208 lines
26 KiB
XML
208 lines
26 KiB
XML
<document id="89273FB4A5C06B6A2ADED8EA18D7D386" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B51FFA6FF79F7CBFC8CFB36" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena biundulata Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="54" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="53" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7BE970EB2A369F263EE39D7E39AE7106">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="A87A38E92823015AE008AB3CDC841255">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="141800985BD71748CF7E2BEE84083080">text</mods:typeOfResource>
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<mods:title id="7E3F2CA60FE7D934B713A25ABCF5DD14">Zootaxa</mods:title>
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<mods:part id="A73B9362E09D7D6B4C0663A61C77C84B">
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<mods:date id="DE20DEF7AE22762413767CFC869DEB75">2011</mods:date>
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<mods:number id="AC759E70DD78A892FD05DF65D90B2B09">2011-07-08</mods:number>
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<mods:number id="024AA51CFF51F4E5DFB1B68D20278088">2944</mods:number>
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<mods:url id="001C1F4BA77A00840DBD6C03777AE888">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="82A3E2A8249D97F190E3E1A0F9A83CC2">journal article</mods:classification>
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<mods:identifier id="B599EC6FB7623FE0C1DEAE6945F37AEA" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="176F05A53B2B09DDDBDDC7CA0D39A8CF" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55B51FFA6FF79F7CBFC8CFB36" ID-DOI="http://doi.org/10.5281/zenodo.5302728" ID-Zenodo-Dep="5302728" LSID="urn:lsid:plazi:treatment:03B087E55B51FFA6FF79F7CBFC8CFB36" httpUri="http://treatment.plazi.org/id/03B087E55B51FFA6FF79F7CBFC8CFB36" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
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<subSubSection id="C30365785B51FFA7FF79F7CBFD8FF996" box="[151,569,1510,1536]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="8BA636F35B51FFA7FF79F7CBFD8FF996" blockId="52.[151,569,1510,1569]" box="[151,569,1510,1536]" pageId="52" pageNumber="53">
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<heading id="D0EE819F5B51FFA7FF79F7CBFD8FF996" bold="true" box="[151,569,1510,1536]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<emphasis id="B96DEAE15B51FFA7FF79F7CBFD8FF996" bold="true" box="[151,569,1510,1536]" pageId="52" pageNumber="53">
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<taxonomicName id="4C194D705B51FFA7FF79F7CBFE2CF996" ID-CoL="3N36L" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,410,1510,1536]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="biundulata" status="sp. nov.">
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<emphasis id="B96DEAE15B51FFA7FF79F7CBFE2CF996" bold="true" box="[151,410,1510,1536]" italics="true" pageId="52" pageNumber="53">Hydraena biundulata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B51FFA7FE47F7CBFD8FF996" box="[425,569,1510,1536]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30365785B51FFA7FF79F424FDB8F9B7" box="[151,526,1545,1569]" pageId="52" pageNumber="53" type="description">
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<paragraph id="8BA636F35B51FFA7FF79F424FDB8F9B7" blockId="52.[151,569,1510,1569]" box="[151,526,1545,1569]" pageId="52" pageNumber="53">
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(
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<figureCitation id="13222A765B51FFA7FF71F424FF4BF9B7" box="[159,253,1545,1569]" captionStart="FIGURE 75" captionStartId="201.[151,250,1753,1776]" captionTargetBox="[234,1356,209,1718]" captionTargetId="figure-26@201.[220,1368,193,1732]" captionTargetPageId="201" captionText="FIGURE 75. Hydraena biundulata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283547" httpUri="https://zenodo.org/record/5283547/files/figure.png" pageId="52" pageNumber="53">Figs. 75</figureCitation>
