205 lines
24 KiB
XML
205 lines
24 KiB
XML
<document id="12236A330242CE2EE690070FB867B282" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55BE3FF14FF79F1F9FE50FDAA" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena tholasoris Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="136" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="135" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D8FD16E37665B89776DE11B125384413">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="9C5F88E7C7C0188108E7ABEC651C3738">Perkins, Philip D.</mods:namePart>
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<mods:title id="A8C81D20D2CF0CE15FF42446698F2E43">Zootaxa</mods:title>
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<mods:part id="383605C045A35936CF8FBD2B8221DF71">
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<mods:date id="B3AD93161FBC1F6A90895E75433065C9">2011</mods:date>
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<mods:number id="EFDFBB326B02C0C9687561D8B8E8ECE5">2011-07-08</mods:number>
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<mods:number id="D7BFF888D771AFF73D5D504FE559B451">2944</mods:number>
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<mods:url id="110FFAFAEED9BBEB6EBD6BC78D9DD0CE">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="94DCC136623552011578D0DDD9970BC5">journal article</mods:classification>
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<mods:identifier id="7C233BB0E04C814DD3B5B67A2FB723F3" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="448766738DB7879FF4D0D01875A5C810" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55BE3FF14FF79F1F9FE50FDAA" ID-DOI="http://doi.org/10.5281/zenodo.5291817" ID-Zenodo-Dep="5291817" LSID="urn:lsid:plazi:treatment:03B087E55BE3FF14FF79F1F9FE50FDAA" httpUri="http://treatment.plazi.org/id/03B087E55BE3FF14FF79F1F9FE50FDAA" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135">
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<subSubSection id="C30365785BE3FF15FF79F1F9FD9AFC7B" box="[151,556,979,1006]" pageId="134" pageNumber="135" type="nomenclature">
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<paragraph id="8BA636F35BE3FF15FF79F1F9FD9AFC7B" blockId="134.[151,556,979,1038]" box="[151,556,979,1006]" pageId="134" pageNumber="135">
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<heading id="D0EE819F5BE3FF15FF79F1F9FD9AFC7B" bold="true" box="[151,556,979,1006]" fontSize="11" level="1" pageId="134" pageNumber="135" reason="1">
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<emphasis id="B96DEAE15BE3FF15FF79F1F9FD9AFC7B" bold="true" box="[151,556,979,1006]" pageId="134" pageNumber="135">
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<taxonomicName id="4C194D705BE3FF15FF79F1F9FE3BFC78" ID-CoL="3N3JM" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,397,980,1006]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="tholasoris" status="sp. nov.">
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<emphasis id="B96DEAE15BE3FF15FF79F1F9FE3BFC78" bold="true" box="[151,397,980,1006]" italics="true" pageId="134" pageNumber="135">Hydraena tholasoris</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5BE3FF15FE72F1FEFD9AFC7B" box="[412,556,979,1005]" pageId="134" pageNumber="135" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785BE3FF15FF79F1DBFE34FB98" box="[151,386,1014,1038]" pageId="134" pageNumber="135" type="description">
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<paragraph id="8BA636F35BE3FF15FF79F1DBFE34FB98" blockId="134.[151,556,979,1038]" box="[151,386,1014,1038]" pageId="134" pageNumber="135">
|
||
(
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<figureCitation id="13222A765BE3FF15FF71F1DBFEBAFB98" box="[159,268,1014,1038]" captionStart="FIGURE 250" captionStartId="332.[151,250,1731,1754]" captionTargetBox="[207,1376,215,1696]" captionTargetId="figure-29@332.[186,1401,193,1710]" captionTargetPageId="332" captionText="FIGURE 250. Hydraena tholasoris, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283900" httpUri="https://zenodo.org/record/5283900/files/figure.png" pageId="134" pageNumber="135">Figs. 250</figureCitation>
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,
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<figureCitation id="13222A765BE3FF15FEF8F1DBFEF5FB98" box="[278,323,1014,1038]" captionStart="FIGURES 252–253" captionStartId="334.[151,264,1650,1673]" captionText="FIGURES 252–253. Aedeagi of holotypes. 252. H. tholasoris. 253. H. hornabrooki." pageId="134" pageNumber="135">252</figureCitation>
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,
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<figureCitation id="13222A765BE3FF15FEA0F1DBFECFFB98" box="[334,377,1014,1038]" captionStart="FIGURES 547–550" captionStartId="412.[151,264,1650,1673]" captionTargetBox="[165,1421,231,1564]" captionTargetId="figure-23@412.[165,1421,193,1629]" captionTargetPageId="412" captionText="FIGURES 547–550. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284145" httpUri="https://zenodo.org/record/5284145/files/figure.png" pageId="134" pageNumber="135">547</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785BE3FF15FF79F612FDDDFB08" pageId="134" pageNumber="135" type="materials_examined">
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<paragraph id="8BA636F35BE3FF15FF79F612FDDDFB08" blockId="134.[151,1436,1086,2010]" pageId="134" pageNumber="135">
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Type Material.
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<materialsCitation id="3B713CAE5BE3FF15FEBBF612FDDDFB08" collectingDate="1965-01-27" collectingDateMax="1965-01-30" collectingDateMin="1965-01-27" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="305" latitude="-5.9833336" location="Gusap" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" pageId="134" pageNumber="135" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
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<typeStatus id="54A288515BE3FF15FEBBF612FE74FBC1" box="[341,450,1087,1111]" pageId="134" pageNumber="135" type="holotype">
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<emphasis id="B96DEAE15BE3FF15FEBBF612FE74FBC1" bold="true" box="[341,450,1087,1111]" pageId="134" pageNumber="135">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15BE3FF15FDDEF612FCBDFBC1" bold="true" box="[560,779,1086,1111]" pageId="134" pageNumber="135">
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||
<collectingRegion id="49DDF8115BE3FF15FDDEF612FCB1FBC1" box="[560,775,1086,1111]" country="Papua New Guinea" name="Morobe" pageId="134" pageNumber="135">Morobe Province</collectingRegion>
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:
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</emphasis>
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<collectingMunicipality id="6BC2AC895BE3FF15FCF1F613FC69FBC0" box="[799,991,1086,1110]" pageId="134" pageNumber="135">Markham Valley</collectingMunicipality>
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,
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<location id="8EC660285BE3FF15FC03F613FB81FBC0" LSID="urn:lsid:plazi:treatment:03B087E55BE3FF14FF79F1F9FE50FDAA:8EC660285BE3FF15FC03F613FB81FBC0" box="[1005,1079,1086,1110]" latitude="-5.9833336" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" name="Gusap" pageId="134" pageNumber="135" stateProvince="Morobe">Gusap</location>
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, c.
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<locationDeviation id="9A142FD15BE3FF15FB8DF613FAE1FBC0" box="[1123,1367,1086,1110]" location="Lae" pageId="134" pageNumber="135">
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<quantity id="4CE19B165BE3FF15FB8DF613FB70FBC0" box="[1123,1222,1086,1110]" metricMagnitude="5" metricUnit="m" metricValue="1.4484096" pageId="134" pageNumber="135" unit="mi" value="90.0">90 miles</quantity>
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NW of Lae
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</locationDeviation>
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, large pond, muddy bottom with thick layer of dead leaves, dense
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<taxonomicName id="4C194D705BE3FF15FCA4F64EFC44FBED" box="[842,1010,1123,1147]" class="Magnoliopsida" family="Ceratophyllaceae" genus="Ceratophyllum" kingdom="Plantae" order="Ceratophyllales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="genus">
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<emphasis id="B96DEAE15BE3FF15FCA4F64EFC44FBED" box="[842,1010,1123,1147]" italics="true" pageId="134" pageNumber="135">Ceratophyllum</emphasis>
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</taxonomicName>
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,
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<quantity id="4CE19B165BE3FF15FBEDF64FFBF8FBEC" box="[1027,1102,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="134" pageNumber="135" unit="m" value="305.0">
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||
<elevation id="0034D1C05BE3FF15FBEDF64FFBF8FBEC" box="[1027,1102,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="134" pageNumber="135" unit="m" value="305.0">305 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345BE3FF15FBB2F64FFB0CFBEC" box="[1116,1210,1122,1146]" degrees="5" direction="south" minutes="59" orientation="latitude" pageId="134" pageNumber="135" precision="925" value="-5.9833336">5° 59' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345BE3FF15FB27F64FFA81FBEC" box="[1225,1335,1122,1146]" degrees="146" direction="east" minutes="5" orientation="longitude" pageId="134" pageNumber="135" precision="925" value="146.08333">146° 5' E</geoCoordinate>
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,
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<date id="FFA710335BE3FF15FAABF64FFF67FB08" pageId="134" pageNumber="135" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">
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||
<collectingDate id="EFE3E9DB5BE3FF15FAABF64FFF67FB08" pageId="134" pageNumber="135" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">27–30 i 1965</collectingDate>
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</date>
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,
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<collectorName id="26EC53255BE3FF15FF33F6AAFE34FB08" box="[221,386,1158,1182]" pageId="134" pageNumber="135">M. E. Bacchus</collectorName>
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(MEB 162) (
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||
<collectionCode id="ED08AE365BE3FF15FDF6F6ABFDE9FB08" box="[536,607,1158,1182]" pageId="134" pageNumber="135">NHM</collectionCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785BE3FF15FF29F687FC76FAE0" pageId="134" pageNumber="135" type="diagnosis">
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<paragraph id="8BA636F35BE3FF15FF29F687FC76FAE0" blockId="134.[151,1436,1086,2010]" pageId="134" pageNumber="135">
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<emphasis id="B96DEAE15BE3FF15FF29F687FE63FB55" bold="true" box="[199,469,1194,1219]" pageId="134" pageNumber="135">Differential Diagnosis.</emphasis>
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Among members of the
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<emphasis id="B96DEAE15BE3FF15FD14F686FCD9FB55" box="[762,879,1195,1219]" italics="true" pageId="134" pageNumber="135">Fundextra</emphasis>
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group, similar to
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<taxonomicName id="4C194D705BE3FF15FBACF686FB64FB55" authorityName="Perkins" authorityYear="2011" box="[1090,1234,1195,1219]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="fundextra">
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<emphasis id="B96DEAE15BE3FF15FBACF686FBE0FB54" box="[1090,1110,1195,1218]" italics="true" pageId="134" pageNumber="135">H</emphasis>
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.
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<emphasis id="B96DEAE15BE3FF15FB89F686FB64FB55" box="[1127,1234,1195,1219]" italics="true" pageId="134" pageNumber="135">fundextra</emphasis>
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</taxonomicName>
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in comparatively large size (ca. 1.50 and 1.54 respectively) and in dorsal sculpture (
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<figureCitation id="13222A765BE3FF15FC6DF6E3FC46FB70" box="[899,1008,1230,1254]" captionStart="FIGURE 242" captionStartId="326.[151,250,1753,1776]" captionTargetBox="[236,1353,218,1722]" captionTargetId="figure-28@326.[214,1372,193,1732]" captionTargetPageId="326" captionText="FIGURE 242. Hydraena fundextra, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283888" httpUri="https://zenodo.org/record/5283888/files/figure.png" pageId="134" pageNumber="135">Figs. 242</figureCitation>
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,
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<figureCitation id="13222A765BE3FF15FC12F6E3FB91FB70" box="[1020,1063,1230,1254]" captionStart="FIGURE 250" captionStartId="332.[151,250,1731,1754]" captionTargetBox="[207,1376,215,1696]" captionTargetId="figure-29@332.[186,1401,193,1710]" captionTargetPageId="332" captionText="FIGURE 250. Hydraena tholasoris, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283900" httpUri="https://zenodo.org/record/5283900/files/figure.png" pageId="134" pageNumber="135">250</figureCitation>
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||
).
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<taxonomicName id="4C194D705BE3FF15FBD3F6E3FA93FB70" authorityName="Perkins" authorityYear="2011" box="[1085,1317,1230,1254]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="tholasoris">
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<emphasis id="B96DEAE15BE3FF15FBD3F6E3FA93FB70" box="[1085,1317,1230,1254]" italics="true" pageId="134" pageNumber="135">Hydraena tholasoris</emphasis>
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</taxonomicName>
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has a pronotal fascia whereas
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<taxonomicName id="4C194D705BE3FF15FE6FF6DEFDB8FA9D" authorityName="Perkins" authorityYear="2011" box="[385,526,1267,1291]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="fundextra">
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||
<emphasis id="B96DEAE15BE3FF15FE6FF6DEFE23FA9C" box="[385,405,1267,1290]" italics="true" pageId="134" pageNumber="135">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15BE3FF15FE4DF6DEFDB8FA9D" box="[419,526,1267,1291]" italics="true" pageId="134" pageNumber="135">fundextra</emphasis>
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</taxonomicName>
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has a macula. The plaque ratios differ in the two species, with
|
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<taxonomicName id="4C194D705BE3FF15FB3FF6DEFAD5FA9D" authorityName="Perkins" authorityYear="2011" box="[1233,1379,1267,1291]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="tholasoris">
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<emphasis id="B96DEAE15BE3FF15FB3FF6DEFB53FA9C" box="[1233,1253,1267,1290]" italics="true" pageId="134" pageNumber="135">H</emphasis>
|
||
.
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<emphasis id="B96DEAE15BE3FF15FB1DF6DEFAD5FA9D" box="[1267,1379,1267,1291]" italics="true" pageId="134" pageNumber="135">tholasoris</emphasis>
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</taxonomicName>
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having shorter and less widely separated plaques compared with P2 (ratios ca. 5/1/7/6 vs. 4/1/8/7); in addition, the plaques are anteriorly convergent in
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<taxonomicName id="4C194D705BE3FF15FDDCF716FD73FAC5" authorityName="Perkins" authorityYear="2011" box="[562,709,1339,1363]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="tholasoris">
|
||
<emphasis id="B96DEAE15BE3FF15FDDCF716FDF0FAC4" box="[562,582,1339,1362]" italics="true" pageId="134" pageNumber="135">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15BE3FF15FDBBF716FD73FAC5" box="[597,709,1339,1363]" italics="true" pageId="134" pageNumber="135">tholasoris</emphasis>
|
||
</taxonomicName>
|
||
and parallel in
|
||
<taxonomicName id="4C194D705BE3FF15FC9BF716FBB5FAC5" authorityName="Perkins" authorityYear="2011" box="[885,1027,1339,1363]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="fundextra">
|
||
<emphasis id="B96DEAE15BE3FF15FC9BF716FC3FFAC4" box="[885,905,1339,1362]" italics="true" pageId="134" pageNumber="135">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15BE3FF15FC76F716FBB5FAC5" box="[920,1027,1339,1363]" italics="true" pageId="134" pageNumber="135">fundextra</emphasis>
|
||
</taxonomicName>
|
||
. The male genitalia of
|
||
<taxonomicName id="4C194D705BE3FF15FAE7F716FA2AFAC5" authorityName="Perkins" authorityYear="2011" box="[1289,1436,1339,1363]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="tholasoris">
|
||
<emphasis id="B96DEAE15BE3FF15FAE7F716FAABFAC4" box="[1289,1309,1339,1362]" italics="true" pageId="134" pageNumber="135">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15BE3FF15FAC2F716FA2AFAC5" box="[1324,1436,1339,1363]" italics="true" pageId="134" pageNumber="135">tholasoris</emphasis>
|
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</taxonomicName>
|
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distinctively differ from those of other members of the group (
|
||
<figureCitation id="13222A765BE3FF15FCBFF773FC05FAE0" box="[849,947,1374,1398]" captionStart="FIGURES 252–253" captionStartId="334.[151,264,1650,1673]" captionText="FIGURES 252–253. Aedeagi of holotypes. 252. H. tholasoris. 253. H. hornabrooki." pageId="134" pageNumber="135">Fig. 252</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785BE3FF14FF29F7AFFC90FE59" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135" type="description">
|
||
<paragraph id="8BA636F35BE3FF15FF29F7AFFB65F990" blockId="134.[151,1436,1086,2010]" pageId="134" pageNumber="135">
|
||
<emphasis id="B96DEAE15BE3FF15FF29F7AFFEEEFA0D" bold="true" box="[199,344,1410,1435]" inLineHeading="true" pageId="134" pageNumber="135" reason="1">Description.</emphasis>
|
||
Size:
|
||
<typeStatus id="54A288515BE3FF15FE42F7AFFDB8FA0C" box="[428,526,1410,1434]" pageId="134" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.50/0.67; head 0.24/0.37; pronotum 0.39/0.53, PA 0.41, PB 0.53; elytra 0.88/0.67. Dorsum of head dark brown; pronotum testaceous in front of and behind diffusely margined, piceous fascia, ratios of color bands, as measured in midline, ca. 6/15/5; elytra brown; legs light brown; maxillary palpi testaceous, tip not darker. Dorsum very weakly shining.
|
||
</paragraph>
|
||
<paragraph id="8BA636F35BE3FF15FF29F43EFCEBF8B0" blockId="134.[151,1436,1086,2010]" pageId="134" pageNumber="135">Frons punctures ca. 1–2xef, slightly smaller and sparser medially; interstices effacedly microreticulate, dull, 0.5–2xpd. Clypeus effacedly microreticulate, very finely sparsely punctulate medially. Mentum very sparsely very finely punctulate, shining. Postmentum effacedly microreticulate in shallow median depression, otherwise finely sparsely punctate. Genae very slightly raised, shining, with shallow, obsolete median impression, without posterior ridge. Pronotum transverse, ca. median 3/4 of anterior margin slightly arcuate to posterior, sides arcuate, slightly emarginate between midlength and posterior angle; punctures on disc much larger than largest frons punctures, interstices strongly shining, ca. 1–3xpd, punctures anteriorly and posteriorly slightly larger than those on disc; PF1 and PF4 absent; PF2 very shallow; PF3 moderately deep, wide.</paragraph>
|
||
<paragraph id="8BA636F35BE3FF15FF29F51FFE78F84C" blockId="134.[151,1436,1086,2010]" pageId="134" pageNumber="135">Elytra with posterior declivity very gradual, summit at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
|
||
<paragraph id="8BA636F35BE2FF14FF29F2BAFC90FE59" blockId="135.[151,1437,151,572]" pageId="135" pageNumber="136">
|
||
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 5/1/7/6. P1 slightly wider than laminate; median carina nearly straight in profile, very slightly arcuate between coxae. P2 markedly concave, length/width ca. 7/5, sides very slightly diverging toward blunt apex. Plaques narrow lines, very slightly raised posteriorly, tapering to points anteriorly, converging slightly anteriorly, at sides of densely punctate median depression. Metaventrite between mesocoxae slightly sloping, confluent with tip of P2. AIS width at straight posterior margin slightly greater than P2. All legs of moderate length. Profemur (male) without tubercle on ventral margin; protibia very slightly arcuate, gradually increasing in width from base to apex, posteromedial margin with small, low carina at about midlength. Meso- and metatibia straight, slender. Abdominal apex asymmetrical; last tergite (male) with deep concavity near apex, offset to left side. Aedeagus as illustrated (
|
||
<figureCitation id="13222A765BE2FF14FD59F39AFCAFFE59" box="[695,793,439,463]" captionStart="FIGURES 252–253" captionStartId="334.[151,264,1650,1673]" captionText="FIGURES 252–253. Aedeagi of holotypes. 252. H. tholasoris. 253. H. hornabrooki." pageId="135" pageNumber="136">Fig. 252</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785BE2FF14FF29F3F1FB13FE65" box="[199,1189,476,500]" pageId="135" pageNumber="136" type="etymology">
|
||
<paragraph id="8BA636F35BE2FF14FF29F3F1FB13FE65" blockId="135.[151,1437,151,572]" box="[199,1189,476,500]" pageId="135" pageNumber="136">
|
||
<emphasis id="B96DEAE15BE2FF14FF29F3F1FEFBFE62" bold="true" box="[199,333,476,500]" inLineHeading="true" pageId="135" pageNumber="136" reason="1">Etymology.</emphasis>
|
||
Judging from the male genitalia this species is the sister species of
|
||
<taxonomicName id="4C194D705BE2FF14FBACF3F0FB2BFE62" authorityName="Perkins" authorityYear="2011" box="[1090,1181,476,500]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="thola">
|
||
<emphasis id="B96DEAE15BE2FF14FBACF3F0FBE0FE62" box="[1090,1110,477,500]" italics="true" pageId="135" pageNumber="136">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15BE2FF14FB8AF3F1FB2BFE62" box="[1124,1181,476,500]" italics="true" pageId="135" pageNumber="136">thola</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785BE2FF14FF29F3D2FE50FDAA" pageId="135" pageNumber="136" type="distribution">
|
||
<paragraph id="8BA636F35BE2FF14FF29F3D2FE50FDAA" blockId="135.[151,1437,151,572]" pageId="135" pageNumber="136">
|
||
<emphasis id="B96DEAE15BE2FF14FF29F3D2FED6FD8E" bold="true" box="[199,352,511,536]" inLineHeading="true" pageId="135" pageNumber="136" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515BE2FF14FD27F02DFD4FFD8E" box="[713,761,512,536]" pageId="135" pageNumber="136">type</typeStatus>
|
||
locality, Markham Valley, Gusap, in the eastern arm of Area 3; elevation
|
||
<quantity id="4CE19B165BE2FF14FECFF009FEDEFDAD" box="[289,360,548,572]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="135" pageNumber="136" unit="m" value="305.0">305 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765BE2FF14FE99F009FE6EFDAA" box="[375,472,548,572]" captionStart="FIGURES 547–550" captionStartId="412.[151,264,1650,1673]" captionTargetBox="[165,1421,231,1564]" captionTargetId="figure-23@412.[165,1421,193,1629]" captionTargetPageId="412" captionText="FIGURES 547–550. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284145" httpUri="https://zenodo.org/record/5284145/files/figure.png" pageId="135" pageNumber="136">Fig. 547</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |