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treatments-xml/data/03/B0/87/03B087E55B14FFE1FF79F6D9FEEAFC82.xml
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<document id="E7DEACECF0768BCE9B5899DBE9E0D694" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B14FFE1FF79F6D9FEEAFC82" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena maculopala Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="115" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="114" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="22D246F4C9772C52A790BC7C8A5F1C11" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5062B95C9F8DD99BBE43410F4E283A77">
<mods:title id="F5F6C4177381A34199804E14A3E4B0ED">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="1E7DE1B6A8572FAF9CDC09CB5C2D2007" type="personal">
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<mods:namePart id="6F0D9BFBB17B2BCA4184F869209D89D4">Perkins, Philip D.</mods:namePart>
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<mods:title id="C10472D88C18C301C9F52E707CF270CB">Zootaxa</mods:title>
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<mods:part id="8659D4009F93776C7893CD0C2E1F97E8">
<mods:date id="82CA3721AF902AE45CF9F9406C27CDBE">2011</mods:date>
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<mods:number id="CBF8C7A5DDE41D26B505238FDBDC7F27">2011-07-08</mods:number>
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<mods:number id="3378533B8E5EB607FD68C094970248E5">2944</mods:number>
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<mods:number id="55E6456B2E3A251E210D3D596F39DF67">1</mods:number>
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<mods:url id="CCE82B54CF44F5057B9186512881496F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
</mods:location>
<mods:classification id="D3D89E61C66D06860292189FCD725FBF">journal article</mods:classification>
<mods:identifier id="6560DF7AE2DDEC70BE37676811C9A711" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="49DBC55F023561266F9F7615052DEE21" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="C6B262A793E94C4D537BCB5837FCAC81" type="Zenodo-Dep">5283420</mods:identifier>
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<treatment id="03B087E55B14FFE1FF79F6D9FEEAFC82" ID-DOI="http://doi.org/10.5281/zenodo.5291741" ID-Zenodo-Dep="5291741" LSID="urn:lsid:plazi:treatment:03B087E55B14FFE1FF79F6D9FEEAFC82" httpUri="http://treatment.plazi.org/id/03B087E55B14FFE1FF79F6D9FEEAFC82" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114">
<subSubSection id="C30365785B14FFE2FF79F6D9FDF5FA9B" box="[151,579,1267,1294]" pageId="113" pageNumber="114" type="nomenclature">
<paragraph id="8BA636F35B14FFE2FF79F6D9FDF5FA9B" blockId="113.[151,579,1267,1326]" box="[151,579,1267,1294]" pageId="113" pageNumber="114">
<heading id="D0EE819F5B14FFE2FF79F6D9FDF5FA9B" bold="true" box="[151,579,1267,1294]" fontSize="11" level="1" pageId="113" pageNumber="114" reason="1">
<emphasis id="B96DEAE15B14FFE2FF79F6D9FDF5FA9B" bold="true" box="[151,579,1267,1294]" pageId="113" pageNumber="114">
<taxonomicName id="4C194D705B14FFE2FF79F6D9FE12FA98" ID-CoL="3N3D4" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,420,1268,1294]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="maculopala" status="sp. nov.">
<emphasis id="B96DEAE15B14FFE2FF79F6D9FE12FA98" bold="true" box="[151,420,1268,1294]" italics="true" pageId="113" pageNumber="114">Hydraena maculopala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B14FFE2FE5DF6DEFDF5FA9B" box="[435,579,1267,1293]" pageId="113" pageNumber="114" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF79F73BFE34FAB8" box="[151,386,1302,1326]" pageId="113" pageNumber="114" type="description">
<paragraph id="8BA636F35B14FFE2FF79F73BFE34FAB8" blockId="113.[151,579,1267,1326]" box="[151,386,1302,1326]" pageId="113" pageNumber="114">
(
<figureCitation id="13222A765B14FFE2FF71F73BFEBDFAB8" box="[159,267,1302,1326]" captionStart="FIGURE 203" captionStartId="297.[151,250,1753,1776]" captionTargetBox="[221,1384,215,1718]" captionTargetId="figure-26@297.[188,1400,193,1732]" captionTargetPageId="297" captionText="FIGURE 203. Hydraena maculopala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283810" httpUri="https://zenodo.org/record/5283810/files/figure.png" pageId="113" pageNumber="114">Figs. 203</figureCitation>
,
<figureCitation id="13222A765B14FFE2FEF8F73BFEF4FAB8" box="[278,322,1302,1326]" captionStart="FIGURES 204–205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204–205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="113" pageNumber="114">205</figureCitation>
,
<figureCitation id="13222A765B14FFE2FEA0F73BFECFFAB8" box="[334,377,1302,1326]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="113" pageNumber="114">531</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF79F772FA99F990" pageId="113" pageNumber="114" type="materials_examined">
<paragraph id="8BA636F35B14FFE2FF79F772FA99F990" blockId="113.[151,1436,1374,2010]" pageId="113" pageNumber="114">
Type Material.
<materialsCitation id="3B713CAE5B14FFE2FEA0F772FD2BFA0C" collectingDate="2006-04-30" collectionCode="ZSM" collectorName="Balke &amp; Sagata" country="Papua New Guinea" elevation="160" latitude="-5.23205" location="Ohu Village" longLatPrecision="1" longitude="145.67938" pageId="113" pageNumber="114" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B14FFE2FEA0F772FE0DFAE1" box="[334,443,1375,1399]" pageId="113" pageNumber="114" type="holotype">
<emphasis id="B96DEAE15B14FFE2FEA0F772FE0DFAE1" bold="true" box="[334,443,1375,1399]" pageId="113" pageNumber="114">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B14FFE2FDF1F772FD4DFAE1" bold="true" box="[543,763,1374,1399]" pageId="113" pageNumber="114">
<collectingRegion id="49DDF8115B14FFE2FDF1F772FD41FAE1" box="[543,759,1374,1399]" country="Papua New Guinea" name="Madang" pageId="113" pageNumber="114">Madang Province</collectingRegion>
:
</emphasis>
<collectingRegion id="49DDF8115B14FFE2FCE6F773FCD1FAE0" box="[776,871,1374,1398]" country="Papua New Guinea" name="Madang" pageId="113" pageNumber="114">Madang</collectingRegion>
,
<location id="8EC660285B14FFE2FC9CF773FC4DFAE0" LSID="urn:lsid:plazi:treatment:03B087E55B14FFE1FF79F6D9FEEAFC82:8EC660285B14FFE2FC9CF773FC4DFAE0" box="[882,1019,1374,1398]" country="Papua New Guinea" latitude="-5.23205" longLatPrecision="1" longitude="145.67938" name="Ohu Village" pageId="113" pageNumber="114" stateProvince="Madang">Ohu Village</location>
,
<quantity id="4CE19B165B14FFE2FBE8F773FBF9FAE0" box="[1030,1103,1374,1398]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="113" pageNumber="114" unit="m" value="160.0">
<elevation id="0034D1C05B14FFE2FBE8F773FBF9FAE0" box="[1030,1103,1374,1398]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="113" pageNumber="114" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B14FFE2FBB4F773FB52FAE0" box="[1114,1252,1374,1398]" degrees="5" direction="south" minutes="13.923" orientation="latitude" pageId="113" pageNumber="114" precision="1" value="-5.23205">5° 13.923' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B14FFE2FB1EF773FA21FAE0" box="[1264,1431,1374,1398]" degrees="145" direction="east" minutes="40.763" orientation="longitude" pageId="113" pageNumber="114" precision="1" value="145.67938">145° 40.763' E</geoCoordinate>
,
<date id="FFA710335B14FFE2FF79F7AFFEA5FA0C" box="[151,275,1410,1434]" pageId="113" pageNumber="114" value="2006-04-30">
<collectingDate id="EFE3E9DB5B14FFE2FF79F7AFFEA5FA0C" box="[151,275,1410,1434]" pageId="113" pageNumber="114" value="2006-04-30">30 iv 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B14FFE2FEF1F7AFFED6FA0C" box="[287,352,1410,1434]" pageId="113" pageNumber="114">Balke</collectorName>
&amp;
<collectorName id="26EC53255B14FFE2FE6BF7AFFE79FA0C" box="[389,463,1410,1434]" pageId="113" pageNumber="114">Sagata</collectorName>
(
<collectingCountry id="F30E76635B14FFE2FE31F7AFFDAFFA0C" box="[479,537,1410,1434]" name="Papua New Guinea" pageId="113" pageNumber="114">PNG</collectingCountry>
49) (
<collectionCode id="ED08AE365B14FFE2FDB8F7AFFD23FA0C" box="[598,661,1410,1434]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="113" pageNumber="114" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B14FFE2FD46F7AEFBE2FA28" collectingDate="2006-05-04" collectionCode="ZSM" collectorName="Balke &amp; Manaono" country="Papua New Guinea" elevation="700" latitude="-4.7035832" location="Adelbert Mts." longLatPrecision="1" longitude="145.41924" municipality="Sewan" pageId="113" pageNumber="114" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B14FFE2FD46F7AEFC96FA0D" bold="true" box="[680,800,1411,1435]" pageId="113" pageNumber="114">
<typeStatus id="54A288515B14FFE2FD46F7AEFC96FA0D" box="[680,800,1411,1435]" pageId="113" pageNumber="114" type="paratype">Paratypes</typeStatus>
</emphasis>
(29):
<emphasis id="B96DEAE15B14FFE2FC80F7AEFBFDFA0D" bold="true" box="[878,1099,1410,1435]" pageId="113" pageNumber="114">
<collectingRegion id="49DDF8115B14FFE2FC80F7AEFBF0FA0D" box="[878,1094,1410,1435]" country="Papua New Guinea" name="Madang" pageId="113" pageNumber="114">Madang Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B14FFE2FBB5F7AFFB4CFA0C" LSID="urn:lsid:plazi:treatment:03B087E55B14FFE1FF79F6D9FEEAFC82:8EC660285B14FFE2FBB5F7AFFB4CFA0C" box="[1115,1274,1410,1434]" country="Papua New Guinea" latitude="-4.7035832" longLatPrecision="1" longitude="145.41924" municipality="Sewan" name="Adelbert Mts." pageId="113" pageNumber="114" stateProvince="Madang">Adelbert Mts.</location>
,
<collectingMunicipality id="6BC2AC895B14FFE2FAE8F7AFFAE7FA0C" box="[1286,1361,1410,1434]" pageId="113" pageNumber="114">Sewan</collectingMunicipality>
–Keki,
<quantity id="4CE19B165B14FFE2FF79F78BFF56FA28" box="[151,224,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="113" pageNumber="114" unit="m" value="700.0">
<elevation id="0034D1C05B14FFE2FF79F78BFF56FA28" box="[151,224,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="113" pageNumber="114" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B14FFE2FF02F78BFEC3FA28" box="[236,373,1446,1470]" degrees="4" direction="south" minutes="42.215" orientation="latitude" pageId="113" pageNumber="114" precision="1" value="-4.7035832">4° 42.215' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B14FFE2FE6CF78BFD9CFA28" box="[386,554,1446,1470]" degrees="145" direction="east" minutes="25.154" orientation="longitude" pageId="113" pageNumber="114" precision="1" value="145.41924">145° 25.154' E</geoCoordinate>
,
<date id="FFA710335B14FFE2FDD8F78BFD2DFA28" box="[566,667,1446,1470]" pageId="113" pageNumber="114" value="2006-05-04">
<collectingDate id="EFE3E9DB5B14FFE2FDD8F78BFD2DFA28" box="[566,667,1446,1470]" pageId="113" pageNumber="114" value="2006-05-04">4 v 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B14FFE2FD48F78BFD51FA28" box="[678,743,1446,1470]" pageId="113" pageNumber="114">Balke</collectorName>
&amp;
<collectorName id="26EC53255B14FFE2FCE5F78AFCC2FA28" box="[779,884,1447,1470]" pageId="113" pageNumber="114">Manaono</collectorName>
(
<collectingCountry id="F30E76635B14FFE2FC6DF78BFC0BFA28" box="[899,957,1446,1470]" name="Papua New Guinea" pageId="113" pageNumber="114">PNG</collectingCountry>
51) (3
<collectionCode id="ED08AE365B14FFE2FBE0F78BFBFAFA28" box="[1038,1100,1446,1470]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="113" pageNumber="114" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B14FFE2FB8EF78BFC4DFA74" collectingDate="2006-11-29" collectionCode="ZSM" collectorName="Binatang Boys" country="Papua New Guinea" elevation="400" latitude="-4.7176332" location="Keki" longLatPrecision="1" longitude="145.40729" municipality="Adelbert Mts." pageId="113" pageNumber="114" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
<location id="8EC660285B14FFE2FB8EF78BFB2FFA28" LSID="urn:lsid:plazi:treatment:03B087E55B14FFE1FF79F6D9FEEAFC82:8EC660285B14FFE2FB8EF78BFB2FFA28" box="[1120,1177,1446,1470]" country="Papua New Guinea" latitude="-4.7176332" longLatPrecision="1" longitude="145.40729" municipality="Adelbert Mts." name="Keki" pageId="113" pageNumber="114" stateProvince="Madang Province">Keki</location>
,
<collectingMunicipality id="6BC2AC895B14FFE2FB4AF78BFAF4FA28" box="[1188,1346,1446,1470]" pageId="113" pageNumber="114">Adelbert Mts.</collectingMunicipality>
,
<quantity id="4CE19B165B14FFE2FAA0F78BFA21FA28" box="[1358,1431,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="113" pageNumber="114" unit="m" value="400.0">
<elevation id="0034D1C05B14FFE2FAA0F78BFA21FA28" box="[1358,1431,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="113" pageNumber="114" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B14FFE2FF79F7E7FE97FA74" box="[151,289,1482,1506]" degrees="4" direction="south" minutes="43.058" orientation="latitude" pageId="113" pageNumber="114" precision="1" value="-4.7176332">4° 43.058' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B14FFE2FEC0F7E7FE60FA74" box="[302,470,1482,1506]" degrees="145" direction="east" minutes="24.437" orientation="longitude" pageId="113" pageNumber="114" precision="1" value="145.40729">145° 24.437' E</geoCoordinate>
,
<date id="FFA710335B14FFE2FE0CF7E7FDEBFA74" box="[482,605,1482,1506]" pageId="113" pageNumber="114" value="2006-11-29">
<collectingDate id="EFE3E9DB5B14FFE2FE0CF7E7FDEBFA74" box="[482,605,1482,1506]" pageId="113" pageNumber="114" value="2006-11-29">29 xi 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B14FFE2FD86F7E7FCBBFA75" box="[616,781,1482,1507]" pageId="113" pageNumber="114">Binatang Boys</collectorName>
(
<collectingCountry id="F30E76635B14FFE2FCF2F7E7FCE0FA74" box="[796,854,1482,1506]" name="Papua New Guinea" pageId="113" pageNumber="114">PNG</collectingCountry>
119) (2
<collectionCode id="ED08AE365B14FFE2FC5BF7E7FC45FA74" box="[949,1011,1482,1506]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="113" pageNumber="114" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B14FFE2FBE9F7E7FA9DF990" collectingDate="2006-11-30" collectionCode="MCZ, NHM, NMW, ZSM" collectorName="Binatang Boys" country="Papua New Guinea" elevation="700" latitude="-4.6967" location="Sewan" longLatPrecision="9" longitude="145.42433" municipality="Adelbert Mts." pageId="113" pageNumber="114" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
Keki–
<location id="8EC660285B14FFE2FBA0F7E7FB2FFA74" LSID="urn:lsid:plazi:treatment:03B087E55B14FFE1FF79F6D9FEEAFC82:8EC660285B14FFE2FBA0F7E7FB2FFA74" box="[1102,1177,1482,1506]" country="Papua New Guinea" latitude="-4.6967" longLatPrecision="9" longitude="145.42433" municipality="Adelbert Mts." name="Sewan" pageId="113" pageNumber="114" stateProvince="Madang Province">Sewan</location>
,
<collectingMunicipality id="6BC2AC895B14FFE2FB4AF7E7FAF4FA74" box="[1188,1346,1482,1506]" pageId="113" pageNumber="114">Adelbert Mts.</collectingMunicipality>
,
<quantity id="4CE19B165B14FFE2FAA0F7E7FA21FA74" box="[1358,1431,1482,1506]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="113" pageNumber="114" unit="m" value="700.0">
<elevation id="0034D1C05B14FFE2FAA0F7E7FA21FA74" box="[1358,1431,1482,1506]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="113" pageNumber="114" unit="m" value="700.0">700 m</elevation>
</quantity>
,
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,
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,
<date id="FFA710335B14FFE2FE3AF7C3FDF9F990" box="[468,591,1518,1542]" pageId="113" pageNumber="114" value="2006-11-30">
<collectingDate id="EFE3E9DB5B14FFE2FE3AF7C3FDF9F990" box="[468,591,1518,1542]" pageId="113" pageNumber="114" value="2006-11-30">30 xi 2006</collectingDate>
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(
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120) (24
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<collectionCode id="ED08AE365B14FFE2FBA1F7C2FB21F990" box="[1103,1175,1519,1542]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="113" pageNumber="114">NMW</collectionCode>
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<collectingCountry id="F30E76635B14FFE2FB4CF7C3FB6CF990" box="[1186,1242,1518,1542]" name="Papua New Guinea" pageId="113" pageNumber="114">PNG</collectingCountry>
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<collectionCode id="ED08AE365B14FFE2FB0BF7C3FA95F990" box="[1253,1315,1518,1542]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="113" pageNumber="114" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF29F43FFA8BF948" pageId="113" pageNumber="114" type="diagnosis">
<paragraph id="8BA636F35B14FFE2FF29F43FFA8BF948" blockId="113.[151,1436,1374,2010]" pageId="113" pageNumber="114">
<emphasis id="B96DEAE15B14FFE2FF29F43FFE65F9BD" bold="true" box="[199,467,1554,1579]" pageId="113" pageNumber="114">Differential Diagnosis.</emphasis>
Similar in general dorsal coloration and pronotal shape and sculpture to
<taxonomicName id="4C194D705B14FFE2FAE1F43EFA2DF9BD" authorityName="Perkins" authorityYear="2011" box="[1295,1435,1555,1579]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nanopala">
<emphasis id="B96DEAE15B14FFE2FAE1F43EFA95F9BC" box="[1295,1315,1555,1578]" italics="true" pageId="113" pageNumber="114">H</emphasis>
.
<emphasis id="B96DEAE15B14FFE2FADCF43EFA2DF9BD" box="[1330,1435,1555,1579]" italics="true" pageId="113" pageNumber="114">nanopala</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B14FFE2FF71F41BFEBDF9D8" box="[159,267,1590,1614]" captionStart="FIGURE 203" captionStartId="297.[151,250,1753,1776]" captionTargetBox="[221,1384,215,1718]" captionTargetId="figure-26@297.[188,1400,193,1732]" captionTargetPageId="297" captionText="FIGURE 203. Hydraena maculopala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283810" httpUri="https://zenodo.org/record/5283810/files/figure.png" pageId="113" pageNumber="114">Figs. 203</figureCitation>
,
<figureCitation id="13222A765B14FFE2FEF9F41BFEF3F9D8" box="[279,325,1590,1614]" captionStart="FIGURE 206" captionStartId="299.[151,250,1753,1776]" captionTargetBox="[218,1381,220,1724]" captionTargetId="figure-25@299.[191,1395,193,1732]" captionTargetPageId="299" captionText="FIGURE 206. Hydraena nanopala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283816" httpUri="https://zenodo.org/record/5283816/files/figure.png" pageId="113" pageNumber="114">206</figureCitation>
); differing therefrom in larger size (ca. 1.32 vs.
<quantity id="4CE19B165B14FFE2FCB0F41BFC70F9D8" box="[862,966,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="113" pageNumber="114" unit="mm" value="1.23">1.23 mm</quantity>
), the presence of an ill-defined macula on the pronotum, the proportionally longer elytra (PW/EL ca. 0.58 vs. 0.63, and the slightly more widely separated plaques (ratios ca. 4/1/5/5 vs. 4/1/5/4). Among
<taxonomicName id="4C194D705B14FFE2FD48F453FD6DF900" box="[678,731,1662,1686]" class="Magnoliopsida" family="Apocynaceae" genus="Pala" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="113" pageNumber="114" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEAE15B14FFE2FD48F453FD6DF900" box="[678,731,1662,1686]" italics="true" pageId="113" pageNumber="114">Pala</emphasis>
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group members, the mesoventral intercoxal process (P2) is the narrowest in these two species. They also show a similarity in basic aedeagal plan, while differing distinctively in details of the distal piece (
<figureCitation id="13222A765B14FFE2FE52F4EBFD9EF948" box="[444,552,1734,1758]" captionStart="FIGURES 204–205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204–205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="113" pageNumber="114">Figs. 205</figureCitation>
,
<figureCitation id="13222A765B14FFE2FDDAF4EBFDD4F948" box="[564,610,1734,1758]" captionStart="FIGURES 208–209" captionStartId="301.[151,264,1650,1673]" captionText="FIGURES 208–209. Aedeagi of holotypes. 208. H. nanopala. 209. H. palamita." pageId="113" pageNumber="114">208</figureCitation>
); reliable determinations will require examination of the aedeagi.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE1FF29F4C7FC97FD31" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114" type="description">
<paragraph id="8BA636F35B14FFE2FF29F4C7FD5EF8F8" blockId="113.[151,1436,1374,2010]" pageId="113" pageNumber="114">
<emphasis id="B96DEAE15B14FFE2FF29F4C7FEEEF895" bold="true" box="[199,344,1770,1795]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B14FFE2FE42F4C7FDB8F894" box="[428,526,1770,1794]" pageId="113" pageNumber="114" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.32/0.58; head 0.23/0.32; pronotum 0.32/0.45, PA 0.35, PB 0.35; elytra 0.78/0.58. Dorsum with frons dark brown to piceous; clypeus brown; labrum testaceous; pronotum testaceous around light brown, ill-defined macula; elytra brown; legs light brown to testaceous; maxillary palpi testaceous, tip not darker.
</paragraph>
<paragraph id="8BA636F35B14FFE1FF29F556FED7FF61" blockId="113.[151,1436,1374,2010]" lastBlockId="114.[151,1436,151,788]" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114">Frons punctures ca. 1–2xef, slightly larger near eyes than medially; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–3xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow, obsolete; PF2 moderately deep; PF3 deep; PF4 very shallow.</paragraph>
<paragraph id="8BA636F35B17FFE1FF29F329FF47FE3A" blockId="114.[151,1436,151,788]" pageId="114" pageNumber="115">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B17FFE1FF29F39AFC97FD31" blockId="114.[151,1436,151,788]" pageId="114" pageNumber="115">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 4/1/5/5. P1 ca. 1/2 P2; median carina straight in profile; postcoxal process medially concave. P2 transversely concave, l/w ca. 5/4, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. 1.5x P2. All legs moderately long and slender. Profemur (male) with very small tubercle next to trochanter; protibia slightly arcuate, gradually increasing in width from base to apex. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B17FFE1FD5CF0A2FCA2FD31" box="[690,788,655,679]" captionStart="FIGURES 204–205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204–205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="114" pageNumber="115">Fig. 205</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B17FFE1FF29F099FB2EFD5A" box="[199,1176,692,716]" pageId="114" pageNumber="115" type="etymology">
<paragraph id="8BA636F35B17FFE1FF29F099FB2EFD5A" blockId="114.[151,1436,151,788]" box="[199,1176,692,716]" pageId="114" pageNumber="115">
<emphasis id="B96DEAE15B17FFE1FF29F099FEFBFD5A" bold="true" box="[199,333,692,716]" inLineHeading="true" pageId="114" pageNumber="115" reason="1">Etymology.</emphasis>
This species is the only member of the
<taxonomicName id="4C194D705B17FFE1FCE5F099FCF6FD5A" box="[779,832,692,716]" class="Magnoliopsida" family="Apocynaceae" genus="Pala" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="114" pageNumber="115" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEAE15B17FFE1FCE5F099FCF6FD5A" box="[779,832,692,716]" italics="true" pageId="114" pageNumber="115">Pala</emphasis>
</taxonomicName>
group with a pronotal macula.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B17FFE1FF29F0FAFEEAFC82" pageId="114" pageNumber="115" type="distribution">
<paragraph id="8BA636F35B17FFE1FF29F0FAFEEAFC82" blockId="114.[151,1436,151,788]" pageId="114" pageNumber="115">
<emphasis id="B96DEAE15B17FFE1FF29F0FAFED6FD66" bold="true" box="[199,352,727,752]" inLineHeading="true" pageId="114" pageNumber="115" reason="1">Distribution.</emphasis>
Currently known from three localities in the northwestern part of Area 5; elevation range
<quantity id="4CE19B165B17FFE1FA8DF0FAFF68FC85" metricMagnitude="2" metricUnit="m" metricValue="4.3" metricValueMax="7.0" metricValueMin="1.6" pageId="114" pageNumber="115" unit="m" value="430.0" valueMax="700.0" valueMin="160.0">160– 700 m</quantity>
(
<figureCitation id="13222A765B17FFE1FF03F0D1FEF9FC82" box="[237,335,764,788]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="114" pageNumber="115">Fig. 531</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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