treatments-xml/data/03/B0/87/03B087E55B2DFFDAFF79F09FFEF2FE62.xml
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<document id="1DB5C47792B11D68BC5E6000B650FADD" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B2DFFDAFF79F09FFEF2FE62" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena okapa Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="74" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="73" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A691B432AF749FA7BA439E7357E3F68B">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
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<mods:namePart id="0E11F35F1A69B06682EEFFA285363AE1">Perkins, Philip D.</mods:namePart>
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<mods:date id="C641E6951EFC60A329DACDA8A1798171">2011</mods:date>
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<mods:number id="5B3A0A811836B286500E451DD5E1FCD5">2011-07-08</mods:number>
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<mods:identifier id="F68FC44D3C46F77DA116B245BD8D5869" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="2EC8D12C45DE6A7BE76B25C6EF015D37" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03B087E55B2DFFDAFF79F09FFEF2FE62" ID-DOI="http://doi.org/10.5281/zenodo.5291596" ID-Zenodo-Dep="5291596" LSID="urn:lsid:plazi:treatment:03B087E55B2DFFDAFF79F09FFEF2FE62" httpUri="http://treatment.plazi.org/id/03B087E55B2DFFDAFF79F09FFEF2FE62" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<subSubSection id="C30365785B2DFFDBFF79F09FFE48FD5A" box="[151,510,690,716]" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="8BA636F35B2DFFDBFF79F09FFE48FD5A" blockId="72.[151,553,690,749]" box="[151,510,690,716]" pageId="72" pageNumber="73">
<heading id="D0EE819F5B2DFFDBFF79F09FFE48FD5A" bold="true" box="[151,510,690,716]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<emphasis id="B96DEAE15B2DFFDBFF79F09FFE48FD5A" bold="true" box="[151,510,690,716]" pageId="72" pageNumber="73">
<taxonomicName id="4C194D705B2DFFDBFF79F09FFEE9FD5A" ID-CoL="6MJCB" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,351,690,716]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="okapa" status="sp. nov.">
<emphasis id="B96DEAE15B2DFFDBFF79F09FFEE9FD5A" bold="true" box="[151,351,690,716]" italics="true" pageId="72" pageNumber="73">Hydraena okapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B2DFFDBFE80F09FFE48FD5A" box="[366,510,690,716]" pageId="72" pageNumber="73" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2DFFDBFF79F0F8FD9FFD7B" box="[151,553,725,749]" pageId="72" pageNumber="73" type="description">
<paragraph id="8BA636F35B2DFFDBFF79F0F8FD9FFD7B" blockId="72.[151,553,690,749]" box="[151,553,725,749]" pageId="72" pageNumber="73">
(
<figureCitation id="13222A765B2DFFDBFF71F0F8FEBCFD7B" box="[159,266,725,749]" captionStart="FIGURE 119" captionStartId="234.[151,250,1753,1776]" captionTargetBox="[263,1338,212,1724]" captionTargetId="figure-29@234.[241,1346,193,1732]" captionTargetPageId="234" captionText="FIGURE 119. Hydraena okapa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283652" httpUri="https://zenodo.org/record/5283652/files/figure.png" pageId="72" pageNumber="73">Figs. 119</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFEF8F0F8FEF4FD7B" box="[278,322,725,749]" captionStart="FIGURES 120121" captionStartId="235.[151,264,1596,1619]" captionTargetBox="[244,1345,239,1485]" captionTargetId="figure-29@235.[226,1360,193,1575]" captionTargetPageId="235" captionText="FIGURES 120121. Aedeagi of holotypes. 120. H. idema. 121. H. okapa." figureDoi="http://doi.org/10.5281/zenodo.5283654" httpUri="https://zenodo.org/record/5283654/files/figure.png" pageId="72" pageNumber="73">121</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFEA0F0F8FE04FD7B" box="[334,434,725,749]" captionStart="FIGURES 322327" captionStartId="358.[151,264,1650,1673]" captionTargetBox="[183,1404,200,1624]" captionTargetId="figure-29@358.[178,1409,193,1629]" captionTargetPageId="358" captionText="FIGURES 322327. Scanning electron micrographs of H. phainops (322323) and H. okapa (324327)." figureDoi="http://doi.org/10.5281/zenodo.5283987" httpUri="https://zenodo.org/record/5283987/files/figure.png" pageId="72" pageNumber="73">324327</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFE53F0F8FE5CFD7B" box="[445,490,725,749]" captionStart="FIGURES 400403" captionStartId="372.[151,264,1650,1673]" captionTargetBox="[271,1319,208,1611]" captionTargetId="figure-28@372.[235,1351,193,1629]" captionTargetPageId="372" captionText="FIGURES 400403. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284025" httpUri="https://zenodo.org/record/5284025/files/figure.png" pageId="72" pageNumber="73">401</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFE1BF0F8FD97FD7B" box="[501,545,725,749]" captionStart="FIGURES 483486" captionStartId="396.[151,264,1650,1673]" captionTargetBox="[165,1421,252,1595]" captionTargetId="figure-23@396.[165,1421,193,1629]" captionTargetPageId="396" captionText="FIGURES 483486. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284099" httpUri="https://zenodo.org/record/5284099/files/figure.png" pageId="72" pageNumber="73">483</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2DFFDBFF79F130FDB9FB9B" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8BA636F35B2DFFDBFF79F130FDB9FB9B" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">
Type Material.
<materialsCitation id="3B713CAE5B2DFFDBFEBBF130FB6EFCCF" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="72" pageNumber="73" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="holotype">
<typeStatus id="54A288515B2DFFDBFEBBF130FE74FCA3" box="[341,450,797,821]" pageId="72" pageNumber="73" type="holotype">
<emphasis id="B96DEAE15B2DFFDBFEBBF130FE74FCA3" bold="true" box="[341,450,797,821]" pageId="72" pageNumber="73">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B2DFFDBFDDEF130FC39FCA0" bold="true" box="[560,911,797,822]" pageId="72" pageNumber="73">
<collectingRegion id="49DDF8115B2DFFDBFDDEF130FC3CFCA0" box="[560,906,797,822]" country="Papua New Guinea" name="Eastern Highlands" pageId="72" pageNumber="73">Eastern Highlands Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B2DFFDBFC4DF130FBEDFCA3" box="[931,1115,797,821]" pageId="72" pageNumber="73">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B2DFFDBFB61F130FB6DFCA3" LSID="urn:lsid:plazi:treatment:03B087E55B2DFFDAFF79F09FFEF2FE62:8EC660285B2DFFDBFB61F130FB6DFCA3" box="[1167,1243,797,821]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="72" pageNumber="73" stateProvince="Eastern Highlands">Okapa</location>
, gravel banks of small swift river,
<quantity id="4CE19B165B2DFFDBFE8CF16CFE0DFCCF" box="[354,443,833,857]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">
<elevation id="0034D1C05B2DFFDBFE8CF16CFE0DFCCF" box="[354,443,833,857]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B2DFFDBFE27F16CFD90FCCF" box="[457,550,833,857]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B2DFFDBFDDBF16CFD07FCCF" box="[565,689,833,857]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B2DFFDBFD50F16CFC9EFCCF" box="[702,808,833,857]" pageId="72" pageNumber="73" value="1965-02-05">
<collectingDate id="EFE3E9DB5B2DFFDBFD50F16CFC9EFCCF" box="[702,808,833,857]" pageId="72" pageNumber="73" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B2DFFDBFCD8F16FFC69FCCF" box="[822,991,833,857]" pageId="72" pageNumber="73">M. E. Bacchus</collectorName>
(MEB 171a) (
<collectionCode id="ED08AE365B2DFFDBFB67F16CFB67FCCF" box="[1161,1233,833,857]" pageId="72" pageNumber="73">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B2DFFDBFB0AF16CFBA6FC37" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="72" pageNumber="73" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
<emphasis id="B96DEAE15B2DFFDBFB0AF16CFAEAFCCF" bold="true" box="[1252,1372,833,857]" pageId="72" pageNumber="73">
<typeStatus id="54A288515B2DFFDBFB0AF16CFAEAFCCF" box="[1252,1372,833,857]" pageId="72" pageNumber="73" type="paratype">Paratypes</typeStatus>
</emphasis>
(96):
<emphasis id="B96DEAE15B2DFFDBFF79F148FE59FCE8" bold="true" box="[151,495,869,894]" pageId="72" pageNumber="73">
<collectingRegion id="49DDF8115B2DFFDBFF79F148FE5FFCE8" box="[151,489,869,894]" country="Papua New Guinea" name="Eastern Highlands" pageId="72" pageNumber="73">Eastern Highlands Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B2DFFDBFE12F148FD19FCEB" box="[508,687,869,893]" pageId="72" pageNumber="73">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B2DFFDBFD33F148FC9FFCEB" LSID="urn:lsid:plazi:treatment:03B087E55B2DFFDAFF79F09FFEF2FE62:8EC660285B2DFFDBFD33F148FC9FFCEB" box="[733,809,869,893]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="72" pageNumber="73" stateProvince="Eastern Highlands">Okapa</location>
, ex dead leaves caught among rocks in small swift river,
<quantity id="4CE19B165B2DFFDBFF79F1A4FF58FC37" box="[151,238,905,929]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">
<elevation id="0034D1C05B2DFFDBFF79F1A4FF58FC37" box="[151,238,905,929]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B2DFFDBFF14F1A4FEE5FC37" box="[250,339,905,929]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B2DFFDBFE8EF1A4FE61FC37" box="[352,471,905,929]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B2DFFDBFE0DF1A4FDFFFC37" box="[483,585,905,929]" pageId="72" pageNumber="73" value="1965-02-05">
<collectingDate id="EFE3E9DB5B2DFFDBFE0DF1A4FDFFFC37" box="[483,585,905,929]" pageId="72" pageNumber="73" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B2DFFDBFDBBF1A7FD4CFC37" box="[597,762,905,929]" pageId="72" pageNumber="73">M. E. Bacchus</collectorName>
(MEB 171b) (11
<collectionCode id="ED08AE365B2DFFDBFC2CF1A4FBBEFC37" box="[962,1032,905,929]" pageId="72" pageNumber="73">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B2DFFDBFBF5F1A4FA9BFC53" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="72" pageNumber="73" specimenCount="1" stateProvince="Eastern Highlands Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895B2DFFDBFBF5F1A4FB66FC37" box="[1051,1232,905,929]" pageId="72" pageNumber="73">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B2DFFDBFB10F1A4FAFCFC37" LSID="urn:lsid:plazi:treatment:03B087E55B2DFFDAFF79F09FFEF2FE62:8EC660285B2DFFDBFB10F1A4FAFCFC37" box="[1278,1354,905,929]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="72" pageNumber="73" stateProvince="Eastern Highlands Province">Okapa</location>
, gravel banks of small swift river,
<quantity id="4CE19B165B2DFFDBFE51F180FDA3FC53" box="[447,533,941,965]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">
<elevation id="0034D1C05B2DFFDBFE51F180FDA3FC53" box="[447,533,941,965]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B2DFFDBFDCFF180FDCFFC53" box="[545,633,941,965]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B2DFFDBFD6BF180FD4DFC53" box="[645,763,941,965]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B2DFFDBFCE8F180FCDDFC53" box="[774,875,941,965]" pageId="72" pageNumber="73" value="1965-02-05">
<collectingDate id="EFE3E9DB5B2DFFDBFCE8F180FCDDFC53" box="[774,875,941,965]" pageId="72" pageNumber="73" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B2DFFDBFC98F183FBACFC53" box="[886,1050,941,965]" pageId="72" pageNumber="73">M. E. Bacchus</collectorName>
(MEB 171a) (71
<collectionCode id="ED08AE365B2DFFDBFB31F180FA93FC53" box="[1247,1317,941,965]" pageId="72" pageNumber="73">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B2DFFDBFAD9F180FDBDFB9B" collectingDate="1965-02-05" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1372" latitude="-6.5333333" location="Okapa" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" pageId="72" pageNumber="73" specimenCount="1" stateProvince="Eastern Highlands Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895B2DFFDBFAD9F180FF56FC7F" pageId="72" pageNumber="73">Wanitabi Valley</collectingMunicipality>
, nr.
<location id="8EC660285B2DFFDBFEE1F1FCFEEDFC7F" LSID="urn:lsid:plazi:treatment:03B087E55B2DFFDAFF79F09FFEF2FE62:8EC660285B2DFFDBFEE1F1FCFEEDFC7F" box="[271,347,977,1001]" latitude="-6.5333333" longLatPrecision="1304" longitude="145.61667" municipality="Wanitabi Valley" name="Okapa" pageId="72" pageNumber="73" stateProvince="Eastern Highlands Province">Okapa</location>
, on wet rocks above waterline in small swift river,
<quantity id="4CE19B165B2DFFDBFC75F1FCFC44FC7F" box="[923,1010,977,1001]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">
<elevation id="0034D1C05B2DFFDBFC75F1FCFC44FC7F" box="[923,1010,977,1001]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="72" pageNumber="73" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B2DFFDBFC10F1FCFBEEFC7F" box="[1022,1112,977,1001]" degrees="6" direction="south" minutes="32" orientation="latitude" pageId="72" pageNumber="73" precision="925" value="-6.5333333">6° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B2DFFDBFB8BF1FCFB68FC7F" box="[1125,1246,977,1001]" degrees="145" direction="east" minutes="37" orientation="longitude" pageId="72" pageNumber="73" precision="925" value="145.61667">145° 37' E</geoCoordinate>
,
<date id="FFA710335B2DFFDBFB04F1FCFAE6FC7F" box="[1258,1360,977,1001]" pageId="72" pageNumber="73" value="1965-02-05">
<collectingDate id="EFE3E9DB5B2DFFDBFB04F1FCFAE6FC7F" box="[1258,1360,977,1001]" pageId="72" pageNumber="73" value="1965-02-05">5 ii 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B2DFFDBFAB3F1FFFF40FB9B" pageId="72" pageNumber="73">M. E. Bacchus</collectorName>
(MEB 171c) (14
<collectionCode id="ED08AE365B2DFFDBFE53F1D8FDB5FB9B" box="[445,515,1013,1037]" pageId="72" pageNumber="73">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2DFFDBFF29F634FF64FA9F" pageId="72" pageNumber="73" type="diagnosis">
<paragraph id="8BA636F35B2DFFDBFF29F634FF64FA9F" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">
<emphasis id="B96DEAE15B2DFFDBFF29F634FE65FBA4" bold="true" box="[199,467,1049,1074]" pageId="72" pageNumber="73">Differential Diagnosis.</emphasis>
A moderately sized, darkly colored species with sexually dimorphic legs (SEM
<figureCitation id="13222A765B2DFFDBFA8BF634FF48FBC3" captionStart="FIGURES 322327" captionStartId="358.[151,264,1650,1673]" captionTargetBox="[183,1404,200,1624]" captionTargetId="figure-29@358.[178,1409,193,1629]" captionTargetPageId="358" captionText="FIGURES 322327. Scanning electron micrographs of H. phainops (322323) and H. okapa (324327)." figureDoi="http://doi.org/10.5281/zenodo.5283987" httpUri="https://zenodo.org/record/5283987/files/figure.png" pageId="72" pageNumber="73">Figs. 325327</figureCitation>
). Bearing some superficial similarity in habitus to
<taxonomicName id="4C194D705B2DFFDBFCD4F613FC72FBC3" authorityName="Perkins" authorityYear="2011" box="[826,964,1085,1109]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="fascinata">
<emphasis id="B96DEAE15B2DFFDBFCD4F613FCF8FBC3" box="[826,846,1086,1109]" italics="true" pageId="72" pageNumber="73">H</emphasis>
.
<emphasis id="B96DEAE15B2DFFDBFCB3F610FC72FBC3" box="[861,964,1085,1109]" italics="true" pageId="72" pageNumber="73">fascinata</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B2DFFDBFC3AF610FBF6FBC3" box="[980,1088,1085,1109]" captionStart="FIGURE 119" captionStartId="234.[151,250,1753,1776]" captionTargetBox="[263,1338,212,1724]" captionTargetId="figure-29@234.[241,1346,193,1732]" captionTargetPageId="234" captionText="FIGURE 119. Hydraena okapa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283652" httpUri="https://zenodo.org/record/5283652/files/figure.png" pageId="72" pageNumber="73">Figs. 119</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFBA2F610FBCCFBC3" box="[1100,1146,1085,1109]" captionStart="FIGURE 122" captionStartId="236.[151,250,1753,1776]" captionTargetBox="[203,1388,215,1715]" captionTargetId="figure-29@236.[187,1400,193,1732]" captionTargetPageId="236" captionText="FIGURE 122. Hydraena fascinata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283658" httpUri="https://zenodo.org/record/5283658/files/figure.png" pageId="72" pageNumber="73">122</figureCitation>
); differentiated therefrom by the very different leg structure and very different aedeagi (
<figureCitation id="13222A765B2DFFDBFCABF64CFC19FBEF" box="[837,943,1121,1145]" captionStart="FIGURES 120121" captionStartId="235.[151,264,1596,1619]" captionTargetBox="[244,1345,239,1485]" captionTargetId="figure-29@235.[226,1360,193,1575]" captionTargetPageId="235" captionText="FIGURES 120121. Aedeagi of holotypes. 120. H. idema. 121. H. okapa." figureDoi="http://doi.org/10.5281/zenodo.5283654" httpUri="https://zenodo.org/record/5283654/files/figure.png" pageId="72" pageNumber="73">Figs. 121</figureCitation>
,
<figureCitation id="13222A765B2DFFDBFC55F64CFC5FFBEF" box="[955,1001,1121,1145]" captionStart="FIGURES 124125" captionStartId="238.[151,264,1650,1673]" captionText="FIGURES 124125. Aedeagi of holotypes. 124. H. fascinata. 125. H. aquila." pageId="72" pageNumber="73">124</figureCitation>
). The female tergite X, gonocoxite and spermatheca are very different in the two species (
<figureCitation id="13222A765B2DFFDBFD22F6A8FCC4FB0B" box="[716,882,1157,1181]" captionStart="FIGURES 400403" captionStartId="372.[151,264,1650,1673]" captionTargetBox="[271,1319,208,1611]" captionTargetId="figure-28@372.[235,1351,193,1629]" captionTargetPageId="372" captionText="FIGURES 400403. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284025" httpUri="https://zenodo.org/record/5284025/files/figure.png" pageId="72" pageNumber="73">Figs. 401, 402</figureCitation>
). The plaques are unusual in that the medial margins are closer to one another in the middle than at either end. The aedeagus (
<figureCitation id="13222A765B2DFFDBFC19F684FBE0FB57" box="[1015,1110,1193,1217]" captionStart="FIGURES 120121" captionStartId="235.[151,264,1596,1619]" captionTargetBox="[244,1345,239,1485]" captionTargetId="figure-29@235.[226,1360,193,1575]" captionTargetPageId="235" captionText="FIGURES 120121. Aedeagi of holotypes. 120. H. idema. 121. H. okapa." figureDoi="http://doi.org/10.5281/zenodo.5283654" httpUri="https://zenodo.org/record/5283654/files/figure.png" pageId="72" pageNumber="73">Fig. 121</figureCitation>
) has unusual parameres. The metaventrite, abdominal apex, and very complicated chaetotaxy of the tibiae are illustrated with SEMs (
<figureCitation id="13222A765B2DFFDBFACAF6E0FF73FA9F" captionStart="FIGURES 322327" captionStartId="358.[151,264,1650,1673]" captionTargetBox="[183,1404,200,1624]" captionTargetId="figure-29@358.[178,1409,193,1629]" captionTargetPageId="358" captionText="FIGURES 322327. Scanning electron micrographs of H. phainops (322323) and H. okapa (324327)." figureDoi="http://doi.org/10.5281/zenodo.5283987" httpUri="https://zenodo.org/record/5283987/files/figure.png" pageId="72" pageNumber="73">Figs. 324 327</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2DFFDAFF29F738FC0CFEA9" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="description">
<paragraph id="8BA636F35B2DFFDBFF29F738FAC5FAC7" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">
<emphasis id="B96DEAE15B2DFFDBFF29F738FEEEFAB8" bold="true" box="[199,344,1301,1326]" inLineHeading="true" pageId="72" pageNumber="73" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B2DFFDBFE42F738FDB8FABB" box="[428,526,1301,1325]" pageId="72" pageNumber="73" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.75/0.72; head 0.25/0.40; pronotum 0.42/0.50, PA 0.43, PB 0.42; elytra 1.08/0.72. Dorsum piceous; legs and maxillary palpi dark brown.
</paragraph>
<paragraph id="8BA636F35B2DFFDBFF29F773FD29F993" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">Frons moderately fine, moderately densely punctate, punctures ca. 1xef; interstices weakly shining, effacedly microreticulate, 14xpd, punctures sparser on disc. Clypeus microreticulate laterally, finely, sparsely punctate medially. Mentum very sparsely very finely punctulate, strongly shining, each puncture with moderately long recumbent seta; postmentum medially effacedly micropunctulate, otherwise finely sparsely punctate, shining. Genae raised, shining, without posterior ridge.</paragraph>
<paragraph id="8BA636F35B2DFFDBFF29F43FFEDDF903" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">Pronotum weakly cordiform, median 2/3 of anterior margin straight or very weakly emarginate; punctures on disc ca. 1xpd largest punctures of frons, interstices effacedly microreticulate, dull, 12xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow; PF2 moderately shallow, very shallowly confluent; PF3 deep; PF4 shallow.</paragraph>
<paragraph id="8BA636F35B2DFFDBFF29F48CFE78F8DF" blockId="72.[151,1437,797,2009]" pageId="72" pageNumber="73">Elytra with summit of posterior declivity slightly past midlength; lateral explanate moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, effacedly microreticulate, dull or very weakly shining, on disc ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B2DFFDAFF29F578FC2AFE8A" blockId="72.[151,1437,797,2009]" lastBlockId="73.[151,1436,151,500]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/4/11/3. P1 laminate; median carina sinuate in profile. P2 raised, l/w ca. 8/3, sides slightly converging toward apex. Plaques wide, flat, closely spaced, medial margins converging slightly toward one another at midlength, located at sides of median depression. Metaventrite without midlongitudinal ridge. AIS width at straight posterior margin ca. 2x P2. Middle and hind legs long and slender, front legs more robust. Profemur (male) with tubercle near trochanter; protibia very slightly arcuate, distal 1/3 widened on medial margin and bearing row of very short spines. Mesotibia very slightly arcuate, medial surface subapically notched. Metatibia straight, medial margin very slightly widened at ca. distal 1/3. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B2CFFDAFCC0F329FC39FE8A" box="[814,911,260,284]" captionStart="FIGURE 119" captionStartId="234.[151,250,1753,1776]" captionTargetBox="[263,1338,212,1724]" captionTargetId="figure-29@234.[241,1346,193,1732]" captionTargetPageId="234" captionText="FIGURE 119. Hydraena okapa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283652" httpUri="https://zenodo.org/record/5283652/files/figure.png" pageId="73" pageNumber="74">Fig. 119</figureCitation>
).
</paragraph>
<paragraph id="8BA636F35B2CFFDAFF79F30AFC0CFEA9" blockId="73.[151,1436,151,500]" box="[151,954,295,319]" pageId="73" pageNumber="74">
Female tergite X, gonocoxite, and spermatheca as illustrated (
<figureCitation id="13222A765B2CFFDAFCA5F30AFC1BFEA9" box="[843,941,295,319]" captionStart="FIGURES 400403" captionStartId="372.[151,264,1650,1673]" captionTargetBox="[271,1319,208,1611]" captionTargetId="figure-28@372.[235,1351,193,1629]" captionTargetPageId="372" captionText="FIGURES 400403. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284025" httpUri="https://zenodo.org/record/5284025/files/figure.png" pageId="73" pageNumber="74">Fig. 401</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2CFFDAFF29F361FCD9FEF2" box="[199,879,332,356]" pageId="73" pageNumber="74" type="etymology">
<paragraph id="8BA636F35B2CFFDAFF29F361FCD9FEF2" blockId="73.[151,1436,151,500]" box="[199,879,332,356]" pageId="73" pageNumber="74">
<emphasis id="B96DEAE15B2CFFDAFF29F361FEFBFEF2" bold="true" box="[199,333,332,356]" inLineHeading="true" pageId="73" pageNumber="74" reason="1">Etymology.</emphasis>
Named in reference to the geographical locality.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B2CFFDAFF29F342FEF2FE62" pageId="73" pageNumber="74" type="distribution">
<paragraph id="8BA636F35B2CFFDAFF29F342FEB8FE3A" blockId="73.[151,1436,151,500]" pageId="73" pageNumber="74">
<emphasis id="B96DEAE15B2CFFDAFF29F342FED6FE1E" bold="true" box="[199,352,367,392]" inLineHeading="true" pageId="73" pageNumber="74" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B2CFFDAFD36F35DFCBEFE1E" box="[728,776,368,392]" pageId="73" pageNumber="74">type</typeStatus>
locality near Okapa, in eastern Area 1; elevation
<quantity id="4CE19B165B2CFFDAFAABF342FA2AFE11" box="[1349,1436,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="73" pageNumber="74" unit="m" value="1372.0">1372 m</quantity>
(
<figureCitation id="13222A765B2CFFDAFF71F3B9FEB4FE3A" box="[159,258,404,428]" captionStart="FIGURES 483486" captionStartId="396.[151,264,1650,1673]" captionTargetBox="[165,1421,252,1595]" captionTargetId="figure-23@396.[165,1421,193,1629]" captionTargetPageId="396" captionText="FIGURES 483486. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284099" httpUri="https://zenodo.org/record/5284099/files/figure.png" pageId="73" pageNumber="74">Fig. 483</figureCitation>
).
</paragraph>
<paragraph id="8BA636F35B2CFFDAFF29F395FEF2FE62" blockId="73.[151,1436,151,500]" pageId="73" pageNumber="74">
<emphasis id="B96DEAE15B2CFFDAFF29F395FEB0FE46" bold="true" box="[199,262,440,464]" pageId="73" pageNumber="74">Note.</emphasis>
In some specimens the medial margins of the plaques are more nearly parallel than they are in the
<typeStatus id="54A288515B2CFFDAFA8FF39AFF71FE62" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
(
<figureCitation id="13222A765B2CFFDAFF38F3F1FE8EFE62" box="[214,312,476,500]" captionStart="FIGURE 119" captionStartId="234.[151,250,1753,1776]" captionTargetBox="[263,1338,212,1724]" captionTargetId="figure-29@234.[241,1346,193,1732]" captionTargetPageId="234" captionText="FIGURE 119. Hydraena okapa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283652" httpUri="https://zenodo.org/record/5283652/files/figure.png" pageId="73" pageNumber="74">Fig. 119</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>