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,
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<figureCitation id="13222A765B51FFA7FEE6F427FE90F9B7" box="[264,294,1546,1569]" captionStart="FIGURES 76–77" captionStartId="202.[151,264,1650,1673]" captionTargetBox="[300,1270,221,1616]" captionTargetId="figure-32@202.[278,1309,193,1629]" captionTargetPageId="202" captionText="FIGURES 76–77. Aedeagi of holotypes. 76. H. insandalia. 77. H. biundulata." figureDoi="http://doi.org/10.5281/zenodo.5283551" httpUri="https://zenodo.org/record/5283551/files/figure.png" pageId="52" pageNumber="53">77</figureCitation>
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,
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<figureCitation id="13222A765B51FFA7FEDCF424FE20F9B7" box="[306,406,1545,1569]" captionStart="FIGURES 280–285" captionStartId="351.[151,264,1650,1673]" captionTargetBox="[170,1416,199,1621]" captionTargetId="figure-26@351.[166,1420,193,1629]" captionTargetPageId="351" captionText="FIGURES 280–285. Scanning electron micrographs of H. manulea (280–283) and H. biundulata (284–285)." figureDoi="http://doi.org/10.5281/zenodo.5283964" httpUri="https://zenodo.org/record/5283964/files/figure.png" pageId="52" pageNumber="53">284, 285</figureCitation>
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,
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<figureCitation id="13222A765B51FFA7FE4CF424FE78F9B7" box="[418,462,1545,1569]" captionStart="FIGURES 396–399" captionStartId="371.[151,264,1650,1673]" captionText="FIGURES 396–399. Female tergite X, gonocoxite, and spermatheca of Hydraena species." pageId="52" pageNumber="53">397</figureCitation>
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,
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<figureCitation id="13222A765B51FFA7FE34F424FDB3F9B7" box="[474,517,1545,1569]" captionStart="FIGURES 467–470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467–470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="52" pageNumber="53">468</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B51FFA7FF79F47CFE2AF927" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="8BA636F35B51FFA7FF79F47CFE2AF927" blockId="52.[151,1436,1617,2037]" pageId="52" pageNumber="53">
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Type Material.
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<materialsCitation id="3B713CAE5B51FFA7FEBEF47CFB25F91B" collectingDate="1964-12-28" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1000" latitude="-6.866667" location="Lae - Bulolo" longLatPrecision="1303" longitude="146.61667" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
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<typeStatus id="54A288515B51FFA7FEBEF47CFE0BF9FF" box="[336,445,1617,1641]" pageId="52" pageNumber="53" type="holotype">
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<emphasis id="B96DEAE15B51FFA7FEBEF47CFE0BF9FF" bold="true" box="[336,445,1617,1641]" pageId="52" pageNumber="53">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B51FFA7FDCAF47CFD4BF9FC" bold="true" box="[548,765,1617,1642]" pageId="52" pageNumber="53">
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<collectingRegion id="49DDF8115B51FFA7FDCAF47CFD41F9FC" box="[548,759,1617,1642]" country="Papua New Guinea" name="Morobe" pageId="52" pageNumber="53">Morobe Province</collectingRegion>
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:
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</emphasis>
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<location id="8EC660285B51FFA7FCE2F47CFC26F9FF" LSID="urn:lsid:plazi:treatment:03B087E55B51FFA6FF79F7CBFC8CFB36:8EC660285B51FFA7FCE2F47CFC26F9FF" box="[780,912,1617,1641]" latitude="-6.866667" longLatPrecision="1303" longitude="146.61667" name="Lae - Bulolo" pageId="52" pageNumber="53" stateProvince="Morobe">Lae–Bulolo</location>
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road, ex stones and gravel bank of large river, [GE est.]
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<quantity id="4CE19B165B51FFA7FEECF458FE39F91B" box="[258,399,1653,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="52" pageNumber="53" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">
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<elevation id="0034D1C05B51FFA7FEECF458FE39F91B" box="[258,399,1653,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="52" pageNumber="53" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B51FFA7FE74F458FE44F91B" box="[410,498,1653,1677]" degrees="6" direction="south" minutes="52" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="-6.866667">6° 52' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B51FFA7FE10F458FDC2F91B" box="[510,628,1653,1677]" degrees="146" direction="east" minutes="37" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="146.61667">146° 37' E</geoCoordinate>
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,
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<date id="FFA710335B51FFA7FD6EF458FCB7F91B" box="[640,769,1653,1677]" pageId="52" pageNumber="53" value="1964-12-28">
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<collectingDate id="EFE3E9DB5B51FFA7FD6EF458FCB7F91B" box="[640,769,1653,1677]" pageId="52" pageNumber="53" value="1964-12-28">28 xii 1964</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B51FFA7FCE5F45BFC19F91B" box="[779,943,1653,1677]" pageId="52" pageNumber="53">M. E. Bacchus</collectorName>
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(MEB 124) (
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<collectionCode id="ED08AE365B51FFA7FBADF458FB3CF91B" box="[1091,1162,1653,1677]" pageId="52" pageNumber="53">NHM</collectionCode>
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||
)
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</materialsCitation>
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.
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<materialsCitation id="3B713CAE5B51FFA7FB72F458FE2EF927" collectingDate="1964-12-28" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1000" latitude="-6.866667" location="Lae - Bulolo" longLatPrecision="1303" longitude="146.61667" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
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<emphasis id="B96DEAE15B51FFA7FB72F458FAA2F91B" bold="true" box="[1180,1300,1653,1677]" pageId="52" pageNumber="53">
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<typeStatus id="54A288515B51FFA7FB72F458FAA2F91B" box="[1180,1300,1653,1677]" pageId="52" pageNumber="53" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: Same data as holotype (11
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<collectionCode id="ED08AE365B51FFA7FEA7F4B4FE26F927" box="[329,400,1689,1713]" pageId="52" pageNumber="53">NHM</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B51FFA7FF29F490FEBCF83B" pageId="52" pageNumber="53" type="diagnosis">
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<paragraph id="8BA636F35B51FFA7FF29F490FEBCF83B" blockId="52.[151,1436,1617,2037]" pageId="52" pageNumber="53">
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<emphasis id="B96DEAE15B51FFA7FF29F490FE62F940" bold="true" box="[199,468,1725,1750]" pageId="52" pageNumber="53">Differential Diagnosis.</emphasis>
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Among members of the
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<taxonomicName id="4C194D705B51FFA7FD1AF493FCF4F943" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[756,834,1726,1749]" class="Insecta" family="Tettigoniidae" genus="Tetana" kingdom="Animalia" order="Orthoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="B96DEAE15B51FFA7FD1AF493FCF4F943" box="[756,834,1726,1749]" italics="true" pageId="52" pageNumber="53">Tetana</emphasis>
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</taxonomicName>
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group, somewhat similar in habitus to
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<taxonomicName id="4C194D705B51FFA7FAECF493FA2DF943" authorityName="Perkins" authorityYear="2011" box="[1282,1435,1725,1749]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insandalia">
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<emphasis id="B96DEAE15B51FFA7FAECF493FAA0F943" box="[1282,1302,1726,1749]" italics="true" pageId="52" pageNumber="53">H</emphasis>
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||
.
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<emphasis id="B96DEAE15B51FFA7FACBF490FA2DF943" box="[1317,1435,1725,1749]" italics="true" pageId="52" pageNumber="53">insandalia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C194D705B51FFA7FF28F4CFFEDBF96F" authorityName="Perkins" authorityYear="2011" box="[198,365,1761,1785]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="penultimata">
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<emphasis id="B96DEAE15B51FFA7FF28F4CFFF6CF96F" box="[198,218,1762,1785]" italics="true" pageId="52" pageNumber="53">H</emphasis>
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||
.
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<emphasis id="B96DEAE15B51FFA7FF09F4CCFEDBF96F" box="[231,365,1761,1785]" italics="true" pageId="52" pageNumber="53">penultimata</emphasis>
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</taxonomicName>
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||
(
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||
<figureCitation id="13222A765B51FFA7FE92F4CCFE6CF96F" box="[380,474,1761,1785]" captionStart="FIGURE 74" captionStartId="200.[151,250,1701,1724]" captionTargetBox="[186,1401,193,1680]" captionTargetId="figure-28@200.[186,1401,193,1680]" captionTargetPageId="200" captionText="FIGURE 74. Hydraena insandalia, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283545" httpUri="https://zenodo.org/record/5283545/files/figure.png" pageId="52" pageNumber="53">Figs. 74</figureCitation>
|
||
,
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||
<figureCitation id="13222A765B51FFA7FE0AF4CCFDB4F96F" box="[484,514,1761,1785]" captionStart="FIGURE 75" captionStartId="201.[151,250,1753,1776]" captionTargetBox="[234,1356,209,1718]" captionTargetId="figure-26@201.[220,1368,193,1732]" captionTargetPageId="201" captionText="FIGURE 75. Hydraena biundulata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283547" httpUri="https://zenodo.org/record/5283547/files/figure.png" pageId="52" pageNumber="53">75</figureCitation>
|
||
,
|
||
<figureCitation id="13222A765B51FFA7FDE0F4CCFD9BF96F" box="[526,557,1761,1785]" captionStart="FIGURE 78" captionStartId="203.[151,250,1690,1713]" captionTargetBox="[186,1401,193,1669]" captionTargetId="figure-26@203.[186,1401,193,1669]" captionTargetPageId="203" captionText="FIGURE 78. Hydraena penultimata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283555" httpUri="https://zenodo.org/record/5283555/files/figure.png" pageId="52" pageNumber="53">78</figureCitation>
|
||
); differing from both compared species by the shape of the pronotum, which is more widely produced in the middle, and nearly equally sinuate in front of and behind the widest part. Additionally, the males of
|
||
<taxonomicName id="4C194D705B51FFA7FEC9F507FE74F8D7" authorityName="Perkins" authorityYear="2011" box="[295,450,1833,1857]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="biundulata">
|
||
<emphasis id="B96DEAE15B51FFA7FEC9F507FE8DF8D7" box="[295,315,1834,1857]" italics="true" pageId="52" pageNumber="53">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B51FFA7FEA7F504FE74F8D7" box="[329,450,1833,1857]" italics="true" pageId="52" pageNumber="53">biundulata</emphasis>
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||
</taxonomicName>
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have tibiae that are shaped differently than the compared species. The male genitalia of the three species are quite distinct, with
|
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<taxonomicName id="4C194D705B51FFA7FDB0F563FD4CF8F3" authorityName="Perkins" authorityYear="2011" box="[606,762,1869,1893]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="biundulata">
|
||
<emphasis id="B96DEAE15B51FFA7FDB0F563FDC4F8F3" box="[606,626,1870,1893]" italics="true" pageId="52" pageNumber="53">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B51FFA7FD6FF560FD4CF8F3" box="[641,762,1869,1893]" italics="true" pageId="52" pageNumber="53">biundulata</emphasis>
|
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</taxonomicName>
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||
being more similar to
|
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<taxonomicName id="4C194D705B51FFA7FC12F563FB23F8F3" authorityName="Perkins" authorityYear="2011" box="[1020,1173,1869,1893]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insandalia">
|
||
<emphasis id="B96DEAE15B51FFA7FC12F563FBA6F8F3" box="[1020,1040,1870,1893]" italics="true" pageId="52" pageNumber="53">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B51FFA7FBF1F560FB23F8F3" box="[1055,1173,1869,1893]" italics="true" pageId="52" pageNumber="53">insandalia</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C194D705B51FFA7FB1DF563FA2AF8F3" authorityName="Perkins" authorityYear="2011" box="[1267,1436,1869,1893]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="penultimata">
|
||
<emphasis id="B96DEAE15B51FFA7FB1DF563FAB1F8F3" box="[1267,1287,1870,1893]" italics="true" pageId="52" pageNumber="53">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B51FFA7FAF8F560FA2AF8F3" box="[1302,1436,1869,1893]" italics="true" pageId="52" pageNumber="53">penultimata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13222A765B51FFA7FF71F55CFE90F81F" box="[159,294,1905,1929]" captionStart="FIGURES 76–77" captionStartId="202.[151,264,1650,1673]" captionTargetBox="[300,1270,221,1616]" captionTargetId="figure-32@202.[278,1309,193,1629]" captionTargetPageId="202" captionText="FIGURES 76–77. Aedeagi of holotypes. 76. H. insandalia. 77. H. biundulata." figureDoi="http://doi.org/10.5281/zenodo.5283551" httpUri="https://zenodo.org/record/5283551/files/figure.png" pageId="52" pageNumber="53">Figs. 76, 77</figureCitation>
|
||
,
|
||
<figureCitation id="13222A765B51FFA7FEDEF55CFEE6F81F" box="[304,336,1905,1929]" captionStart="FIGURES 80–81" captionStartId="205.[151,264,1650,1673]" captionTargetBox="[301,1290,208,1616]" captionTargetId="figure-29@205.[276,1311,193,1629]" captionTargetPageId="205" captionText="FIGURES 80–81. Aedeagi of holotypes. 80. H. penultimata. 81. H. acumena." figureDoi="http://doi.org/10.5281/zenodo.5283561" httpUri="https://zenodo.org/record/5283561/files/figure.png" pageId="52" pageNumber="53">80</figureCitation>
|
||
). The metaventrite and the chaetotaxy of the meso- and metatibiae are illustrated with SEMs (
|
||
<figureCitation id="13222A765B51FFA7FA8DF55CFF4BF83B" captionStart="FIGURES 280–285" captionStartId="351.[151,264,1650,1673]" captionTargetBox="[170,1416,199,1621]" captionTargetId="figure-26@351.[166,1420,193,1629]" captionTargetPageId="351" captionText="FIGURES 280–285. Scanning electron micrographs of H. manulea (280–283) and H. biundulata (284–285)." figureDoi="http://doi.org/10.5281/zenodo.5283964" httpUri="https://zenodo.org/record/5283964/files/figure.png" pageId="52" pageNumber="53">Figs. 284, 285</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B51FFA6FF29F594FE33FB99" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
|
||
<paragraph id="8BA636F35B51FFA6FF29F594FC66FF42" blockId="52.[151,1436,1617,2037]" lastBlockId="53.[151,1437,151,1184]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
|
||
<emphasis id="B96DEAE15B51FFA7FF29F594FEEEF844" bold="true" box="[199,344,1977,2002]" inLineHeading="true" pageId="52" pageNumber="53" reason="1">Description.</emphasis>
|
||
Size:
|
||
<typeStatus id="54A288515B51FFA7FE42F594FDB8F847" box="[428,526,1977,2001]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.55/0.63; head 0.26/0.37; pronotum 0.40/0.48, PA 0.40, PB 0.38; elytra 0.90/0.63. Dorsum of head piceous; pronotum light brown in front of and behind piceous, diffusely margined, fascia, ratios of color bands, as measured in midline, ca. 6/15/5; elytra dark brown; legs and maxillary palpi dark brown. Dorsum moderately shining.
|
||
</paragraph>
|
||
<paragraph id="8BA636F35B50FFA6FF29F2CDFCEBFE62" blockId="53.[151,1437,151,1184]" pageId="53" pageNumber="54">Frons punctures ca. 2xef near eyes, smaller and sparser medially; interstices shining, medially 3–7xpd. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and shining medially. Mentum very sparsely very finely punctulate, strongly shining. Postmentum microreticulate in shallow median depression, otherwise moderately coarsely punctate. Genae very slightly raised, shining, with shallow, obsolete anteromedian impression, without posterior ridge. Pronotum subcordiform, distinctively narrowed both in front of and behind widest area; median 2/3 of anterior margin emarginate; punctures on disc slightly smaller than largest frons punctures, interstices shining, ca. 2–5xpd, punctures anteriorly and posteriorly ca. 2xpd those on disc, and denser; PF1 absent; PF2 and PF4 very shallow; PF3 moderately deep, wide.</paragraph>
|
||
<paragraph id="8BA636F35B50FFA6FF29F3D2FD24FD31" blockId="53.[151,1437,151,1184]" pageId="53" pageNumber="54">Elytra widest at about midlength; summit of posterior declivity slightly posterior to midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
|
||
<paragraph id="8BA636F35B50FFA6FF29F099FE33FB99" blockId="53.[151,1437,151,1184]" pageId="53" pageNumber="54">
|
||
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/3/9/4. P1 laminate; median carina sinuate in profile, slightly raised between coxae. P2 length/width ca. 5/3, transversely very slightly concave, sides very slightly diverging toward blunt apex. Plaques large, raised slightly more laterally than medially, at sloping sides of median depression, anteriorly slightly tapering and convergent; each plaque with tiny papilla slightly behind midlength. Metaventrite between mesocoxae very weakly raised in midline. AIS width at straight posterior margin ca. 1.5x P2. All legs rather long and slender. Profemur (male) with tubercle on ventral margin near trochanter; protibia arcuate, increasing in width from near base to subapically. Mesotibia slightly arcuate. Metatibia sinuate, slender except apically widened and bearing few setae. Abdominal apex very slightly asymmetrical, if at all; last tergite (male) with very small apicomedian notch. Aedeagus as illustrated (
|
||
<figureCitation id="13222A765B50FFA6FCE6F1F9FCEAFC7A" box="[776,860,980,1004]" captionStart="FIGURES 76–77" captionStartId="202.[151,264,1650,1673]" captionTargetBox="[300,1270,221,1616]" captionTargetId="figure-32@202.[278,1309,193,1629]" captionTargetPageId="202" captionText="FIGURES 76–77. Aedeagi of holotypes. 76. H. insandalia. 77. H. biundulata." figureDoi="http://doi.org/10.5281/zenodo.5283551" httpUri="https://zenodo.org/record/5283551/files/figure.png" pageId="53" pageNumber="54">Fig. 77</figureCitation>
|
||
). Female tergite X, gonocoxite, and spermatheca as illustrated (
|
||
<figureCitation id="13222A765B50FFA6FEF8F1DAFECEFB99" box="[278,376,1015,1039]" captionStart="FIGURES 396–399" captionStartId="371.[151,264,1650,1673]" captionText="FIGURES 396–399. Female tergite X, gonocoxite, and spermatheca of Hydraena species." pageId="53" pageNumber="54">Fig. 397</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B50FFA6FF29F631FEE5FBC1" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="8BA636F35B50FFA6FF29F631FEE5FBC1" blockId="53.[151,1437,151,1184]" pageId="53" pageNumber="54">
|
||
<emphasis id="B96DEAE15B50FFA6FF29F631FEFBFBA2" bold="true" box="[199,333,1052,1076]" inLineHeading="true" pageId="53" pageNumber="54" reason="1">Etymology.</emphasis>
|
||
Named in reference to the undulating medial surface of the male metatibiae and undulating margin of the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B50FFA6FF29F64EFC8CFB36" pageId="53" pageNumber="54" type="distribution">
|
||
<paragraph id="8BA636F35B50FFA6FF29F64EFC8CFB36" blockId="53.[151,1437,151,1184]" pageId="53" pageNumber="54">
|
||
<emphasis id="B96DEAE15B50FFA6FF29F64EFED6FBEA" bold="true" box="[199,352,1123,1148]" inLineHeading="true" pageId="53" pageNumber="54" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B50FFA6FD21F649FD49FBEA" box="[719,767,1124,1148]" pageId="53" pageNumber="54">type</typeStatus>
|
||
locality on the Lae–Bulolo road, at the eastern end of Area 3 where it borders Area 6; elevation
|
||
<quantity id="4CE19B165B50FFA6FDDFF6A5FD0AFB09" box="[561,700,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="53" pageNumber="54" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B50FFA6FD25F6A5FC98FB36" box="[715,814,1160,1184]" captionStart="FIGURES 467–470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467–470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="53" pageNumber="54">Fig. 468</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |