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<document id="8E5C7CC601803B126AC1E601B3C13875" ID-CLB-Dataset="46283" ID-DOI="10.5281/zenodo.278802" ID-GBIF-Dataset="275e1f6f-69ff-4f2e-a9a9-0d1cb7d286d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="278802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460392962328" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2011" docId="03BB87D5FF8EFF93FF7F2032FE549FF2" docLanguage="en" docName="zt03041p038.pdf" docOrigin="Zootaxa 3041" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fenneralpheus Felder &amp; Manning 1986" docType="treatment" docVersion="13" lastPageNumber="28" masterDocId="FF82FFADFF94FF88FFE82158FFA79D55" masterDocTitle="Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="27" updateTime="1698298436616" updateUser="ExternalLinkService">
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<mods:title id="4103B0E71A19AA138B0952EB35D7DC0F">Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda)</mods:title>
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<mods:namePart id="74900682553C20F075FE95C3305830DC">Anker, Arthur</mods:namePart>
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<mods:date id="E0CAB1BCC2F06BDB203AB999AEB86904">2011</mods:date>
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<mods:classification id="35B7D314BD637F4906E18824C8D8A8F5">journal article</mods:classification>
<mods:identifier id="AFD93F4F2BDAA8357F2290570DE18680" type="CLB-Dataset">46283</mods:identifier>
<mods:identifier id="22EFD5477476F9113B935D5521BA8F4D" type="DOI">10.5281/zenodo.278802</mods:identifier>
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<treatment id="03BB87D5FF8EFF93FF7F2032FE549FF2" ID-DOI="http://doi.org/10.5281/zenodo.4893579" ID-GBIF-Taxon="119417244" ID-Zenodo-Dep="4893579" LSID="urn:lsid:plazi:treatment:03BB87D5FF8EFF93FF7F2032FE549FF2" httpUri="http://treatment.plazi.org/id/03BB87D5FF8EFF93FF7F2032FE549FF2" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="C3086548FF8EFF92FF7F2032FD2D9CD1" box="[151,650,362,388]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BAD36C3FF8EFF92FF7F2032FD2D9CD1" blockId="26.[151,650,362,388]" box="[151,650,362,388]" pageId="26" pageNumber="27">
<heading id="D0E581AFFF8EFF92FF7F2032FD2D9CD1" bold="true" box="[151,650,362,388]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<taxonomicName id="4C124D40FF8EFF92FF7F2032FD2D9CD1" ID-CoL="84NPF" authority="Felder &amp; Manning, 1986" authorityName="Felder &amp; Manning" authorityYear="1986" box="[151,650,362,388]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FF7F2032FD2D9CD1" bold="true" box="[151,650,362,388]" pageId="26" pageNumber="27">
<emphasis id="B966EAD1FF8EFF92FF7F2032FEE89CD1" bold="true" box="[151,335,362,388]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
<bibRefCitation id="EF834B32FF8EFF92FEBE2032FD2D9CD1" author="Felder" box="[342,650,362,388]" pageId="26" pageNumber="27" refString="Felder, D. L. &amp; Manning R. B. (1986) A new genus and two new species of alpheid shrimps (Decapoda: Caridea) from South Florida. Journal of Crustacean Biology, 6, 497 - 508." type="journal article" year="1986">Felder &amp; Manning, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF8EFF92FF7F20E9FB0098C5" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="8BAD36C3FF8EFF92FF7F20E9FDCC9C92" blockId="26.[151,619,433,455]" box="[151,619,433,455]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB0211EDFF8EFF92FF7F20E9FDCC9C92" box="[151,619,433,455]" pageId="26" pageNumber="27">
<taxonomicName id="4C124D40FF8EFF92FF7F20E9FDC09C92" ID-CoL="84NPF" authority="Felder &amp; Manning, 1986: 497" authorityName="Felder &amp; Manning" authorityPageNumber="497" authorityYear="1986" box="[151,615,433,455]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FF7F20E9FE889C92" box="[151,303,433,455]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
<treatmentCitation id="0AB310D2FF8EFF92FEDD20E9FDC09C92" author="Felder" box="[309,615,433,455]" page="497" pageId="26" pageNumber="27" year="1986">
<bibRefCitation id="EF834B32FF8EFF92FEDD20E9FD939C92" author="Felder" box="[309,564,433,455]" pageId="26" pageNumber="27" refString="Felder, D. L. &amp; Manning R. B. (1986) A new genus and two new species of alpheid shrimps (Decapoda: Caridea) from South Florida. Journal of Crustacean Biology, 6, 497 - 508." type="journal article" year="1986">Felder &amp; Manning, 1986</bibRefCitation>
: 497
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAD36C3FF8EFF92FF7F20ACFE179839" blockId="26.[151,1437,499,2037]" pageId="26" pageNumber="27">
<emphasis id="B966EAD1FF8EFF92FF7F20ACFE209F59" bold="true" box="[151,391,499,524]" pageId="26" pageNumber="27">Emended diagnosis.</emphasis>
Carapace glabrous, not setose; frontal margin rounded, without orbital teeth, rostral projection, orbital crests; anterolateral suture present; pterygostomial angle subacutely protruding anteriorly or broadly rounded; branchiostegial margin with broad, somewhat membranous ventral lip; posterior margin with deep cardiac notch. Sixth abdominal somite with posteroventral articulated plate. Telson with two pairs of spiniform setae dorsally; posterior margin rounded, with two pairs of posterolateral spiniform setae; anal tubercles present. Eyes concealed in dorsal and lateral views; anteromesial margin bluntly projecting. Antennular peduncles flattened dorsoventrally; ventromesial carina of first article with large tooth; stylocerite appressed, not exceeding distal margin of first article; second article elongate, much longer than wide; lateral antennular flagellum with short secondary ramus. Antenna with stout basicerite bearing distoventral tooth; scaphocerite with small distolateral tooth; carpocerite overreaching scaphocerite. Mouthparts not especially modified; mandible with two-articulated palp; first maxilliped with elongate palp and broad caridean lobe; second maxilliped with elongate epipod. Third maxilliped with lateral plate on coxa moderately produced dorsally; tip of ultimate article unarmed. First pereiopods (chelipeds) very unequal in size and asymmetrical in shape, carried flexed; major cheliped stout, with enlarged, elongate chela; merus depressed ventrally, margins smooth; carpus short, more or less cup-shaped; chela with palm smooth, excavated ventromesially; fingers with armature consisting of large teeth with distal ridges, tooth-fossa system absent; adhesive discs, if present, feebly developed. Second pereiopod with carpus composed of five articles. Third pereiopod and fourth pereiopods very stout, strongly compressed laterally; ischium unarmed or with small spiniform seta; carpus short, with strong distoventral spiniform seta; propodus with strong spiniform setae on ventral margin; dactylus simple, subconical or subspatulate. Fifth pereiopod not compressed, shorter and more slender than third and fourth pereiopods; propodus with several setal rows distolaterally. Male second pleopod with appendix masculina exceeding appendix interna; female second pleopod with appendix interna only. Uropod with lateral lobe of protopod bifid distally; exopod distally truncate; diaeresis with deep incision and large triangular tooth near mesial margin of exopod. Gill/exopod formula: 5 pleurobranchs (P1–5), 1 arthrobranch (Mxp3), 2 lobe-like epipods (Mxp1–2), 0 podobranchs, 5 mastigobranchs (Mxp3, P1–4), 5 setobranchs (P1–5), 3–4 exopods (always Mxp1–3, sometimes P1).
</paragraph>
<paragraph id="8BAD36C3FF8EFF92FF2D242FFB0098C5" blockId="26.[151,1437,499,2037]" box="[197,1191,1399,1424]" pageId="26" pageNumber="27">
<emphasis id="B966EAD1FF8EFF92FF2D242FFE3698C5" bold="true" box="[197,401,1399,1424]" pageId="26" pageNumber="27">Species included.</emphasis>
<taxonomicName id="4C124D40FF8EFF92FE7F2421FCBE98C5" ID-CoL="85296" authority="Felder &amp; Manning, 1986" authorityName="Felder &amp; Manning" authorityYear="1986" box="[407,793,1400,1424]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B966EAD1FF8EFF92FE7F2421FE5B98C5" box="[407,508,1400,1424]" italics="true" pageId="26" pageNumber="27">F. chacei</emphasis>
<bibRefCitation id="EF834B32FF8EFF92FDEB2420FCBE98C5" author="Felder" box="[515,793,1400,1424]" pageId="26" pageNumber="27" refString="Felder, D. L. &amp; Manning R. B. (1986) A new genus and two new species of alpheid shrimps (Decapoda: Caridea) from South Florida. Journal of Crustacean Biology, 6, 497 - 508." type="journal article" year="1986">Felder &amp; Manning, 1986</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54A98861FF8EFF92FCC02420FCFD98C5" box="[808,858,1400,1424]" pageId="26" pageNumber="27">type</typeStatus>
species),
<taxonomicName id="4C124D40FF8EFF92FC212421FBE098C5" ID-CoL="85R33" box="[969,1095,1400,1424]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="orabovis" status="sp. nov.">
<emphasis id="B966EAD1FF8EFF92FC212421FBE098C5" box="[969,1095,1400,1424]" italics="true" pageId="26" pageNumber="27">F. orabovis</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF8EFF92FBA62420FB0098C5" bold="true" box="[1102,1191,1400,1424]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A25557AAFF8EFF92FBA62420FB0098C5" box="[1102,1191,1400,1424]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF8EFF92FF2D24C3FC5B98E1" box="[197,1020,1435,1460]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BAD36C3FF8EFF92FF2D24C3FC5B98E1" blockId="26.[151,1437,499,2037]" box="[197,1020,1435,1460]" pageId="26" pageNumber="27">
<emphasis id="B966EAD1FF8EFF92FF2D24C3FEF998E1" bold="true" box="[197,350,1435,1460]" pageId="26" pageNumber="27">Distribution.</emphasis>
Western Atlantic: southeastern
<collectingCountry id="F3057653FF8EFF92FD2A24C4FD5D98E1" box="[706,762,1436,1460]" name="United States of America" pageId="26" pageNumber="27">USA</collectingCountry>
(Florida) and
<collectingCountry id="F3057653FF8EFF92FC7324C5FC5098E1" box="[923,1015,1437,1460]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF8EFF92FF2D2498FA3B9AA1" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BAD36C3FF8EFF92FF2D2498FE469A49" blockId="26.[151,1437,499,2037]" pageId="26" pageNumber="27">
<emphasis id="B966EAD1FF8EFF92FF2D2498FE80988D" bold="true" box="[197,295,1472,1496]" pageId="26" pageNumber="27">Remark</emphasis>
s.
<bibRefCitation id="EF834B32FF8EFF92FED72498FDC7988D" author="Felder" box="[319,608,1472,1496]" pageId="26" pageNumber="27" refString="Felder, D. L. &amp; Manning R. B. (1986) A new genus and two new species of alpheid shrimps (Decapoda: Caridea) from South Florida. Journal of Crustacean Biology, 6, 497 - 508." type="journal article" year="1986">Felder &amp; Manning (1986)</bibRefCitation>
listed the following differences between
<taxonomicName id="4C124D40FF8EFF92FBC52498FB72988D" box="[1069,1237,1472,1496]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FBC52498FB72988D" box="[1069,1237,1472,1496]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D40FF8EFF92FAE32498FA33988D" box="[1291,1428,1472,1496]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FAE32498FA33988D" box="[1291,1428,1472,1496]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
: (1) the pterygostomial angle strongly produced anteriorly in
<taxonomicName id="4C124D40FF8EFF92FCB224BCFBA598A9" box="[858,1026,1508,1532]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FCB224BCFBA598A9" box="[858,1026,1508,1532]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(not in
<taxonomicName id="4C124D40FF8EFF92FB8C24BCFB4A98A9" box="[1124,1261,1508,1532]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FB8C24BCFB4A98A9" box="[1124,1261,1508,1532]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
); (2) the branchiostegial margin with a distinct ventral lip in
<taxonomicName id="4C124D40FF8EFF92FD5C2750FCFB9B75" box="[692,860,1544,1568]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FD5C2750FCFB9B75" box="[692,860,1544,1568]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(not in
<taxonomicName id="4C124D40FF8EFF92FC5F2750FBE79B75" box="[951,1088,1544,1568]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FC5F2750FBE79B75" box="[951,1088,1544,1568]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
); (3) the antennular peduncles depressed and with the second article “much the longest” in
<taxonomicName id="4C124D40FF8EFF92FCD22774FC459B11" box="[826,994,1580,1604]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FCD22774FC459B11" box="[826,994,1580,1604]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(not in
<emphasis id="B966EAD1FF8EFF92FBD22774FB1C9B11" box="[1082,1211,1580,1604]" italics="true" pageId="26" pageNumber="27">Leptapheus</emphasis>
); (4) the scaphocerite “spatulate” and without “anterolateral spine” (= distolateral tooth), but with an “anterior emargination” in
<taxonomicName id="4C124D40FF8EFF92FF7F272CFE989BD9" box="[151,319,1652,1676]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FF7F272CFE989BD9" box="[151,319,1652,1676]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(vs. scaphocerite non-spatulate and with a well-developed distolateral tooth in
<taxonomicName id="4C124D40FF8EFF92FB5A272CFA9C9BD9" box="[1202,1339,1652,1676]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FB5A272CFA9C9BD9" box="[1202,1339,1652,1676]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
); (5) the fingers of the major chela much longer, evenly toothed, not gaping in
<taxonomicName id="4C124D40FF8EFF92FC5B27C0FBFC9BE5" box="[947,1115,1688,1712]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FC5B27C0FBFC9BE5" box="[947,1115,1688,1712]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(vs. with the fingers shorter and gaping in
<taxonomicName id="4C124D40FF8EFF92FED127E4FE659B81" box="[313,450,1724,1748]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FED127E4FE659B81" box="[313,450,1724,1748]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
); (6) the much stouter third and fourth pereiopods in
<taxonomicName id="4C124D40FF8EFF92FBC827E4FB6F9B81" box="[1056,1224,1724,1748]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FBC827E4FB6F9B81" box="[1056,1224,1724,1748]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(relatively slender in
<taxonomicName id="4C124D40FF8EFF92FF5B27B8FE9B9BAD" box="[179,316,1760,1784]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FF5B27B8FE9B9BAD" box="[179,316,1760,1784]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
); and (7) the much larger size of
<taxonomicName id="4C124D40FF8EFF92FD4627B8FCF19BAD" box="[686,854,1760,1784]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FD4627B8FCF19BAD" box="[686,854,1760,1784]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
(cl
<quantity id="4CEA9B26FF8EFF92FC6927B8FBB69BAD" box="[897,1041,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.28" metricValueMin="0.9199999999999999" pageId="26" pageNumber="27" unit="mm" value="11.0" valueMax="12.8" valueMin="9.2">9.2-12.8 mm</quantity>
in
<taxonomicName id="4C124D40FF8EFF92FBDB27B9FB309BAD" box="[1075,1175,1760,1784]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B966EAD1FF8EFF92FBDB27B9FB309BAD" box="[1075,1175,1760,1784]" italics="true" pageId="26" pageNumber="27">F. chacei</emphasis>
</taxonomicName>
vs.
<quantity id="4CEA9B26FF8EFF92FB2C27B8FAE19BAD" box="[1220,1350,1760,1784]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="7.8" metricValueMin="2.9" pageId="26" pageNumber="27" unit="mm" value="5.35" valueMax="7.8" valueMin="2.9">2.9-7.8 mm</quantity>
in
<taxonomicName id="4C124D40FF8EFF92FA8127B9FF519A49" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FA8127B9FF519A49" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
spp. known to date).
</paragraph>
<paragraph id="8BAD36C3FF8EFF92FF2D2670FA3B9AA1" blockId="26.[151,1437,499,2037]" pageId="26" pageNumber="27">
The pterygostomial angle is not anteriorly produced in the below-described new species of
<taxonomicName id="4C124D40FF8EFF92FB052670FA329A15" box="[1261,1429,1832,1856]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FB052670FA329A15" box="[1261,1429,1832,1856]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
, thus eliminating the first difference between the two genera. The antennular peduncles are somewhat depressed and the second article distinctly longer than the first or third in
<taxonomicName id="4C124D40FF8EFF92FCC72629FC399ADD" box="[815,926,1904,1928]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B966EAD1FF8EFF92FCC72629FC399ADD" box="[815,926,1904,1928]" italics="true" pageId="26" pageNumber="27">L. forceps</emphasis>
</taxonomicName>
; therefore the third difference is also invalid. The size of the distolateral tooth and the shape of the blade of the scaphocerite appear to be quite variable in species of
<taxonomicName id="4C124D40FF8EFF92FF5D26E0FE999A85" box="[181,318,1976,2000]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FF5D26E0FE999A85" box="[181,318,1976,2000]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
, whilst a small but distinct tooth is present in the new species of
<taxonomicName id="4C124D40FF8EFF92FBF426E0FB639A85" box="[1052,1220,1976,2000]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FBF426E0FB639A85" box="[1052,1220,1976,2000]" italics="true" pageId="26" pageNumber="27">Fenneralpheus</emphasis>
</taxonomicName>
, making the fourth difference unreliable. The fingers of the major cheliped are not gaping in several species of
<taxonomicName id="4C124D40FF8EFF92FB532684FAE39AA1" box="[1211,1348,2012,2036]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8EFF92FB532684FAE39AA1" box="[1211,1348,2012,2036]" italics="true" pageId="26" pageNumber="27">Leptalpheus</emphasis>
</taxonomicName>
, e.g.
<emphasis id="B966EAD1FF8EFF92FA6D2685FA3B9AA1" box="[1413,1436,2013,2036]" italics="true" pageId="26" pageNumber="27">L.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3086548FF8FFF93FF7F21C0FE549FF2" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BAD36C3FF8FFF93FF7F21C0FEB59CF9" blockId="27.[151,1436,151,680]" pageId="27" pageNumber="28">
<taxonomicName id="4C124D40FF8FFF93FF7F21C0FEA89DE5" ID-CoL="85TJM" box="[151,271,152,176]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="marginalis" status="sp. nov.">
<emphasis id="B966EAD1FF8FFF93FF7F21C0FEA89DE5" box="[151,271,152,176]" italics="true" pageId="27" pageNumber="28">marginalis</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF8FFF93FEFE21C0FED79DE5" bold="true" box="[278,368,152,176]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A25557AAFF8FFF93FEFE21C0FED79DE5" box="[278,368,152,176]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C124D40FF8FFF93FE4F21C1FD9C9DE5" ID-CoL="85TJL" box="[423,571,152,176]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="hendrickxi" status="sp. nov.">
<emphasis id="B966EAD1FF8FFF93FE4F21C1FD9C9DE5" box="[423,571,152,176]" italics="true" pageId="27" pageNumber="28">L. hendrickxi</emphasis>
</taxonomicName>
<emphasis id="B966EAD1FF8FFF93FDAA21C0FD3B9DE5" bold="true" box="[578,668,152,176]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A25557AAFF8FFF93FDAA21C0FD3B9DE5" box="[578,668,152,176]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see above), and only slightly gaping in
<taxonomicName id="4C124D40FF8FFF93FB8E21C1FB589DE5" ID-CoL="85TJT" box="[1126,1279,152,176]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pierrenoeli">
<emphasis id="B966EAD1FF8FFF93FB8E21C1FB589DE5" box="[1126,1279,152,176]" italics="true" pageId="27" pageNumber="28">L. pierrenoeli</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF834B32FF8FFF93FAE621CFFEAB9D81" author="Anker" pageId="27" pageNumber="28" refString="Anker A. (2008) The alpheid shrimp genus Leptalpheus Williams, 1965 in the southwestern Caribbean Sea, with description of one new species from Panama (Crustacea: Decapoda). Zoosystema, 30, 781 - 794." type="journal article" year="2008">Anker 2008, fig. 2C, D</bibRefCitation>
). In fact, the major cheliped of
<taxonomicName id="4C124D40FF8FFF93FD8721E5FCAF9D81" ID-CoL="85TJT" box="[623,776,188,212]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pierrenoeli">
<emphasis id="B966EAD1FF8FFF93FD8721E5FCAF9D81" box="[623,776,188,212]" italics="true" pageId="27" pageNumber="28">L. pierrenoeli</emphasis>
</taxonomicName>
has some similarities to that of
<taxonomicName id="4C124D40FF8FFF93FB9921E5FB719D81" ID-CoL="85296" box="[1137,1238,188,212]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B966EAD1FF8FFF93FB9921E5FB719D81" box="[1137,1238,188,212]" italics="true" pageId="27" pageNumber="28">F. chacei</emphasis>
</taxonomicName>
, especially in the general shape and proportions of the fingers to the palm, however, differing in the armature of the finger cutting edges (
<bibRefCitation id="EF834B32FF8FFF93FF0D205CFE579C49" author="Anker" box="[229,496,260,284]" pageId="27" pageNumber="28" refString="Anker A. (2008) The alpheid shrimp genus Leptalpheus Williams, 1965 in the southwestern Caribbean Sea, with description of one new species from Panama (Crustacea: Decapoda). Zoosystema, 30, 781 - 794." type="journal article" year="2008">Anker 2008, fig. 2A–D</bibRefCitation>
). Finally, the difference in size is eliminated by the fairly large Indo-West Pacific
<taxonomicName id="4C124D40FF8FFF93FA6D205DFF5B9C15" ID-CoL="8533K" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B966EAD1FF8FFF93FA6D205DFF5B9C15" italics="true" pageId="27" pageNumber="28">L. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D40FF8FFF93FEDD2071FE7A9C15" ID-CoL="85TJH" box="[309,477,296,320]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B966EAD1FF8FFF93FEDD2071FE7A9C15" box="[309,477,296,320]" italics="true" pageId="27" pageNumber="28">L. denticulatus</emphasis>
</taxonomicName>
, with the
<typeStatus id="54A98861FF8FFF93FDB82070FD179C15" box="[592,688,296,320]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
of the latter species, at cl
<quantity id="4CEA9B26FF8FFF93FC36207FFBE49C6A" box="[990,1091,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="27" pageNumber="28" unit="mm" value="14.5">14.5 mm</quantity>
(
<bibRefCitation id="EF834B32FF8FFF93FBBA207FFAEE9C6A" author="Anker" box="[1106,1353,295,319]" pageId="27" pageNumber="28" refString="Anker A, Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
), being larger than the largest known specimen of
<taxonomicName id="4C124D40FF8FFF93FD982015FD729C31" ID-CoL="85296" box="[624,725,332,356]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B966EAD1FF8FFF93FD982015FD729C31" box="[624,725,332,356]" italics="true" pageId="27" pageNumber="28">F. chacei</emphasis>
</taxonomicName>
. Generally, size cannot be a character of generic importance, as exemplified by extreme variation within the genera
<taxonomicName id="4C124D40FF8FFF93FD322028FC4F9CD2" ID-CoL="84LB5" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[730,1000,367,392]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FD322028FC939CDD" box="[730,820,368,392]" italics="true" pageId="27" pageNumber="28">Alpheus</emphasis>
Fabricius, 1798
</taxonomicName>
and
<taxonomicName id="4C124D40FF8FFF93FBF72028FAB99CD2" ID-CoL="84TZP" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[1055,1310,367,392]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FBF72028FB399CDD" box="[1055,1182,368,392]" italics="true" pageId="27" pageNumber="28">Synalpheus</emphasis>
Bate, 1888
</taxonomicName>
(A. Anker, pers. obs.).
</paragraph>
<paragraph id="8BAD36C3FF8FFF93FF2D20EFFE549FF2" blockId="27.[151,1436,151,680]" pageId="27" pageNumber="28">
Thus, the only morphological characters that allow distinguishing effectively
<taxonomicName id="4C124D40FF8FFF93FBCF20E0FB689C85" ID-CoL="84NPF" box="[1063,1231,440,464]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FBCF20E0FB689C85" box="[1063,1231,440,464]" italics="true" pageId="27" pageNumber="28">Fenneralpheus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C124D40FF8FFF93FAFB20E0FA3B9C85" ID-CoL="84PMC" box="[1299,1436,440,464]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FAFB20E0FA3B9C85" box="[1299,1436,440,464]" italics="true" pageId="27" pageNumber="28">Leptalpheus</emphasis>
</taxonomicName>
(as currently defined) are the presence of a large ventral lip on the branchiostegial margin and the very stout third and fourth pereiopods, with short carpi and propodi, both armed with unusually robust spiniform setae. The presence of a large, somewhat membranous lip on the branchiostegial margin is shared with the Indo-West Pacific
<taxonomicName id="4C124D40FF8FFF93FF7F2310FDBE9F0A" ID-CoL="85HYX" authority="Anker &amp; Jeng, 2006" authorityName="Anker &amp; Jeng" authorityYear="2006" box="[151,537,583,608]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FF7F2310FE869F35" box="[151,289,584,608]" italics="true" pageId="27" pageNumber="28">Richalpheus</emphasis>
<bibRefCitation id="EF834B32FF8FFF93FEC3231FFDBE9F0A" author="Anker" box="[299,537,583,608]" pageId="27" pageNumber="28" refString="Anker, A. &amp; Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391." type="journal article" year="2006">Anker &amp; Jeng, 2006</bibRefCitation>
</taxonomicName>
, which can be easily distinguished from
<taxonomicName id="4C124D40FF8FFF93FC1B2310FB3C9F35" ID-CoL="84NPF" box="[1011,1179,584,608]" class="Malacostraca" family="Alpheidae" genus="Fenneralpheus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAD1FF8FFF93FC1B2310FB3C9F35" box="[1011,1179,584,608]" italics="true" pageId="27" pageNumber="28">Fenneralpheus</emphasis>
</taxonomicName>
by the absence of the mastigobranch-setobranch complex (Mxp3, P1-5) and the very different shape and finger armature of the major cheliped (
<bibRefCitation id="EF834B32FF8FFF93FEEC23D7FE409FF2" author="Anker" box="[260,487,655,680]" pageId="27" pageNumber="28" refString="Anker, A. &amp; Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391." type="journal article" year="2006">Anker &amp; Jeng 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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