160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="475D3608C366E8E1C0EF2DA847DAF23C" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B10FFE5FF79F1DAFDF4FED6" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena torosopala Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="119" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="118" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="501ED45A0D157896769A06DA0BB6CAA9">
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<mods:title id="BB057C88FA8A41DB0238396A0F8FA66F">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="0CD6A9618F0197BC08E760A04555ACEC">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="F0CC546C96BE5983357F0694545A18BE">text</mods:typeOfResource>
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<mods:title id="20D655997E34B7FEE3C7F76FC49C7F69">Zootaxa</mods:title>
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<mods:part id="1B84EE3B3F48039100FD96FF3EFA2C57">
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<mods:date id="B0C0AC7F1837DC7DA12A0E81E7336DE1">2011</mods:date>
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<mods:number id="13AC1E1798850AEDEF329107775AA7AE">2011-07-08</mods:number>
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<mods:number id="C674AD00DB63AA18FEAECFBA7C2E18DA">2944</mods:number>
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<mods:url id="3CE412723C0B4686CC3F7158C1515EAA">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="1151AB312B10BA2B26C44A9DBC86E61E">journal article</mods:classification>
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<mods:identifier id="BF100B199065A744C51C8A168EA4A2FD" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<treatment id="03B087E55B10FFE5FF79F1DAFDF4FED6" ID-DOI="http://doi.org/10.5281/zenodo.5291755" ID-Zenodo-Dep="5291755" LSID="urn:lsid:plazi:treatment:03B087E55B10FFE5FF79F1DAFDF4FED6" httpUri="http://treatment.plazi.org/id/03B087E55B10FFE5FF79F1DAFDF4FED6" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118">
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<subSubSection id="C30365785B10FFE6FF79F1DAFD82FB87" box="[151,564,1015,1041]" pageId="117" pageNumber="118" type="nomenclature">
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<paragraph id="8BA636F35B10FFE6FF79F1DAFD82FB87" blockId="117.[151,564,1015,1074]" box="[151,564,1015,1041]" pageId="117" pageNumber="118">
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<heading id="D0EE819F5B10FFE6FF79F1DAFD82FB87" bold="true" box="[151,564,1015,1041]" fontSize="11" level="1" pageId="117" pageNumber="118" reason="1">
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<emphasis id="B96DEAE15B10FFE6FF79F1DAFD82FB87" bold="true" box="[151,564,1015,1041]" pageId="117" pageNumber="118">
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<taxonomicName id="4C194D705B10FFE6FF79F1DAFE23FB87" ID-CoL="3N3JT" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,405,1015,1041]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="torosopala" status="sp. nov.">
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<emphasis id="B96DEAE15B10FFE6FF79F1DAFE23FB87" bold="true" box="[151,405,1015,1041]" italics="true" pageId="117" pageNumber="118">Hydraena torosopala</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B10FFE6FE4AF1DAFD82FB87" box="[420,564,1015,1041]" pageId="117" pageNumber="118" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B10FFE6FF79F637FE34FBA4" box="[151,386,1050,1074]" pageId="117" pageNumber="118" type="description">
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<paragraph id="8BA636F35B10FFE6FF79F637FE34FBA4" blockId="117.[151,564,1015,1074]" box="[151,386,1050,1074]" pageId="117" pageNumber="118">
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(
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<figureCitation id="13222A765B10FFE6FF71F637FEBAFBA4" box="[159,268,1050,1074]" captionStart="FIGURE 214" captionStartId="305.[151,250,1753,1776]" captionTargetBox="[275,1312,198,1724]" captionTargetId="figure-26@305.[259,1328,193,1732]" captionTargetPageId="305" captionText="FIGURE 214. Hydraena torosopala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283834" httpUri="https://zenodo.org/record/5283834/files/figure.png" pageId="117" pageNumber="118">Figs. 214</figureCitation>
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,
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<figureCitation id="13222A765B10FFE6FEF8F637FEF5FBA4" box="[278,323,1050,1074]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="117" pageNumber="118">216</figureCitation>
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,
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<figureCitation id="13222A765B10FFE6FEA0F637FECFFBA4" box="[334,377,1050,1074]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="117" pageNumber="118">532</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B10FFE6FF79F64EFD28FB08" pageId="117" pageNumber="118" type="materials_examined">
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<paragraph id="8BA636F35B10FFE6FF79F64EFD28FB08" blockId="117.[151,1436,1122,2010]" pageId="117" pageNumber="118">
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Type Material.
