217 lines
20 KiB
XML
217 lines
20 KiB
XML
<document id="E613499EEA2CA823089E4D0C82762F88" ID-DOI="10.11646/zootaxa.4007.2.10" ID-GBIF-Dataset="164eccb8-f9c5-4766-8803-7c5d65650e77" ID-ISSN="1175-5326" ID-Zenodo-Dep="238866" ID-ZooBank="0A5D7B5A-17D2-4F97-AD23-3510ADC6228B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461218563564" checkinUser="plazi" docAuthor="Yan, Zhu-Qi & Li, Li-Zhen" docDate="2015" docId="03BB87875209DF452CF965BFFA466626" docLanguage="en" docName="zt04007p282.pdf" docOrigin="Zootaxa 4007 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetrabothrus" docType="key" docVersion="9" lastPageNumber="281" masterDocId="FF82FFFF520DDF402C6E610FFFDD6704" masterDocTitle="New species and additional records of Tetrabothrus Bernhauer from China (Coleoptera, Staphylinidae, Aleocharinae)" masterLastPageNumber="282" masterPageNumber="276" pageNumber="280" updateTime="1698603212977" updateUser="plazi">
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<mods:titleInfo id="5D81AA2BDC7B3F9E7EB021D05B6AF190">
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<mods:title id="170B086C8B15D0207B4CD39748E93134">New species and additional records of Tetrabothrus Bernhauer from China (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
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<mods:namePart id="EA7B143D76F81D84E99371DA7EE38116">Yan, Zhu-Qi</mods:namePart>
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<mods:namePart id="E5A75F6A3098FB4D00FC0DFBECEFC697">Li, Li-Zhen</mods:namePart>
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<mods:title id="63EE4119B49CE389D19B98D7A892B146">Zootaxa</mods:title>
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<mods:part id="35EBE13549B58714D2B3EB4A5B92B197">
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<mods:date id="1C1924CA94125C114198D7DD4DBE2CB4">2015</mods:date>
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<mods:number id="C7A936EB27883957DAF4C80570938317">4007</mods:number>
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<mods:number id="A43AB8A10A6A6BE280D39AF4777AE8AE">2</mods:number>
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<mods:identifier id="65D95D897C981CDC080F60331A65C400" type="DOI">10.11646/zootaxa.4007.2.10</mods:identifier>
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<treatment id="03BB87875209DF452CF965BFFA466626" ID-DOI="http://doi.org/10.5281/zenodo.5481388" ID-GBIF-Taxon="119631260" ID-Zenodo-Dep="5481388" LSID="urn:lsid:plazi:treatment:03BB87875209DF452CF965BFFA466626" httpUri="http://treatment.plazi.org/id/03BB87875209DF452CF965BFFA466626" lastPageId="5" lastPageNumber="281" pageId="4" pageNumber="280">
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<subSubSection id="C308651A5209DF442CF965BFFD8263CD" box="[151,607,1200,1225]" pageId="4" pageNumber="280" type="nomenclature">
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<paragraph id="8BAD36915209DF442CF965BFFD8263CD" blockId="4.[151,607,1200,1225]" box="[151,607,1200,1225]" pageId="4" pageNumber="280">
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<heading id="D0E581FD5209DF442CF965BFFD8263CD" bold="true" box="[151,607,1200,1225]" fontSize="10" level="2" pageId="4" pageNumber="280" reason="2">
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<emphasis id="B966EA835209DF442CF965BFFD8263CD" bold="true" box="[151,607,1200,1225]" pageId="4" pageNumber="280">
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Key to Chinese species of
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<taxonomicName id="4C124D125209DF442DA465BFFD8263CD" ID-CoL="8VSGM" box="[458,607,1200,1225]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="genus">
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<emphasis id="B966EA835209DF442DA465BFFD8263CD" bold="true" box="[458,607,1200,1225]" italics="true" pageId="4" pageNumber="280">Tetrabothrus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C308651A5209DF452CF965F7FA466626" lastPageId="5" lastPageNumber="281" pageId="4" pageNumber="280" type="key">
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<keyStep id="30E62E345209DF442CF965F7FA46622E" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF965F7FA466209" blockId="4.[151,1435,1272,2026]" box="[151,1435,1272,1293]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF965F7FA466209" box="[151,1435,1272,1293]" pageId="4" pageNumber="280">1 Legs distinctly intramuscular; femora broad................................................................ 2</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF9641AFA46622E" blockId="4.[151,1435,1272,2026]" box="[151,1435,1301,1322]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9641AFA46622E" box="[151,1435,1301,1322]" pageId="4" pageNumber="280">- Legs normal; femora narrow............................................................................ 3</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="30E62E345209DF442CF9643CFA466286" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF9643CFA46624C" blockId="4.[151,1435,1272,2026]" box="[151,1435,1331,1352]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9643CFA46624C" box="[151,1435,1331,1352]" pageId="4" pageNumber="280">
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2 The length of tubus of aedeagus as long as the bulbus, and much narrower than the bulbus. (Yunnan, Guangdong)...
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<taxonomicName id="4C124D125209DF442925643CFA46624C" box="[1355,1435,1331,1352]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="puezti">
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<emphasis id="B966EA835209DF442925643CFA46624C" box="[1355,1435,1331,1352]" italics="true" pageId="4" pageNumber="280">T. puezti</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF9645FFA466286" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9645FFA466286" pageId="4" pageNumber="280">
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- The length of tubus of aedeagus approximately 0.8 time as long as the bulbus, and little narrower than the bulbus. (
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<collectingCountry id="F30576015209DF442940645FFAAA6261" box="[1326,1399,1360,1381]" name="Taiwan" pageId="4" pageNumber="280">Taiwan</collectingCountry>
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)...........................................................................................
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<taxonomicName id="4C124D125209DF44297C6462FA466286" ID-CoL="55L32" box="[1298,1435,1389,1410]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B966EA835209DF44297C6462FA466286" box="[1298,1435,1389,1410]" italics="true" pageId="4" pageNumber="280">T. taiwanensis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E62E345209DF442CF96484FA4662B9" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF96484FA4662A4" blockId="4.[151,1435,1272,2026]" box="[151,1435,1419,1440]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF96484FA4662A4" box="[151,1435,1419,1440]" pageId="4" pageNumber="280">3 Elytra approximately less than 0.65 times as long as pronotum (EL/PL<0.65)..................................... 4</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF964A7FA4662B9" blockId="4.[151,1435,1272,2026]" box="[151,1435,1448,1469]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF964A7FA4662B9" box="[151,1435,1448,1469]" pageId="4" pageNumber="280">- Elytra more than 0.80 times as long as pronotum (EL/PL>0.80)................................................ 6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E62E345209DF442CF964CAFA4862FC" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF964CAFA4862DE" blockId="4.[151,1435,1272,2026]" box="[151,1429,1477,1498]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF964CAFA4862DE" box="[151,1429,1477,1498]" pageId="4" pageNumber="280">
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4
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<collectingCountry id="F30576015209DF442C8F64CAFECC62DE" box="[225,273,1477,1498]" name="India" pageId="4" pageNumber="280">Hind</collectingCountry>
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wings slightly longer than elytra (
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<collectingCountry id="F30576015209DF442E3764CAFD7F62DE" box="[601,674,1477,1498]" name="Taiwan" pageId="4" pageNumber="280">Taiwan</collectingCountry>
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)...................................................
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<taxonomicName id="4C124D125209DF44296764CAFA4862DE" ID-CoL="55L2X" box="[1289,1429,1477,1498]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="semiapterus">
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<emphasis id="B966EA835209DF44296764CAFA4862DE" box="[1289,1429,1477,1498]" italics="true" pageId="4" pageNumber="280">T. semiapterus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF964ECFA4862FC" blockId="4.[151,1435,1272,2026]" box="[151,1429,1507,1528]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF964ECFA4862FC" box="[151,1429,1507,1528]" pageId="4" pageNumber="280">
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-
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<collectingCountry id="F30576015209DF442C8F64ECFECC62FC" box="[225,273,1507,1528]" name="India" pageId="4" pageNumber="280">Hind</collectingCountry>
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wings entirely reduced............................................................................ 5
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="30E62E345209DF442CF9670FFA466169" pageId="4" pageNumber="280">
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||
<paragraph id="8BAD36915209DF442CF9670FFA466136" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9670FFA466136" pageId="4" pageNumber="280">
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5 Larger body size 6.3–7.0 mm; legs reddish-brown; tubus of aedeagus almost straight and pointed in lateral view (Yunnan)..............................................................................................
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<taxonomicName id="4C124D125209DF4429736712FA466136" ID-CoL="55L24" box="[1309,1435,1565,1586]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="brevalatus">
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<emphasis id="B966EA835209DF4429736712FA466136" box="[1309,1435,1565,1586]" italics="true" pageId="4" pageNumber="280">T. brevalatus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF96734FA466169" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF96734FA466169" pageId="4" pageNumber="280">
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- Smaller body size
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<quantity id="4CEA9B745209DF442DFB6734FDFE6154" box="[405,547,1595,1616]" metricMagnitude="-3" metricUnit="m" metricValue="4.255" metricValueMax="4.56" metricValueMin="3.95" pageId="4" pageNumber="280" unit="mm" value="4.255" valueMax="4.56" valueMin="3.95">3.95–4.56 mm</quantity>
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; legs yellowish-brown; tubus of aedeagus curved ventrad at base and pointed at apex in lat- eral view (Xizang)..........................................................................
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<taxonomicName id="4C124D125209DF44297B6757FA466169" ID-CoL="55L36" box="[1301,1435,1624,1645]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="xizangensis">
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<emphasis id="B966EA835209DF44297B6757FA466169" box="[1301,1435,1624,1645]" italics="true" pageId="4" pageNumber="280">T. xizangensis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E62E345209DF442CF9677AFA4661AC" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF9677AFA46618E" blockId="4.[151,1435,1272,2026]" box="[151,1435,1653,1674]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9677AFA46618E" box="[151,1435,1653,1674]" pageId="4" pageNumber="280">6 Forebody distinctly bicoloured........................................................................... 7</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF9679CFA4661AC" blockId="4.[151,1435,1272,2026]" box="[151,1435,1683,1704]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9679CFA4661AC" box="[151,1435,1683,1704]" pageId="4" pageNumber="280">- Forebody not distinctly bicoloured, more or less uniformly reddish to dark-brown.................................. 8</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E62E345209DF442CF967BFFA466019" pageId="4" pageNumber="280">
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<paragraph id="8BAD36915209DF442CF967BFFA4661E6" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF967BFFA4661E6" pageId="4" pageNumber="280">
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7 Pronotum pale-reddish, strongly contrasting with the blackish posterior portion of head and blackish elytra; longitudinal diam- eter of eye nearly thrice as long as postocular part (Northern Sichuan)...................................
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<taxonomicName id="4C124D125209DF44297167C2FA4661E6" ID-CoL="55L2W" box="[1311,1435,1741,1762]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="rubricollis">
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<emphasis id="B966EA835209DF44297167C2FA4661E6" box="[1311,1435,1741,1762]" italics="true" pageId="4" pageNumber="280">T. rubricollis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF967E4FA466019" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF967E4FA466019" pageId="4" pageNumber="280">
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- Pronotum yellowish, different from reddish-brown head and elytra; longitudinal diameter of eye nearly twice as long as pos- tocular part (wide spread)......................................................................
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<taxonomicName id="4C124D125209DF4429496607FA466019" ID-CoL="55L2G" box="[1319,1435,1800,1821]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="B966EA835209DF4429496607FA466019" box="[1319,1435,1800,1821]" italics="true" pageId="4" pageNumber="280">T. japonicus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E62E345209DF442CF9662AFA46605C" pageId="4" pageNumber="280">
|
||
<paragraph id="8BAD36915209DF442CF9662AFA46603E" blockId="4.[151,1435,1272,2026]" box="[151,1435,1829,1850]" pageId="4" pageNumber="280">
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||
<keyLead id="30E395A45209DF442CF9662AFA46603E" box="[151,1435,1829,1850]" pageId="4" pageNumber="280">8 Legs pale-yellowish, apices of the femora weakly to distinctly infuscate..........................................9</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF9664CFA46605C" blockId="4.[151,1435,1272,2026]" box="[151,1435,1859,1880]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9664CFA46605C" box="[151,1435,1859,1880]" pageId="4" pageNumber="280">- Legs reddish-brown to brown, bases of the femora sometimes paler............................................. 10</keyLead>
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</paragraph>
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</keyStep>
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<paragraph id="8BAD36915209DF442CF9666FFA4B60B4" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
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<keyLead id="30E395A45209DF442CF9666FFA4B60B4" pageId="4" pageNumber="280">
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<keyStep id="30E62E345209DF442CF9666FFA4B60B4" pageId="4" pageNumber="280">
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9 Apices of the femora very narrowly and indistinctly infuscate; anterior angles of pronotum visible in dorsal view; median lobe of aedeagus
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<quantity id="4CEA9B745209DF442D376672FE736096" box="[345,430,1917,1938]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="280" unit="mm" value="0.45">0.45 mm</quantity>
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long and with short, strongly angled ventral process; paramere small and with shorter apical lobe (wide spead)......................................................................................
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<taxonomicName id="4C124D125209DF44295D6694FA4B60B4" ID-CoL="55L2F" box="[1331,1430,1947,1968]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="inflexus">
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<emphasis id="B966EA835209DF44295D6694FA4B60B4" box="[1331,1430,1947,1968]" italics="true" pageId="4" pageNumber="280">T. inflexus</emphasis>
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</taxonomicName>
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</keyStep>
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</keyLead>
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</paragraph>
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<paragraph id="8BAD36915209DF442CF966B7FA4660EE" blockId="4.[151,1435,1272,2026]" pageId="4" pageNumber="280">
|
||
<keyLead id="30E395A45209DF442CF966B7FA4660EE" pageId="4" pageNumber="280">
|
||
- Apices of the femora broadly and distinctly infuscate; anterior angles of pronotum not visible in.dorsal view; median lobe of aedeagus
|
||
<quantity id="4CEA9B745209DF442D5166DAFE4E60EE" box="[319,403,2005,2026]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="280" unit="mm" value="0.5">0.50 mm</quantity>
|
||
long, and with evenly curved ventral process; paramere larger and with longer apical lobe (Yunnan, North
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAD36915208DF452C8F6198FA4867E3" blockId="5.[151,1436,151,290]" pageId="5" pageNumber="281">
|
||
Sichuan).......................................................................................
|
||
<taxonomicName id="4C124D125208DF4529296198FA4867A8" ID-CoL="55L27" box="[1351,1429,151,172]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="cavus">
|
||
<emphasis id="B966EA835208DF4529296198FA4867A8" box="[1351,1429,151,172]" italics="true" pageId="5" pageNumber="281">T. cavus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<keyLead id="30E395A45208DF452CF961BAFA4867E3" pageId="5" pageNumber="281">
|
||
<keyStep id="30E62E345208DF452CF961BAFA4867E3" pageId="5" pageNumber="281">
|
||
10 Head with obtusely marked posterior angle; bulbus of aedeagus narrow at base and the tubus long and straight in lateral view (Shaanxi)...................................................................................
|
||
<taxonomicName id="4C124D125208DF45294A61DDFA4867E3" ID-CoL="55L28" box="[1316,1429,210,231]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B966EA835208DF45294A61DDFA4867E3" box="[1316,1429,210,231]" italics="true" pageId="5" pageNumber="281">T. chinensis</emphasis>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAD36915208DF452CF961E0FA466626" blockId="5.[151,1436,151,290]" pageId="5" pageNumber="281">
|
||
<keyLead id="30E395A45208DF452CF961E0FA466626" pageId="5" pageNumber="281">
|
||
- Head with transverse posterior angles; bulbus of aedeagus rounded at base and the tubus short and curved in lateral view (
|
||
<collectingCountry id="F30576015208DF452C896002FE8D6626" box="[231,336,269,290]" name="Hong Kong" pageId="5" pageNumber="281">Hongkong</collectingCountry>
|
||
,
|
||
<collectingCountry id="F30576015208DF452D346002FE7E6626" box="[346,419,269,290]" name="Taiwan" pageId="5" pageNumber="281">Taiwan</collectingCountry>
|
||
)........................................................................
|
||
<taxonomicName id="4C124D125208DF4529786002FA466626" ID-CoL="55L2V" box="[1302,1435,269,290]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="rougemonti">
|
||
<emphasis id="B966EA835208DF4529786002FA466626" box="[1302,1435,269,290]" italics="true" pageId="5" pageNumber="281">T. rougemonti</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |