCE8632FFC6C" blockId="13.[189,1399,815,947]" pageId="13" pageNumber="14">
HEAD. Labrum, mandible, genae, anteclypeus dark red except for upper genae; upper genae black. Space between eye margins, genae shining black (
<figureCitation id="132942F92265CA6ECB7EFC8A620EFCB3" box="[783,879,850,876]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="13" pageNumber="14">Fig. 9A</figureCitation>
). Frontal, dorsal surfaces of head, including postclypeus, antennae, matt black except for large orange transverse stripe running from eye margins across anterior ocellus (
<figureCitation id="132942F92265CA6EC9A2FC416320FC6C" box="[467,577,921,947]" captionStart="Fig" captionStartId="15.[189,232,1896,1922]" captionTargetBox="[196,1391,1266,1855]" captionTargetId="figure@15.[189,1398,1263,1858]" captionTargetPageId="15" captionText="Fig. 12. Head of Prodasineura Cowley, 1934 spp., frontal view. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov. C. holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916152" httpUri="https://zenodo.org/record/5916152/files/figure.png" pageId="13" pageNumber="14">Fig. 12A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFC0B646DFB89" blockId="13.[189,1399,979,1111]" pageId="13" pageNumber="14">
THORAX. Prothorax black with orange markings on distal margin of each pronotal lobe as follows: thin stripe on anterior lobe, small spots on posterior lobe, large oval spot on middle pronotal lobe (
<figureCitation id="132942F92265CA6ECD7AFC2E640BFBCF" box="[1291,1386,1014,1040]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="13" pageNumber="14">Fig. 9A</figureCitation>
). Propleuron entirely black. Synthorax black with narrow orange antehumeral stripe; two other distinct stripes: paler orange on metepisternum, yellowish on lower margin of metepimeron (
<figureCitation id="132942F92265CA6ECCD1FBE46461FB89" box="[1184,1280,1084,1110]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="13" pageNumber="14">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2265CA6EC8CCFBAF60AFFB6C" blockId="13.[189,1399,1142,1203]" pageId="13" pageNumber="14">LEGS. Mostly black except for some small dark yellow spots on coxa, trochanter, femur; flexor surface of tibia dark yellowish.</paragraph>
<caption id="DF6D0EF42265CA6EC8CCF854602BF832" ID-DOI="http://doi.org/10.5281/zenodo.5916144" ID-Zenodo-Dep="5916144" httpUri="https://zenodo.org/record/5916144/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1932,1958]" targetBox="[214,1348,1250,1882]" targetPageId="13">
<paragraph id="8BAD5E7C2265CA6EC8CCF854602BF832" blockId="13.[189,1399,1932,2029]" pageId="13" pageNumber="14">
<emphasis id="B966826E2265CA6EC8CCF8546076F879" bold="true" box="[189,279,1932,1958]" pageId="13" pageNumber="14">Fig. 10.</emphasis>
Wings base and genital ligula of
<taxonomicName id="4C1225FF2265CA6ECAFEF85462B4F879" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[655,981,1932,1958]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2265CA6ECAFEF854624EF879" box="[655,815,1932,1958]" italics="true" pageId="13" pageNumber="14">Prodasineura</emphasis>
<bibRefCitation id="EF83238D2265CA6ECB44F85462B4F879" author="Cowley J." box="[821,981,1932,1958]" pageId="13" pageNumber="14" pagination="200 - 205" refId="ref10504" refString="Cowley J. 1934. Changes in the generic names of the Odonata. Entomologist 67: 200 - 205." type="journal article" year="1934">Cowley, 1934</bibRefCitation>
</taxonomicName>
spp., males, holotypes
<emphasis id="B966826E2265CA6ECCAEF8546477F879" bold="true" box="[1247,1302,1932,1958]" pageId="13" pageNumber="14">A–B</emphasis>
.
<taxonomicName id="4C1225FF2265CA6ECD52F8556417F879" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1315,1398,1932,1958]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2265CA6ECD52F8556417F879" box="[1315,1398,1932,1958]" italics="true" pageId="13" pageNumber="14">P. kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152265CA6EC8CCF868607AF815" box="[189,283,1968,1994]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019071811-ODO.
<emphasis id="B966826E2265CA6ECA0CF87763D7F816" bold="true" box="[637,694,1967,1993]" pageId="13" pageNumber="14">C–D</emphasis>
.
<taxonomicName id="4C1225FF2265CA6ECAB7F8686239F816" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[710,856,1967,1993]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2265CA6ECAB7F8686239F816" box="[710,856,1967,1993]" italics="true" pageId="13" pageNumber="14">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152265CA6ECB10F86862DEF815" box="[865,959,1968,1994]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019051212-ODO. Images not to scale.
</paragraph>
</caption>
<paragraph id="8BAD5E7C2266CA6DC8CCFED46021FE96" blockId="14.[189,1399,267,329]" pageId="14" pageNumber="15">
WINGS. Completely hyaline, Px 16, 14 in forewings, hindwings respectively. Pterostigma black (
<figureCitation id="132942F92266CA6DC8B7FEF66052FE96" box="[198,307,302,329]" captionStart="Fig" captionStartId="13.[189,232,1932,1958]" captionTargetBox="[214,1348,1250,1882]" captionTargetId="figure@13.[189,1398,1241,1895]" captionTargetPageId="13" captionText="Fig. 10. Wings base and genital ligula of Prodasineura Cowley, 1934 spp., males, holotypes A–B. P. kong sp. nov., ZCDTU 2019071811-ODO. C–D. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916144" httpUri="https://zenodo.org/record/5916144/files/figure.png" pageId="14" pageNumber="15">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2266CA6DC8CCFEA863C7FE0F" blockId="14.[189,1399,368,464]" pageId="14" pageNumber="15">
ABDOMEN. Mostly black except: yellow triangular spots on lower margins of S1; yellowish bands on ventro-lateral surfaces of S2; small spot on anterior margin of S3, whitish spot on dorsal anterior margin of S4–7; S8–10 entirely black (
<figureCitation id="132942F92266CA6DCA5AFE6E63F8FE0F" box="[555,665,438,464]" captionStart="Fig" captionStartId="17.[189,232,1894,1920]" captionTargetBox="[222,1369,1178,1818]" captionTargetId="figure@17.[189,1398,1165,1857]" captionTargetPageId="17" captionText="Fig. 14. Habitus of Prodasineura Cowley, 1934 spp. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov., C. ♂, holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5916162" httpUri="https://zenodo.org/record/5916162/files/figure.png" pageId="14" pageNumber="15">Fig. 14A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2266CA6DC8CCFE2063E7FD87" blockId="14.[189,1399,504,601]" pageId="14" pageNumber="15">
ANAL APPENDAGES. Typical of members of genus
<taxonomicName id="4C1225FF2266CA6DCA97FE2062E7FDCD" authorityName="Cowley" authorityYear="1934" box="[742,902,504,530]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2266CA6DCA97FE2062E7FDCD" box="[742,902,504,530]" italics="true" pageId="14" pageNumber="15">Prodasineura</emphasis>
</taxonomicName>
with cerci hammer-like shaped; paraprocts broad at base, tapering abruptly to rounded tip; cerci black excepted tip dark yellowish; paraprocts black at base; remain yellowish (
<figureCitation id="132942F92266CA6DC989FDE6635CFD87" box="[504,573,574,600]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
C–D).
</paragraph>
<paragraph id="8BAD5E7C2266CA6DC8CCFD586384FD62" blockId="14.[189,1398,640,701]" pageId="14" pageNumber="15">
GENITAL LIGULA. Latero-apically with two robust flagella with pointed tips; subbasal shorter flagellum, expanding apically as rounded lobe (
<figureCitation id="132942F92266CA6DCA1CFD7B63B9FD62" box="[621,728,675,701]" captionStart="Fig" captionStartId="13.[189,232,1932,1958]" captionTargetBox="[214,1348,1250,1882]" captionTargetId="figure@13.[189,1398,1241,1895]" captionTargetPageId="13" captionText="Fig. 10. Wings base and genital ligula of Prodasineura Cowley, 1934 spp., males, holotypes A–B. P. kong sp. nov., ZCDTU 2019071811-ODO. C–D. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916144" httpUri="https://zenodo.org/record/5916144/files/figure.png" pageId="14" pageNumber="15">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2266CA6DC89BFD3C6390FD21" blockId="14.[189,1110,740,802]" box="[234,753,740,767]" pageId="14" pageNumber="15">
<emphasis id="B966826E2266CA6DC89BFD3C6024FD21" bold="true" box="[234,325,740,766]" pageId="14" pageNumber="15">Female</emphasis>
(
<figureCitation id="132942F92266CA6DC924FD3C6366FD20" box="[341,519,740,767]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="14" pageNumber="15">Figs 11A, C, E</figureCitation>
,
<figureCitation id="132942F92266CA6DCA62FD3C6326FD21" box="[531,583,740,766]" captionStart="Fig" captionStartId="15.[189,232,1896,1922]" captionTargetBox="[196,1391,1266,1855]" captionTargetId="figure@15.[189,1398,1263,1858]" captionTargetPageId="15" captionText="Fig. 12. Head of Prodasineura Cowley, 1934 spp., frontal view. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov. C. holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916152" httpUri="https://zenodo.org/record/5916152/files/figure.png" pageId="14" pageNumber="15">12B</figureCitation>
,
<figureCitation id="132942F92266CA6DCA22FD3C630FFD21" box="[595,622,740,766]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="14" pageNumber="15">13</figureCitation>
C–D, 14B)
</paragraph>
<paragraph id="8BAD5E7C2266CA6DC8CCFCD06537FCFD" blockId="14.[189,1110,740,802]" box="[189,1110,775,802]" pageId="14" pageNumber="15">
MEASUREMENTS. Hindwing
<quantity id="4CEAF3992266CA6DC989FCDF6305FCFD" box="[504,612,775,802]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="14" pageNumber="15" unit="mm" value="20.5">20.5 mm</quantity>
, abdomen (including appendages)
<quantity id="4CEAF3992266CA6DCB8DFCD06530FCFD" box="[1020,1105,776,802]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="14" pageNumber="15" unit="mm" value="31.0">31 mm</quantity>
.
</paragraph>
<paragraph id="8BAD5E7C2266CA6CC8CCFC916079FE50" blockId="14.[189,1398,841,903]" lastBlockId="15.[189,1399,267,399]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
HEAD
<figureCitation id="132942F92266CA6DC97BFC91601AFCBC" box="[266,379,841,867]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="14" pageNumber="15">Figs 11A</figureCitation>
,
<figureCitation id="132942F92266CA6DC9E0FC9160A3FCBC" box="[401,450,841,867]" captionStart="Fig" captionStartId="15.[189,232,1896,1922]" captionTargetBox="[196,1391,1266,1855]" captionTargetId="figure@15.[189,1398,1263,1858]" captionTargetPageId="15" captionText="Fig. 12. Head of Prodasineura Cowley, 1934 spp., frontal view. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov. C. holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916152" httpUri="https://zenodo.org/record/5916152/files/figure.png" pageId="14" pageNumber="15">12B</figureCitation>
. Labrum yellowish with black dot beneath upper margin; mandibles, genae yellowish except for small black dot near anteclypeus; yellow colour extends from compound eyes just over postclypeus to level of base of antennae; antennae black; anteclypeus dark yellowish; postclypeus yellow with two separated black spots, black stripe centrally. Antennae black except for second segment, yellowish anteriorly. Front, rear of head black with large transverse yellowish stripe at level of anterior ocellus.
</paragraph>
<caption id="DF6D0EF42266CA6DC8CCF8546417F832" ID-DOI="http://doi.org/10.5281/zenodo.5916150" ID-Zenodo-Dep="5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1932,1958]" targetBox="[199,1388,972,1891]" targetPageId="14">
<paragraph id="8BAD5E7C2266CA6DC8CCF8546417F832" blockId="14.[189,1399,1932,2029]" pageId="14" pageNumber="15">
<emphasis id="B966826E2266CA6DC8CCF854607FF879" bold="true" box="[189,286,1932,1958]" pageId="14" pageNumber="15">Fig. 11.</emphasis>
<taxonomicName id="4C1225FF2266CA6DC95AF85463E2F879" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[299,643,1932,1958]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2266CA6DC95AF85460AAF879" box="[299,459,1932,1958]" italics="true" pageId="14" pageNumber="15">Prodasineura</emphasis>
<bibRefCitation id="EF83238D2266CA6DC9A8F85463E2F879" author="Cowley J." box="[473,643,1932,1958]" pageId="14" pageNumber="15" pagination="200 - 205" refId="ref10504" refString="Cowley J. 1934. Changes in the generic names of the Odonata. Entomologist 67: 200 - 205." type="journal article" year="1934">Cowley, 1934</bibRefCitation>
</taxonomicName>
spp., female paratypes.
<emphasis id="B966826E2266CA6DCBC9F8546540F879" bold="true" box="[952,1057,1932,1958]" pageId="14" pageNumber="15">A, C, E</emphasis>
.
<taxonomicName id="4C1225FF2266CA6DCC44F85565F0F879" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1077,1169,1932,1958]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2266CA6DCC44F85565F0F879" box="[1077,1169,1932,1958]" italics="true" pageId="14" pageNumber="15">P. kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152266CA6DCCEEF8556462F878" box="[1183,1283,1933,1959]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019071817-ODO.
<emphasis id="B966826E2266CA6DC9D6F868609EF815" bold="true" box="[423,511,1968,1994]" pageId="14" pageNumber="15">B, D, F</emphasis>
.
<taxonomicName id="4C1225FF2266CA6DCA7CF86863FCF816" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[525,669,1967,1993]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2266CA6DCA7CF86863FCF816" box="[525,669,1967,1993]" italics="true" pageId="14" pageNumber="15">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152266CA6DCAD4F8686260F815" box="[677,769,1968,1994]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale.
</paragraph>
</caption>
<paragraph id="8BAD5E7C2267CA6CC8CCFE77624FFDEC" blockId="15.[189,1398,431,563]" pageId="15" pageNumber="16">
THORAX
<figureCitation id="132942F92267CA6CC953FE7760F0FE16" box="[290,401,431,457]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="15" pageNumber="16">Figs 11A</figureCitation>
,
<figureCitation id="132942F92267CA6CC9D2FE7760DCFE16" box="[419,445,431,457]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="15" pageNumber="16">13</figureCitation>
C–D. Prothorax black, laterally yellowish, forming large oval stripe on middle pronotal lobe. In dorso-oblique view, posterior lobe with pair of raised robust anterior projections, two ear-like processes (
<figureCitation id="132942F92267CA6CC9A8FE2D634FFDD0" box="[473,558,501,527]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
C–D). In dorsal view, projections sharply curving upward (
<figureCitation id="132942F92267CA6CCC88FE2D6408FDD0" box="[1273,1385,501,527]" captionStart="Fig" captionStartId="16.[189,232,1896,1922]" captionTargetBox="[238,1320,930,1852]" captionTargetId="figure@16.[189,1398,923,1859]" captionTargetPageId="16" captionText="Fig. 13. Posterior pronotal lobe of prothorax of Prodasineura Cowley, 1934 spp., females, lateral and dorsal views. A–B. P. croconota Ris, 1916, ZCDTU 2017060802-ODO. C–D. P. kong sp. nov., ZCDTU 019071817-ODO. E–F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. G–H. P. verticalis Selys, 1860, ZCDTU 2018030611-ODO. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916160" httpUri="https://zenodo.org/record/5916160/files/figure.png" pageId="15" pageNumber="16">Fig. 13D</figureCitation>
). Synthorax black with three yellow stripes (
<figureCitation id="132942F92267CA6CCAC4FDC06240FDEC" box="[693,801,536,563]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="15" pageNumber="16">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2267CA6CC8CCFD8B602FFD50" blockId="15.[189,1398,594,655]" pageId="15" pageNumber="16">LEGS. Coxae yellowish with large outer black spot; trochanter yellow with flexor side black; tibia, tarsus, claws black.</paragraph>
<paragraph id="8BAD5E7C2267CA6CC8CCFD68635FFD33" blockId="15.[189,1398,687,748]" pageId="15" pageNumber="16">
WINGS
<figureCitation id="132942F92267CA6CC961FD68601BFD16" box="[272,378,687,714]" captionStart="Fig" captionStartId="17.[189,232,1894,1920]" captionTargetBox="[222,1369,1178,1818]" captionTargetId="figure@17.[189,1398,1165,1857]" captionTargetPageId="17" captionText="Fig. 14. Habitus of Prodasineura Cowley, 1934 spp. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov., C. ♂, holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5916162" httpUri="https://zenodo.org/record/5916162/files/figure.png" pageId="15" pageNumber="16">Fig. 14B</figureCitation>
. Hyaline, forewings slightly longer than hindwings, Px 15, 13 in forewings, hindwings respectively. Pterostigma brown.
</paragraph>
<paragraph id="8BAD5E7C2267CA6CC8CCFCD5647FFC6C" blockId="15.[189,1399,780,948]" pageId="15" pageNumber="16">
ABDOMEN
<figureCitation id="132942F92267CA6CC945FCD560A3FCF8" box="[308,450,780,807]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="15" pageNumber="16">Figs 11C, E</figureCitation>
,
<figureCitation id="132942F92267CA6CC9A2FCD56365FCF8" box="[467,516,781,807]" captionStart="Fig" captionStartId="17.[189,232,1894,1920]" captionTargetBox="[222,1369,1178,1818]" captionTargetId="figure@17.[189,1398,1165,1857]" captionTargetPageId="17" captionText="Fig. 14. Habitus of Prodasineura Cowley, 1934 spp. A–B. P. kong sp. nov. A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO. C–D. P. lancastrei sp. nov., C. ♂, holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5916162" httpUri="https://zenodo.org/record/5916162/files/figure.png" pageId="15" pageNumber="16">14B</figureCitation>
. Black with yellowish pattern: S1 with heart-shaped spot on lateral sides; S2 with narrow yellowish stripes along lower margins; ventro-lateral halves of S3–6 brown-yellowish with oval yellowish spot on anterior margin of each segments; S7 dark brownish and S8–9 black with yellowish stripe on lower margin; dorsal side of S8 with oval yellow spot posteriorly but extending to mid-dorsally on S9; lower margin of S10 black but with yellowish marking dorsally (
<figureCitation id="132942F92267CA6CCCD7FC416471FC6C" box="[1190,1296,921,947]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="15" pageNumber="16">Fig. 11C</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2267CA6CC8CCFC3A6368FBFF" blockId="15.[189,1399,994,1056]" pageId="15" pageNumber="16">
APPENDAGES. Yellowish except for lower margin of cerci; cerci dark brownish; ovipositor black, distal end yellowish (
<figureCitation id="132942F92267CA6CC900FBDD609DFBFF" box="[369,508,1029,1056]" captionStart="Fig" captionStartId="14.[189,232,1932,1958]" captionTargetBox="[199,1388,972,1891]" captionTargetId="figure@14.[189,1398,968,1894]" captionTargetPageId="14" captionText="Fig. 11. Prodasineura Cowley, 1934 spp., female paratypes. A, C, E. P. kong sp. nov., ZCDTU 2019071817-ODO. B, D, F. P. lancastrei sp. nov., ZCDTU 2019051217-ODO. A–B = head and thorax, lateral view; C–D = abdominal tip, dorsal view; E–F = abdominal tip, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916150" httpUri="https://zenodo.org/record/5916150/files/figure.png" pageId="15" pageNumber="16">Fig. 11C, E</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2267CA6CC89BFB9663B9FBB6" blockId="15.[189,1399,1102,1199]" box="[234,728,1102,1129]" pageId="15" pageNumber="16">
<emphasis id="B966826E2267CA6CC89BFB9663B9FBB6" bold="true" box="[234,728,1102,1129]" pageId="15" pageNumber="16">
Variation in male and female
<typeStatus id="54A9E0DE2267CA6CCA2DFB9763B9FBB6" box="[604,728,1103,1129]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD5E7C2267CA6CC8CCFBAA6416FB70" blockId="15.[189,1399,1102,1199]" pageId="15" pageNumber="16">
<typeStatus id="54A9E0DE2267CA6CC8CCFBAA604EFB53" box="[189,303,1138,1164]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
from
<collectingRegion id="49D6909E2267CA6CC904FBAA60BBFB53" box="[373,474,1138,1164]" country="Vietnam" name="Bac Kan" pageId="15" pageNumber="16">Bac Kan</collectingRegion>
with some orange on dorsum of cerci; male from type location with incomplete transverse band between compound eyes over anterior ocellus. Male from
<collectingRegion id="49D6909E2267CA6CCC59FB4D65FCFB70" box="[1064,1181,1173,1199]" country="Vietnam" name="Cao Bang" pageId="15" pageNumber="16">Cao Bang</collectingRegion>
with small orange
</paragraph>
<caption id="DF6D0EF42267CA6CC8CCF8B0619EF834" ID-DOI="http://doi.org/10.5281/zenodo.5916152" ID-Zenodo-Dep="5916152" httpUri="https://zenodo.org/record/5916152/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1896,1922]" targetBox="[196,1391,1266,1855]" targetPageId="15">
<paragraph id="8BAD5E7C2267CA6CC8CCF8B0619EF834" blockId="15.[189,1399,1896,2027]" pageId="15" pageNumber="16">
<emphasis id="B966826E2267CA6CC8CCF8B0607BF85D" bold="true" box="[189,282,1896,1922]" pageId="15" pageNumber="16">Fig. 12.</emphasis>
Head of
<taxonomicName id="4C1225FF2267CA6CC9F9F8B063B5F85D" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[392,724,1896,1922]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2267CA6CC9F9F8B06349F85D" box="[392,552,1896,1922]" italics="true" pageId="15" pageNumber="16">Prodasineura</emphasis>
<bibRefCitation id="EF83238D2267CA6CCA41F8B063B5F85D" author="Cowley J." box="[560,724,1896,1922]" pageId="15" pageNumber="16" pagination="200 - 205" refId="ref10504" refString="Cowley J. 1934. Changes in the generic names of the Odonata. Entomologist 67: 200 - 205." type="journal article" year="1934">Cowley, 1934</bibRefCitation>
</taxonomicName>
spp., frontal view.
<emphasis id="B966826E2267CA6CCBC9F8B0628EF85D" bold="true" box="[952,1007,1896,1922]" pageId="15" pageNumber="16">A–B</emphasis>
.
<taxonomicName id="4C1225FF2267CA6CCB8EF8B16534F85D" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1023,1109,1896,1922]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="kong" status="sp. nov.">
<emphasis id="B966826E2267CA6CCB8EF8B16534F85D" box="[1023,1109,1896,1922]" italics="true" pageId="15" pageNumber="16">P. kong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152267CA6CCC2CF8B065D8F85D" box="[1117,1209,1896,1922]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
A. ♂, holotype, ZCDTU 2019071811-ODO. B. ♀, paratype, ZCDTU 2019071817-ODO.
<emphasis id="B966826E2267CA6CCC40F853650BF87A" bold="true" box="[1073,1130,1931,1957]" pageId="15" pageNumber="16">C–D</emphasis>
.
<taxonomicName id="4C1225FF2267CA6CCC0DF854646EF87A" authority="Phan &amp; Ngo, 2020" authorityName="Phan &amp; Ngo" authorityYear="2020" box="[1148,1295,1931,1957]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="lancastrei" status="sp. nov.">
<emphasis id="B966826E2267CA6CCC0DF854646EF87A" box="[1148,1295,1931,1957]" italics="true" pageId="15" pageNumber="16">P. lancastrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2553F152267CA6CCD6BF8546416F879" box="[1306,1399,1932,1958]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
C. holotype, ZCDTU 2019051212-ODO. D. ♀, paratype, ZCDTU 2019051217-ODO. Images not to scale.
</paragraph>
</caption>
<paragraph id="8BAD5E7C2278CA73C8CCFED462A7FE50" blockId="16.[189,1399,267,399]" pageId="16" pageNumber="17">
spot on posterodorsal corner of propleuron. Other male
<typeStatus id="54A9E0DE2278CA73CB06FED46286FEF9" box="[887,999,268,294]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
from type location provided no significant differential features from
<typeStatus id="54A9E0DE2278CA73CA14FEF663ADFE97" box="[613,716,302,328]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
male. Compound eyes bright red in living male (
<figureCitation id="132942F92278CA73CC8FFEF66408FE96" box="[1278,1385,302,329]" captionStart="Fig" captionStartId="18.[189,232,2003,2029]" captionTargetBox="[189,1398,334,1962]" captionTargetId="figure@18.[186,1401,331,1964]" captionTargetPageId="18" captionText="Fig. 15. Prodasineura Cowley, 1934 spp., males, in nature. A. P. kong sp. nov. B. P. lancastrei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5916166" httpUri="https://zenodo.org/record/5916166/files/figure.png" pageId="16" pageNumber="17">Fig. 15A</figureCitation>
). Stripes on male synthorax bright orange in life (
<figureCitation id="132942F92278CA73CA87FE8A6202FEB2" box="[758,867,338,365]" captionStart="Fig" captionStartId="18.[189,232,2003,2029]" captionTargetBox="[189,1398,334,1962]" captionTargetId="figure@18.[186,1401,331,1964]" captionTargetPageId="18" captionText="Fig. 15. Prodasineura Cowley, 1934 spp., males, in nature. A. P. kong sp. nov. B. P. lancastrei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5916166" httpUri="https://zenodo.org/record/5916166/files/figure.png" pageId="16" pageNumber="17">Fig. 15A</figureCitation>
), but stripe on lower margin of metepimeron becoming yellowish after acetone treatment in
<typeStatus id="54A9E0DE2278CA73CA95FEAD622AFE50" box="[740,843,373,399]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(
<figureCitation id="132942F92278CA73CB2BFEAD62DBFE50" box="[858,954,373,399]" captionStart="Fig" captionStartId="12.[189,232,1896,1922]" captionTargetBox="[203,1384,1207,1855]" captionTargetId="figure@12.[189,1398,1203,1859]" captionTargetPageId="12" captionText="Fig. 9. Structures of Prodasineura Cowley, 1934 spp., males, holotypes. A, C–D. P. kong sp. nov., ZCDTU 2019071811-ODO. B, E–F. P. lancastrei sp. nov., ZCDTU 2019051212-ODO. A–B = head and thorax, lateral view; C, E = appendages, lateral view; D, F = appendages, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5916142" httpUri="https://zenodo.org/record/5916142/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BAD5E7C2278CA73C8CCFE6665FEFE07" blockId="16.[189,1183,446,473]" box="[189,1183,446,473]" pageId="16" pageNumber="17">
MEASUREMENTS. Hind wing
<quantity id="4CEAF3992278CA73C98EFE6663E0FE07" box="[511,641,446,472]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="16" pageNumber="17" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
, abdomen (including appendages)
<quantity id="4CEAF3992278CA73CC68FE6665FBFE07" box="[1049,1178,446,472]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="16" pageNumber="17" unit="mm" value="30.5" valueMax="31.0" valueMin="30.0">30–31 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3080DF72278CA73C8CCFDDF62C4FC8B" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BAD5E7C2278CA73C8CCFDDF6037FDFE" blockId="16.[189,342,519,545]" box="[189,342,519,545]" pageId="16" pageNumber="17">
<heading id="D0E5E9102278CA73C8CCFDDF6037FDFE" bold="true" box="[189,342,519,545]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B966826E2278CA73C8CCFDDF6037FDFE" bold="true" box="[189,342,519,545]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD5E7C2278CA73C8CCFDED60C3FDAC" blockId="16.[189,1399,565,628]" pageId="16" pageNumber="17">
<collectingCountry id="F3051EEC2278CA73C8CCFDED6041FD90" box="[189,288,565,591]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
(
<figureCitation id="132942F92278CA73C940FDED60EBFD8F" box="[305,394,565,592]" captionStart="Fig" captionStartId="21.[189,232,1932,1958]" captionTargetBox="[357,1189,265,1880]" captionTargetId="figure@21.[357,1216,261,1894]" captionTargetPageId="21" captionText="Fig. 18. Distribution map of Prodasineura Cowley, 1934 spp. in Vietnam (orange group species). (●) P. croconota Ris, 1916. (●) P. verticalis Selys, 1860. (●) Prodasineura kong sp. nov. (●) P. lancastrei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5916174" httpUri="https://zenodo.org/record/5916174/files/figure.png" pageId="16" pageNumber="17">Fig. 18</figureCitation>
):
<collectingRegion id="49D6909E2278CA73C9D0FDEE6376FD8F" box="[417,535,566,592]" country="Vietnam" name="Cao Bang" pageId="16" pageNumber="17">Cao Bang</collectingRegion>
(
<collectingRegion id="49D6909E2278CA73CA56FDEE63FEFD8F" box="[551,671,566,592]" country="Vietnam" name="Cao Bang" pageId="16" pageNumber="17">Cao Bang</collectingRegion>
City),
<collectingRegion id="49D6909E2278CA73CA82FDEE623BFD8F" box="[755,858,566,592]" country="Vietnam" name="Bac Kan" pageId="16" pageNumber="17">Bac Kan</collectingRegion>
(Ba Be District) and
<collectingRegion id="49D6909E2278CA73CC2DFDEE658AFD90" box="[1116,1259,565,592]" country="Vietnam" name="Quang Binh" pageId="16" pageNumber="17">Quang Binh</collectingRegion>
(Minh Hoa District) Provinces.
</paragraph>
<paragraph id="8BAD5E7C2278CA73C8CCFD7960D9FD63" blockId="16.[189,440,673,700]" box="[189,440,673,700]" pageId="16" pageNumber="17">
<heading id="D0E5E9102278CA73C8CCFD7960D9FD63" bold="true" box="[189,440,673,700]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B966826E2278CA73C8CCFD7960D9FD63" bold="true" box="[189,440,673,700]" pageId="16" pageNumber="17">Habitat and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD5E7C2278CA73C8CCFD0862C4FC8B" blockId="16.[189,1398,720,852]" pageId="16" pageNumber="17">
The new species was collected in
<collectingRegion id="49D6909E2278CA73CA33FD0863AEFD35" box="[578,719,720,746]" country="Vietnam" name="Quang Binh" pageId="16" pageNumber="17">Quang Binh</collectingRegion>
from an open stream (about
<quantity id="4CEAF3992278CA73CC6BFD086503FD34" box="[1050,1122,720,747]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="16" pageNumber="17" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</quantity>
wide) with fast running water a low limestone mountain area (
<figureCitation id="132942F92278CA73CAFAFD2B6399FCD1" box="[651,760,755,782]" captionStart="Fig" captionStartId="19.[189,232,1790,1816]" captionTargetBox="[227,1357,268,1747]" captionTargetId="figure@19.[208,1376,262,1753]" captionTargetPageId="19" captionText="Fig. 16. Habitat of Prodasineura Cowley, 1934 spp. in Vietnam. A. Gia Trang stream in Chu Mom Ray National Park, home of P. doisuthepensis Hoess, 2007 and P. verticalis Selys, 1860. B. Typical stream where P. lancastrei sp. nov., P. autumnalis (Fraser, 1922) and P. croconota Ris, 1916 occur, in Pu Mat National Park. C. Open limestone stream in Tan Hoa Commune, Minh Hoa District of Quang Binh Province with three species: P. autumnalis, P. kong sp. nov. and P. coerulescens (Fraser, 1932) coexisting. D. Habitat of P. hoffmanni Kosterin, 2015 and P. autumnalis in Mang Canh, Kon Plong District of Kon Tum Province." figureDoi="http://doi.org/10.5281/zenodo.5916170" httpUri="https://zenodo.org/record/5916170/files/figure.png" pageId="16" pageNumber="17">Fig. 16B</figureCitation>
). This species usually flies over the surface of water, fighting with two other species of
<taxonomicName id="4C1225FF2278CA73CA3AFCCF638AFCEE" authorityName="Cowley" authorityYear="1934" box="[587,747,791,817]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B966826E2278CA73CA3AFCCF638AFCEE" box="[587,747,791,817]" italics="true" pageId="16" pageNumber="17">Prodasineura</emphasis>
</taxonomicName>
present, for occupation of the habitat near the border of the stream, the blue
<taxonomicName id="4C1225FF2278CA73C9DBFCE3633EFC8B" baseAuthorityName="Fraser" baseAuthorityYear="1932" box="[426,607,826,852]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="coerulescens">
<emphasis id="B966826E2278CA73C9DBFCE3633EFC8B" box="[426,607,826,852]" italics="true" pageId="16" pageNumber="17">P. coerulescens</emphasis>
</taxonomicName>
and the dark
<emphasis id="B966826E2278CA73CA8FFCE362C4FC8B" box="[766,933,826,852]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C1225FF2278CA73CA8FFCE362C0FC8B" authorityName="" baseAuthorityName="Fraser" baseAuthorityYear="1922" box="[766,929,826,852]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="autumnalis">P. autumnalis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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