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<materialsCitation id="3B713CAE5B10FFE6FEA1F64EFD28FB08" collectingDate="2006-11-29" collectionCode="ZSM" country="Papua New Guinea" county="Binatang Boys" elevation="400" latitude="-4.7176332" location="Keki" longLatPrecision="1" longitude="145.40729" municipality="Adelbert Mts." pageId="117" pageNumber="118" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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||
<typeStatus id="54A288515B10FFE6FEA1F64EFE0AFBED" box="[335,444,1123,1147]" pageId="117" pageNumber="118" type="holotype">
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<emphasis id="B96DEAE15B10FFE6FEA1F64EFE0AFBED" bold="true" box="[335,444,1123,1147]" pageId="117" pageNumber="118">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B10FFE6FDCAF64EFCB7FBED" bold="true" box="[548,769,1122,1147]" pageId="117" pageNumber="118">
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<collectingRegion id="49DDF8115B10FFE6FDCAF64EFD4AFBED" box="[548,764,1122,1147]" country="Papua New Guinea" name="Madang" pageId="117" pageNumber="118">Madang Province</collectingRegion>
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:
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</emphasis>
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<location id="8EC660285B10FFE6FCFEF64FFCFFFBEC" LSID="urn:lsid:plazi:treatment:03B087E55B10FFE5FF79F1DAFDF4FED6:8EC660285B10FFE6FCFEF64FFCFFFBEC" box="[784,841,1122,1146]" country="Papua New Guinea" county="Binatang Boys" latitude="-4.7176332" longLatPrecision="1" longitude="145.40729" municipality="Adelbert Mts." name="Keki" pageId="117" pageNumber="118" stateProvince="Madang">Keki</location>
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,
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<collectingMunicipality id="6BC2AC895B10FFE6FCBBF64FFC42FBEC" box="[853,1012,1122,1146]" pageId="117" pageNumber="118">Adelbert Mts.</collectingMunicipality>
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,
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||
<quantity id="4CE19B165B10FFE6FBEEF64FFBFFFBEC" box="[1024,1097,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="117" pageNumber="118" unit="m" value="400.0">
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<elevation id="0034D1C05B10FFE6FBEEF64FFBFFFBEC" box="[1024,1097,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="117" pageNumber="118" unit="m" value="400.0">400 m</elevation>
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</quantity>
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,
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||
<geoCoordinate id="EE2D50345B10FFE6FBB8F64FFB57FBEC" box="[1110,1249,1122,1146]" degrees="4" direction="south" minutes="43.058" orientation="latitude" pageId="117" pageNumber="118" precision="1" value="-4.7176332">4° 43.058' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B10FFE6FB00F64FFA21FBEC" box="[1262,1431,1122,1146]" degrees="145" direction="east" minutes="24.437" orientation="longitude" pageId="117" pageNumber="118" precision="1" value="145.40729">145° 24.437' E</geoCoordinate>
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,
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||
<date id="FFA710335B10FFE6FF79F6ABFEA7FB08" box="[151,273,1158,1182]" pageId="117" pageNumber="118" value="2006-11-29">
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||
<collectingDate id="EFE3E9DB5B10FFE6FF79F6ABFEA7FB08" box="[151,273,1158,1182]" pageId="117" pageNumber="118" value="2006-11-29">29 xi 2006</collectingDate>
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</date>
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||
,
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||
<collectingCounty id="62C74E7F5B10FFE6FEF2F6ABFE77FB09" box="[284,449,1158,1183]" pageId="117" pageNumber="118">Binatang Boys</collectingCounty>
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||
(
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||
<collectingCountry id="F30E76635B10FFE6FE3FF6ABFDBDFB08" box="[465,523,1158,1182]" name="Papua New Guinea" pageId="117" pageNumber="118">PNG</collectingCountry>
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119) (
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<collectionCode id="ED08AE365B10FFE6FDBDF6ABFD24FB08" box="[595,658,1158,1182]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="117" pageNumber="118" type="Museum">ZSM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B10FFE6FF29F687FF64FAB8" pageId="117" pageNumber="118" type="diagnosis">
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<paragraph id="8BA636F35B10FFE6FF29F687FF64FAB8" blockId="117.[151,1436,1122,2010]" pageId="117" pageNumber="118">
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<emphasis id="B96DEAE15B10FFE6FF29F687FE63FB55" bold="true" box="[199,469,1194,1219]" pageId="117" pageNumber="118">Differential Diagnosis.</emphasis>
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Similar to
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<taxonomicName id="4C194D705B10FFE6FDB2F686FD6AFB54" authorityName="Perkins" authorityYear="2011" box="[604,732,1195,1218]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="tritropis">
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<emphasis id="B96DEAE15B10FFE6FDB2F686FDC6FB54" box="[604,624,1195,1218]" italics="true" pageId="117" pageNumber="118">H</emphasis>
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.
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<emphasis id="B96DEAE15B10FFE6FD6FF686FD6AFB54" box="[641,732,1195,1218]" italics="true" pageId="117" pageNumber="118">tritropis</emphasis>
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</taxonomicName>
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in dorsal color and plaque ratios; differing therefrom by the larger size (1.45 vs.
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<quantity id="4CE19B165B10FFE6FE6CF6E3FE5AFB70" box="[386,492,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="117" pageNumber="118" unit="mm" value="1.29">1.29 mm</quantity>
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), the proportionally smaller pronotum (PW/EL ca. 0.62 vs. 0.66), and the concave prosternal intercoxal process (
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<figureCitation id="13222A765B10FFE6FE07F6DFFDE2FA9C" box="[489,596,1266,1290]" captionStart="FIGURE 211" captionStartId="303.[151,250,1753,1776]" captionTargetBox="[261,1342,226,1718]" captionTargetId="figure-26@303.[228,1360,193,1732]" captionTargetPageId="303" captionText="FIGURE 211. Hydraena tritropis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283828" httpUri="https://zenodo.org/record/5283828/files/figure.png" pageId="117" pageNumber="118">Figs. 211</figureCitation>
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,
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<figureCitation id="13222A765B10FFE6FD8EF6DFFD38FA9C" box="[608,654,1266,1290]" captionStart="FIGURE 214" captionStartId="305.[151,250,1753,1776]" captionTargetBox="[275,1312,198,1724]" captionTargetId="figure-26@305.[259,1328,193,1732]" captionTargetPageId="305" captionText="FIGURE 214. Hydraena torosopala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283834" httpUri="https://zenodo.org/record/5283834/files/figure.png" pageId="117" pageNumber="118">214</figureCitation>
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). The male genitalia of the two species distinctively differ (
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<figureCitation id="13222A765B10FFE6FAC5F6DFFA21FA9C" box="[1323,1431,1266,1290]" captionStart="FIGURES 212–213" captionStartId="304.[151,264,1650,1673]" captionTargetBox="[250,1326,204,1607]" captionTargetId="figure-32@304.[226,1360,193,1629]" captionTargetPageId="304" captionText="FIGURES 212–213. Aedeagi of holotypes. 212. H. formosopala. 213. H. tritropis." figureDoi="http://doi.org/10.5281/zenodo.5283830" httpUri="https://zenodo.org/record/5283830/files/figure.png" pageId="117" pageNumber="118">Figs. 213</figureCitation>
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,
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<figureCitation id="13222A765B10FFE6FF79F73BFF70FAB8" box="[151,198,1302,1326]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="117" pageNumber="118">216</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B10FFE5FF29F717FDCFFF42" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118" type="description">
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<paragraph id="8BA636F35B10FFE6FF29F717FC5EFA28" blockId="117.[151,1436,1122,2010]" pageId="117" pageNumber="118">
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<emphasis id="B96DEAE15B10FFE6FF29F717FEEEFAC5" bold="true" box="[199,344,1338,1363]" inLineHeading="true" pageId="117" pageNumber="118" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B10FFE6FE42F717FDB8FAC4" box="[428,526,1338,1362]" pageId="117" pageNumber="118" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.45/0.69; head 0.22/0.36; pronotum 0.36/0.53, PA 0.39, PB 0.43; elytra 0.86/0.69. Dorsum of head with frons dark brown to dark reddish brown, clypeus lighter, brown, labrum testaceous; pronotum light brown to testaceous; elytra dark brown to dark reddish brown; legs and maxillary palpi testaceous, tip of palpus not darker.
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</paragraph>
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<paragraph id="8BA636F35B10FFE6FF29F7E6FBD0F9BC" blockId="117.[151,1436,1122,2010]" pageId="117" pageNumber="118">Frons punctures ca. 1xef, very slightly, if at all, larger and denser near eyes than medially; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge.</paragraph>
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<paragraph id="8BA636F35B10FFE6FF29F41BFB50F8F8" blockId="117.[151,1436,1122,2010]" pageId="117" pageNumber="118">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 7/1/5/5. P1 ca. 4/7 P2; median carina anterior to coxae, straight in profile; intercoxal process wide, medially concave. P2 transversely concave, l/w ca. 5/7, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. equal P2. All legs moderately long and slender. Profemur (male) without tubercle next to trochanter; protibia slightly arcuate, anterior margin slightly widened subapically. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Pronotum very transverse, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–3xpd, punctures very slightly, if at all, larger and denser at anterior and posterior; PF1 absent; PF2 very shallow, obsolete; PF3 shallow; PF4 absent.</paragraph>
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<paragraph id="8BA636F35B10FFE5FF29F557FDCFFF42" blockId="117.[151,1436,1122,2010]" lastBlockId="118.[151,1436,151,320]" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118">
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Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins rather wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another. Aedeagus as illustrated (
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<figureCitation id="13222A765B13FFE5FDE4F291FDDBFF42" box="[522,621,188,212]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="118" pageNumber="119">Fig. 216</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF29F2CDFAC7FF61" box="[199,1393,223,248]" pageId="118" pageNumber="119" type="etymology">
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<paragraph id="8BA636F35B13FFE5FF29F2CDFAC7FF61" blockId="118.[151,1436,151,320]" box="[199,1393,223,248]" pageId="118" pageNumber="119">
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<emphasis id="B96DEAE15B13FFE5FF29F2CDFEFBFF6E" bold="true" box="[199,333,224,248]" inLineHeading="true" pageId="118" pageNumber="119" reason="1">Etymology.</emphasis>
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"Muscular, bulging"; named in reference to the thick cuticle between the pro- and mesocoxae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF29F32EFDF4FED6" pageId="118" pageNumber="119" type="distribution">
|
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<paragraph id="8BA636F35B13FFE5FF29F32EFDF4FED6" blockId="118.[151,1436,151,320]" pageId="118" pageNumber="119">
|
||
<emphasis id="B96DEAE15B13FFE5FF29F32EFED6FE8A" bold="true" box="[199,352,259,284]" inLineHeading="true" pageId="118" pageNumber="119" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B13FFE5FD24F329FD4CFE8A" box="[714,762,260,284]" pageId="118" pageNumber="119">type</typeStatus>
|
||
locality, Keki, in the Adelbert Mts., in the northwestern part of Area 5; elevation
|
||
<quantity id="4CE19B165B13FFE5FE93F30AFE72FEA9" box="[381,452,295,319]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="118" pageNumber="119" unit="m" value="400.0">400 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B13FFE5FE3DF30AFD84FEA9" box="[467,562,295,319]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="118" pageNumber="119">Fig. 532</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |