treatments-xml/data/03/B0/87/03B087DCFFE0FFF1FF52F53B7C58FF09.xml

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<mods:title id="A78AFBC5A2E03EB97185EB6AC9286659">Bidessus anatolicus adiyaman ssp. n. from Adıyaman province, southern Turkey (Coleoptera, Dytiscidae, Bidessini)</mods:title>
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<mods:name id="402280508957153C22C5AAD46B7B5250" type="personal">
<mods:role id="14C98EC01A99BB913EC7E30012B3AB3A">
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<mods:namePart id="4DA0CFC77FB34149D6D56E6C0A8A56B7">Aykut, Medeni</mods:namePart>
<mods:nameIdentifier id="B7EBA0FF45504EAC261A3F8BA566A7BE" type="ORCID">0000-0001-6824-1394</mods:nameIdentifier>
<mods:affiliation id="46D56873FE6AFE6E1CE5AC7491621E29">Dicle University, Ziya Gökalp Education Faculty, Department of Mathematics and Science, TR- 21280 Diyarbakır, Turkey. medeniaykut @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6824 - 1394</mods:affiliation>
<mods:nameIdentifier id="23B627C87FC6F5E37D724A2EFB191EBD" type="email">medeniaykut@hotmail.com</mods:nameIdentifier>
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<mods:name id="12E9622FC11EAF26C751B70AFC79436A" type="personal">
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<mods:namePart id="A2C3D02D783AC57285E048FD439F1F6C">Taşar, Gani Erhan</mods:namePart>
<mods:affiliation id="2FCD8DB66C27949FB0E5A52116C8E6F8">Adıyaman University, Kahta Vocational High School, TR- 02400 Adıyaman, Turkey.</mods:affiliation>
</mods:name>
<mods:name id="C82B4BD2BE6A31A164858650BD446E2A" type="personal">
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<mods:namePart id="39290C10EB4482DFCB630D2761975A5F">Fery, Hans</mods:namePart>
<mods:affiliation id="FA09C0A98E639B997EB338E2A791AC33">Räuschstrasse 73, D- 13509 Berlin, Germany.</mods:affiliation>
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<mods:title id="D7607310AEEA15B3FEDEC4DA8BDED49F">Zootaxa</mods:title>
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<mods:part id="76AD13ACE0571F38C6A23EE5865616C7">
<mods:date id="E0613F531F6D9E45E145EF1F967BDC4B">2021</mods:date>
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<mods:number id="B7E5C1F6000F9643075432BAA3F254FF">2021-09-02</mods:number>
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<mods:number id="1B22C1BF3FB8588D01C3D278CA73DE61">5027</mods:number>
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<mods:identifier id="C5D71DECE201B178100C7412FE233B56" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B087DCFFE0FFF1FF52F53B7C58FF09" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09" httpUri="http://treatment.plazi.org/id/03B087DCFFE0FFF1FF52F53B7C58FF09" lastPageId="11" lastPageNumber="574" pageId="1" pageNumber="564">
<subSubSection id="C3036541FFE0FFFBFF52F53B7F4BFBC5" box="[151,601,1051,1078]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="8BA636CAFFE0FFFBFF52F53B7F4BFBC5" blockId="1.[151,601,1051,1078]" box="[151,601,1051,1078]" pageId="1" pageNumber="564">
<heading id="D0EE81A6FFE0FFFBFF52F53B7F4BFBC5" bold="true" box="[151,601,1051,1078]" fontSize="11" level="1" pageId="1" pageNumber="564" reason="1">
<emphasis id="B96DEAD8FFE0FFFBFF52F53B7F4BFBC5" bold="true" box="[151,601,1051,1078]" pageId="1" pageNumber="564">
<taxonomicName id="4C194D49FFE0FFFBFF52F53B7F16FBC6" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[151,516,1051,1078]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE0FFFBFF52F53B7F16FBC6" bold="true" box="[151,516,1051,1078]" italics="true" pageId="1" pageNumber="564">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57A3FFE0FFFBFDC9F53B7F4BFBC5" box="[524,601,1051,1077]" pageId="1" pageNumber="564" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BA636CAFFE0FFFBFF52F5437C5FFB51" blockId="1.[151,1437,1123,2013]" pageId="1" pageNumber="564">
<materialsCitation id="3B713C97FFE0FFFBFF52F5437C5BFB51" country="Turkey" latitude="37.73" location="Inekli Lake" longLatPrecision="708" longitude="37.5" municipality="Golbasi district" pageId="1" pageNumber="564" specimenCount="1" stateProvince="Adiyaman" typeStatus="holotype">
<emphasis id="B96DEAD8FFE0FFFBFF52F5437C23FB8D" bold="true" box="[151,305,1123,1149]" pageId="1" pageNumber="564">
<typeStatus id="54A28868FFE0FFFBFF52F5437DC0FB8D" box="[151,210,1123,1149]" pageId="1" pageNumber="564">Type</typeStatus>
locality
</emphasis>
: 
<collectingCountry id="F30E765AFFE0FFFBFE84F5437C80FB8D" box="[321,402,1123,1149]" name="Turkey" pageId="1" pageNumber="564">Turkey</collectingCountry>
, 
<collectingRegion id="49DDF828FFE0FFFBFE58F5437F92FB8D" box="[413,640,1123,1149]" country="Turkey" name="Adiyaman" pageId="1" pageNumber="564">Adıyaman province</collectingRegion>
, 
<collectingMunicipality id="6BC2ACB0FFE0FFFBFD49F5437E2FFB8D" box="[652,829,1123,1149]" pageId="1" pageNumber="564">Gölbaşı district</collectingMunicipality>
, near 
<location id="8EC66011FFE0FFFBFC47F5457915FB8D" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE0FFFBFC47F5457915FB8D" box="[898,1031,1123,1152]" country="Turkey" latitude="37.73" longLatPrecision="708" longitude="37.5" municipality="Golbasi district" name="Inekli Lake" pageId="1" pageNumber="564" stateProvince="Adiyaman">İnekli Lake</location>
, ca. 
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<geoCoordinate id="EE2D500DFFE0FFFBFB5DF54379FAFB8E" box="[1176,1256,1123,1150]" degrees="37.50" direction="east" orientation="longitude" pageId="1" pageNumber="564" precision="555" value="37.5">37.50E</geoCoordinate>
(see 
<figureCitation id="13222A4FFFE0FFFBFAE3F5437879FB8E" box="[1318,1387,1123,1150]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="1" pageNumber="564">Fig. 7</figureCitation>
, red star, and 
<figureCitation id="13222A4FFFE0FFFBFF39F5A77C2DFB51" box="[252,319,1159,1185]" captionStart="FIGURES 8" captionStartId="10.[151,264,1491,1515]" captionTargetBox="[164,1423,194,1453]" captionTargetId="figure-30@10.[151,1436,181,1467]" captionTargetPageId="10" captionText="FIGURES 8a–8d. Around the type locality of Bidessus anatolicus adiyaman ssp. n. at the northern part of the İnekli Lake)." figureDoi="http://doi.org/10.5281/zenodo.5453463" httpUri="https://zenodo.org/record/5453463/files/figure.png" pageId="1" pageNumber="564">Fig. 8</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636CAFFE0FFFBFF02F58B7CA7FA89" blockId="1.[151,1437,1123,2013]" pageId="1" pageNumber="564">
<emphasis id="B96DEAD8FFE0FFFBFF02F58B7C7CFB35" bold="true" box="[199,366,1195,1221]" pageId="1" pageNumber="564">Type material</emphasis>
: 
<materialsCitation id="3B713C97FFE0FFFBFEB9F58B7ED6FAFD" collectingDate="2017-05-24" collectionCode="NMW" collectorName="Tasar, G. E." country="Turkey" determinerName="Fery" elevation="875" latitude="37.725" location="Inekli Lake" longLatPrecision="70" longitude="37.5" municipality="Golbasi district" pageId="1" pageNumber="564" specimenCount="1" specimenCount-male="1" stateProvince="Adiyaman" typeStatus="holotype">
<emphasis id="B96DEAD8FFE0FFFBFEB9F58B7CFBFB35" bold="true" box="[380,489,1195,1221]" pageId="1" pageNumber="564">
<typeStatus id="54A28868FFE0FFFBFEB9F58B7CFBFB35" box="[380,489,1195,1221]" pageId="1" pageNumber="564" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<specimenCount id="9D1FFD43FFE0FFFBFE3DF58C7F1DFB36" box="[504,527,1196,1222]" pageId="1" pageNumber="564" type="male">♂</specimenCount>
, “ 
<date id="FFA7100AFFE0FFFBFDE1F58B7FB7FB36" box="[548,677,1195,1222]" pageId="1" pageNumber="564" value="2017-05-24">
<collectingDate id="EFE3E9E2FFE0FFFBFDE1F58B7FB7FB36" box="[548,677,1195,1222]" pageId="1" pageNumber="564" value="2017-05-24">24.05.2017</collectingDate>
</date>
<collectingCountry id="F30E765AFFE0FFFBFD69F58B7FEEFB35" box="[684,764,1195,1221]" name="Turkey" pageId="1" pageNumber="564">Turkey</collectingCountry>
/ 
<collectingRegion id="49DDF828FFE0FFFBFCD5F58B7ED6FB36" box="[784,964,1195,1222]" country="Turkey" name="Adiyaman" pageId="1" pageNumber="564">Adıyaman prov.</collectingRegion>
/ 
<collectingMunicipality id="6BC2ACB0FFE0FFFBFC1CF58B799AFB35" box="[985,1160,1195,1221]" pageId="1" pageNumber="564">Gölbaşı district</collectingMunicipality>
, near 
<location id="8EC66011FFE0FFFBFB0FF58D7858FB35" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE0FFFBFB0FF58D7858FB35" box="[1226,1354,1195,1224]" country="Turkey" latitude="37.725" longLatPrecision="70" longitude="37.5" municipality="Golbasi district" name="Inekli Lake" pageId="1" pageNumber="564" stateProvince="Adiyaman">İnekli Lake</location>
/ small puddles / 
<geoCoordinate id="EE2D500DFFE0FFFBFEC3F5EF7C75FB1A" box="[262,359,1231,1258]" degrees="37.725" direction="north" orientation="latitude" pageId="1" pageNumber="564" precision="55" value="37.725">37.725N</geoCoordinate>
<geoCoordinate id="EE2D500DFFE0FFFBFEAAF5EF7CDCFB1A" box="[367,462,1231,1258]" degrees="37.500" direction="east" orientation="longitude" pageId="1" pageNumber="564" precision="55" value="37.5">37.500E</geoCoordinate>
/ 
<quantity id="4CE19B2FFFE0FFFBFE20F5EF7F3FFB1A" box="[485,557,1231,1258]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="1" pageNumber="564" unit="m" value="875.0">
<elevation id="0034D1F9FFE0FFFBFE20F5EF7F3FFB1A" box="[485,557,1231,1258]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="1" pageNumber="564" unit="m" value="875.0">875 m</elevation>
</quantity>
(Google-Earth), 
<collectorName id="26EC531CFFE0FFFBFD2AF5F07E7EFB19" box="[751,876,1231,1258]" pageId="1" pageNumber="564">Taşar, G.E.</collectorName>
leg.” [printed], “ 
<typeStatus id="54A28868FFE0FFFBFBE9F5EF7985FB19" box="[1068,1175,1231,1257]" pageId="1" pageNumber="564" type="holotype">Holotype</typeStatus>
/ 
<taxonomicName id="4C194D49FFE0FFFBFB6AF5EF7801FB19" authorityName="Sharp" authorityYear="1880" box="[1199,1299,1231,1257]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
/ anatolicus adiyaman 
<emphasis id="B96DEAD8FFE0FFFBFECEF5D37C47FAFD" bold="true" box="[267,341,1267,1293]" pageId="1" pageNumber="564">ssp. n.</emphasis>
/ Aykut, Taşar &amp; 
<determinerName id="7E7ADA0EFFE0FFFBFDDDF5D47F59FAFE" box="[536,587,1268,1294]" pageId="1" pageNumber="564">Fery</determinerName>
det. 2021” [red, printed] (
<collectionCode id="ED08AE0FFFE0FFFBFCABF5D37EAEFAFD" box="[878,956,1267,1293]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="564">NMW</collectionCode>
)
</materialsCitation>
. 
<emphasis id="B96DEAD8FFE0FFFBFC0BF5D37954FAFD" bold="true" box="[974,1094,1267,1293]" pageId="1" pageNumber="564">
<typeStatus id="54A28868FFE0FFFBFC0BF5D37954FAFD" box="[974,1094,1267,1293]" pageId="1" pageNumber="564" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D1FFD43FFE0FFFBFB90F5D37992FAFE" box="[1109,1152,1267,1294]" pageId="1" pageNumber="564" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D1FFD43FFE0FFFBFB49F5D379A7FAFE" box="[1164,1205,1267,1294]" pageId="1" pageNumber="564" type="female">4 ♀</specimenCount>
, 4 exs, same first la- bel as 
<typeStatus id="54A28868FFE0FFFBFF18F4377C51FAC1" box="[221,323,1303,1329]" pageId="1" pageNumber="564" type="holotype">holotype</typeStatus>
; 
<materialsCitation id="3B713C97FFE0FFFBFE88F4377CA3FA89" collectingDate="2013-07-16" collectionCode="CGT, CHF, CMA" collectorName="Tasar, G. E." country="Turkey" county="Golbasi district" elevation="875" latitude="37.744" location="Google-Earth" longLatPrecision="70" longitude="37.536" municipality="Azapli Lake" pageId="1" pageNumber="564" specimenCount="1" specimenCount-male="1" stateProvince="Adiyaman" typeStatus="paratype">
<specimenCount id="9D1FFD43FFE0FFFBFE88F4377C6BFAC2" box="[333,377,1303,1330]" pageId="1" pageNumber="564" type="male">1 ♂</specimenCount>
, 7 exs, “ 
<date id="FFA7100AFFE0FFFBFE10F4377F5AFAC1" box="[469,584,1303,1329]" pageId="1" pageNumber="564" value="2013-07-16">
<collectingDate id="EFE3E9E2FFE0FFFBFE10F4377F5AFAC1" box="[469,584,1303,1329]" pageId="1" pageNumber="564" value="2013-07-16">16.7.2013</collectingDate>
</date>
<collectingCountry id="F30E765AFFE0FFFBFD88F4377F8FFAC1" box="[589,669,1303,1329]" name="Turkey" pageId="1" pageNumber="564">Turkey</collectingCountry>
/ 
<collectingRegion id="49DDF828FFE0FFFBFD75F4377E76FAC2" box="[688,868,1303,1330]" country="Turkey" name="Adiyaman" pageId="1" pageNumber="564">Adıyaman prov.</collectingRegion>
/ 
<collectingCounty id="62C74E46FFE0FFFBFCBDF4377934FAC1" box="[888,1062,1303,1329]" pageId="1" pageNumber="564">Gölbaşı district</collectingCounty>
, near 
<collectingMunicipality id="6BC2ACB0FFE0FFFBFBA0F43779FCFAC1" box="[1125,1262,1303,1329]" pageId="1" pageNumber="564">Azaplı Lake</collectingMunicipality>
/ small puddles / 
<geoCoordinate id="EE2D500DFFE0FFFBFF63F41B7C15FAA5" box="[166,263,1339,1365]" degrees="37.744" direction="north" orientation="latitude" pageId="1" pageNumber="564" precision="55" value="37.744">37.744N</geoCoordinate>
<geoCoordinate id="EE2D500DFFE0FFFBFECBF41B7C7EFAA6" box="[270,364,1339,1366]" degrees="37.536" direction="east" orientation="longitude" pageId="1" pageNumber="564" precision="55" value="37.536">37.536E</geoCoordinate>
/ 
<quantity id="4CE19B2FFFE0FFFBFE47F41B7CDAFAA6" box="[386,456,1339,1366]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="1" pageNumber="564" unit="m" value="875.0">
<elevation id="0034D1F9FFE0FFFBFE47F41B7CDAFAA6" box="[386,456,1339,1366]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="1" pageNumber="564" unit="m" value="875.0">875 m</elevation>
</quantity>
(
<location id="8EC66011FFE0FFFBFE12F41B7F64FAA6" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE0FFFBFE12F41B7F64FAA6" box="[471,630,1339,1366]" country="Turkey" county="Golbasi district" latitude="37.744" longLatPrecision="70" longitude="37.536" municipality="Azapli Lake" name="Google-Earth" pageId="1" pageNumber="564" stateProvince="Adiyaman">Google-Earth</location>
), 
<collectorName id="26EC531CFFE0FFFBFD4DF41C7E16FAA5" box="[648,772,1339,1366]" pageId="1" pageNumber="564">Taşar, G.E.</collectorName>
leg.” [printed]; all 
<typeStatus id="54A28868FFE0FFFBFC1EF41C7954FAA6" box="[987,1094,1340,1366]" pageId="1" pageNumber="564" type="paratype">paratypes</typeStatus>
with a respective red 
<typeStatus id="54A28868FFE0FFFBFAF9F41C788EFAA6" box="[1340,1436,1340,1366]" pageId="1" pageNumber="564" type="paratype">paratype</typeStatus>
label (
<collectionCode id="ED08AE0FFFE0FFFBFF19F47F7C04FA89" box="[220,278,1375,1401]" pageId="1" pageNumber="564">CGT</collectionCode>
, 
<collectionCode id="ED08AE0FFFE0FFFBFEE7F47F7C4BFA89" box="[290,345,1375,1401]" pageId="1" pageNumber="564">CHF</collectionCode>
, 
<collectionCode id="ED08AE0FFFE0FFFBFEA1F47F7CBBFA89" box="[356,425,1375,1401]" pageId="1" pageNumber="564">CMA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036541FFE0FFF2FF02F4A37E3BFA55" lastPageId="8" lastPageNumber="571" pageId="1" pageNumber="564" type="description">
<paragraph id="8BA636CAFFE0FFFBFF02F4A379E8F9A2" blockId="1.[151,1437,1123,2013]" pageId="1" pageNumber="564">
<emphasis id="B96DEAD8FFE0FFFBFF02F4A37F92FA6D" bold="true" box="[199,640,1411,1437]" pageId="1" pageNumber="564">
Description of the 
<typeStatus id="54A28868FFE0FFFBFE6FF4A37F01FA6D" box="[426,531,1411,1437]" pageId="1" pageNumber="564" type="holotype">holotype</typeStatus>
. Habitus
</emphasis>
very broadly oval, sides of elytra centrally appearing almost inflated (
<figureCitation id="13222A4FFFE0FFFBFF65F4877DFDFA31" box="[160,239,1447,1473]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="1" pageNumber="564">Fig. 1a</figureCitation>
); maximum width of body before midlength of elytra, here outline of body strongly curved; posteriorly, in weakly curved line converging to shortly rounded apex. Body outline at junction of pronotum and elytra with weak discontinuity because outline of elytra not following outline of pronotum, instead being slightly convex at base of elytra (
<figureCitation id="13222A4FFFE0FFFBFEC2F7337C4FF9DD" box="[263,349,1555,1581]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="1" pageNumber="564">Figs 1a</figureCitation>
, 
<figureCitation id="13222A4FFFE0FFFBFEA9F7337C9BF9DD" box="[364,393,1555,1581]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="1" pageNumber="564">2a</figureCitation>
, 
<figureCitation id="13222A4FFFE0FFFBFE53F7337CA1F9DD" box="[406,435,1555,1581]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="1" pageNumber="564">3a</figureCitation>
). Dorsal surface shiny, brownish yellow on head and pronotum and dark brownish with brownish yellow pattern on elytra. Ventral surface (
<figureCitation id="13222A4FFFE0FFFBFD12F7177E37F9A1" box="[727,805,1591,1617]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="1" pageNumber="564">Fig. 1c</figureCitation>
) shiny and light brownish to large extent.
</paragraph>
<paragraph id="8BA636CAFFE0FFFBFF02F77B7F08F82D" blockId="1.[151,1437,1123,2013]" pageId="1" pageNumber="564">
<emphasis id="B96DEAD8FFE0FFFBFF02F77B7CD3F985" bold="true" box="[199,449,1627,1653]" pageId="1" pageNumber="564">Dorsal surface: Head</emphasis>
with anterior margin almost evenly rounded; posterior to margin with two distinct clypeal grooves above insertion of antennae; grooves connected by a weak impression, thus clypeus behind anterior margin appearing somewhat vaulted. Transverse occipital (cervical) line connecting hind margins of eyes (
<figureCitation id="13222A4FFFE0FFFBFACEF7837870F94D" box="[1291,1378,1699,1725]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="1" pageNumber="564">Figs 1a</figureCitation>
, 
<figureCitation id="13222A4FFFE0FFFBFAB7F783789DF94D" box="[1394,1423,1699,1725]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="1" pageNumber="564">3a</figureCitation>
). Punctation generally sparse, behind occipital line absent; diameter of punctures on clypeus and frons equalling more or less that of meshes of reticulation, distance between punctures about 4–5x that of diameter, in clypeal grooves punctures denser; here and near eyes interspersed with a few very small punctures; alongside inner margin of eyes with distinct line of larger punctures. Reticulation generally polygonal, but behind anterior margin (on slightly vaulted part) and behind occipital line (on vertex) with very narrow elongate transverse meshes. Surface brownish yellow with diffusely delimited more brownish areas above insertion of antennae and near eyes. First four antenno- meres brownish yellow, others brownish. Setation on dorsal surface almost imperceptible, only faintly recognisable in SEM photographs (
<figureCitation id="13222A4FFFE0FFFBFE4BF6E37CF6F82D" box="[398,484,1987,2013]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="1" pageNumber="564">Figs 2a</figureCitation>
, 
<figureCitation id="13222A4FFFE0FFFBFE2AF6E37F1EF82D" box="[495,524,1987,2013]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="1" pageNumber="564">3a</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F1B67883FE78" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F1B67C52FF40" bold="true" box="[199,320,150,176]" pageId="2" pageNumber="565">Pronotum</emphasis>
with distinct basal striae (
<figureCitation id="13222A4FFFE3FFF8FDADF1B67FAFFF40" box="[616,701,150,176]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Figs 1a</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FD0EF1B67FFAFF40" box="[715,744,150,176]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="2" pageNumber="565">3a</figureCitation>
), reaching anteriad beyond mid-length of pronotum; between striae not impressed; lateral sides of pronotum almost straight in posterior 3/4, converging anteriad, near anterior corners curved inwards; margins rimmed, rim near anterior corners becoming narrower. Punctation between striae very sparse and fine, only before base with coarser punctures, these more numerous and larger near striae; punctures between striae and lateral sides also sparse, but larger than finer punctures between striae; narrow stripe behind anterior margin with several small and densely arranged punctures. Reticulation on pronotum absent. Surface brownish yellow, lighter (more yellowish) than on head. Between striae before base dark brownish, in particular near striae.
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F0B27F46FD3C" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F0B27C01FE5C" bold="true" box="[199,275,402,428]" pageId="2" pageNumber="565">Elytra</emphasis>
with distinct basal striae (
<figureCitation id="13222A4FFFE3FFF8FDF0F0B27F9BFE5C" box="[565,649,402,428]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Figs 1a</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FD50F0B27FA0FE5C" box="[661,690,402,428]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">2a</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FD78F0B27FC8FE5C" box="[701,730,402,428]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="2" pageNumber="565">3a</figureCitation>
); these somewhat longer than pronotal striae, not reaching until midlength of elytra; sutural puncture lines present, posteriad reaching beyond midlength of elytra. Sides of elytra distinctly projecting over faintly rimmed side margin of elytra; thus, in dorsal view actual side margin only visible at shoulders and before apex. In lateral view elytral margin moderately ascending towards shoulders; epipleuron not visible until shoulders. Punctation between discal striae rather dense, distance between punctures more or less equal to their diameter, punctures more or less as large as punctures near pronotal striae; punctures between striae and elytral outline smaller and sparser. Reticulation absent on entire elytra. Surface blackish brown with three yellow bands; one band before midlength of elytra, second band short behind midlength, and third band short before apex; all bands interrupted medially (
<figureCitation id="13222A4FFFE3FFF8FE33F3927F54FD3C" box="[502,582,690,716]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F3F67FA1FCAD" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F3F67FA6FD00" bold="true" box="[199,692,726,752]" pageId="2" pageNumber="565">Ventral surface: Colour and reticulation:</emphasis>
Predominantly brownish (
<figureCitation id="13222A4FFFE3FFF8FC25F3F67923FD00" box="[992,1073,726,752]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Fig. 1c</figureCitation>
); head, prosternum, propleuron, hypomeron (= pronotal epipleuron), elytral epipleuron and legs more yellowish brown; surface shiny. Reticulation only on pro- and mesotibiae as well as on pro- and mesofemora recognisable, else absent. Setation generally sparse, on most parts of surface absent (cf. 
<figureCitation id="13222A4FFFE3FFF8FDE2F2627F68FCAC" box="[551,634,834,860]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Figs 1c</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FD42F2627FB4FCAD" box="[647,678,834,861]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">2b</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F2467F16FC1D" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F2467C15FC70" bold="true" box="[199,263,870,896]" pageId="2" pageNumber="565">Head</emphasis>
: Gula laterally with very few punctures. Genae with crease behind eyes, anteriorly with line of few small punctures. First and second antennomeres relatively long, third shorter and distinctly narrower, fourth much shorter than third, more or less as long as wide, fifth to ninth distinctly larger than fourth, tenth again narrower, eleventh slender and longer (
<figureCitation id="13222A4FFFE3FFF8FEB0F2F27CDEFC1C" box="[373,460,978,1004]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">Figs 2b</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FE1DF2F27CE5FC1D" box="[472,503,978,1005]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="2" pageNumber="565">3b</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F2D6789BFAFC" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F2D67C53FBE0" bold="true" box="[199,321,1014,1040]" pageId="2" pageNumber="565">Prothorax</emphasis>
: Prosternum anteromedially (before procoxae) somewhat vaulted and coarsely sculptured; laterally (before propleuron) smooth and shiny, without punctures. Hypomeron in anterior half laterally narrowly flattened, becoming broader anteriad; flat region with some coarse lengthy punctures and long setae (
<figureCitation id="13222A4FFFE3FFF8FB55F51E79F1FBA9" box="[1168,1251,1086,1113]" captionStart="FIGURES 3" captionStartId="5.[151,264,1920,1944]" captionTargetBox="[166,1421,197,1878]" captionTargetId="figure-17@5.[151,1436,181,1897]" captionTargetPageId="5" captionText="FIGURES 3a–3b. Bidessus anatolicus adiyaman ssp. n. (paratype): (a) oblique view on head, pronotum and anterior part of elytra; (b) ventral view on parts of head and prosternum (propleuron and hypomeron)." figureDoi="http://doi.org/10.5281/zenodo.5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="2" pageNumber="565">Fig. 3b</figureCitation>
); rest of hypomeron and propleuron smooth and shiny. Prosternal column ascending posteriad (as viewed if beetle turned upside down), between procoxae with a few transverse ridges. Prosternal process weakly inclined, broadly lanceolate, apex broadly rounded (
<figureCitation id="13222A4FFFE3FFF8FE5DF58A7CF9FB35" box="[408,491,1194,1221]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">Fig. 2b</figureCitation>
); with distinct broad longitudinal parallel-sided furrow; furrow formed by a few very coarse and closely arranged punctures, sides of furrow strongly rimmed; right and left of furrow process strongly descending. Apex of process reaching between mesocoxae and here touching anteromedial process of metaventrite.
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F4367DD6F94D" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
<emphasis id="B96DEAD8FFE3FFF8FF02F4367C45FAC0" bold="true" box="[199,343,1302,1328]" pageId="2" pageNumber="565">Pterothorax</emphasis>
: Metepisternum triangular, before anterior margin deeply impressed and with a few coarse punctures. Anteromedial process of metaventrite broad (more or less 1/3 of maximum width of mesocoxa), anteriorly reaching beyond middle of mesocoxae; distally with short broad furrow for reception of prosternal process. Suture between metaventrite and metacoxal plates almost imperceptible; metaventrite behind mesocoxae strongly rimmed; directly behind rim with line of punctures; rim with very distinct backwards directed extension, reaching anterior part of metacoxal plate and separating central part of metaventrite from its lateral wing. Metaventrite right and left of midline with line of coarse punctures, lines diverging posteriad, reaching until anterior margin of metacoxal plates; another line of coarse punctures alongside border to metacoxal plates. Metacoxal plates centrally with sev- eral coarse punctures. Metacoxal lines diverging anteriad, relatively short, maximum distance between lines about as long as midline; some coarse punctures alongside metacoxal lines. Posterior margin of metacoxal processes obliquely incised, each process postero-laterally without lobes, and thus not covering bases of metatrochanters (
<figureCitation id="13222A4FFFE3FFF8FAAAF75E7DA5F94D" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">Fig. 2b</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE3FFF8FF02F7E67864F849" blockId="2.[151,1437,150,1977]" pageId="2" pageNumber="565">
Epipleura anteriorly rather broad (broader than mesotibiae), becoming narrower at level of first abdominal ven- trite (
<figureCitation id="13222A4FFFE3FFF8FF10F7CA7C34F8F4" box="[213,294,1770,1796]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Fig. 1c</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FEF1F7CA7C41F8F5" box="[308,339,1770,1797]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">2b</figureCitation>
); with some small punctures alongside margins; without transverse carina at humeral angle (
<figureCitation id="13222A4FFFE3FFF8FAAAF7CA7DA5F8D9" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">Fig. 2b</figureCitation>
). Elytral surface between lateral outline and actual margin of elytra (alongside epipleura, where sides of elytra projecting over rimmed margin; see 
<figureCitation id="13222A4FFFE3FFF8FDECF6127F67F8BC" box="[553,629,1842,1868]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Fig. 1c</figureCitation>
) with only very few punctures except a line of small punctures alongside actual margin of elytra (
<figureCitation id="13222A4FFFE3FFF8FE67F6767CE4F881" box="[418,502,1878,1905]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">Fig. 2b</figureCitation>
). Notes: According to 
<bibRefCitation id="EF884B3BFFE3FFF8FD35F6767954F880" author="Miller, K. B. &amp; Bergsten, J." box="[752,1094,1878,1905]" pageId="2" pageNumber="565" refId="ref5205" refString="Miller, K. B. &amp; Bergsten, J. (2016) Diving beetles of the world: Systematics and biology of the Dytiscidae. Johns Hopkins Uni- versity Press, Baltimore, Maryland, ix + 320 pp." type="book" year="2016">Miller &amp; Bergsten (2016: 219)</bibRefCitation>
most genera of Bidessini have the metacoxae fused to the first visible abdominal ventrite; this character was not checked as it would have damaged a specimen. We can, however, state that the suture between the two structures is easily visible (see 
<figureCitation id="13222A4FFFE3FFF8FB2FF6BE782FF848" box="[1258,1341,1950,1976]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="2" pageNumber="565">Figs 1c</figureCitation>
, 
<figureCitation id="13222A4FFFE3FFF8FA8CF6BE787BF849" box="[1353,1385,1950,1977]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="2" pageNumber="565">2b</figureCitation>
).
</paragraph>
<caption id="DF666642FFE2FFF9FF52F6B779A6F824" ID-DOI="http://doi.org/10.5281/zenodo.5453449" ID-Zenodo-Dep="5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="3" pageNumber="566" startId="3.[151,264,1943,1967]" targetBox="[170,1413,182,1919]" targetPageId="3">
<paragraph id="8BA636CAFFE2FFF9FF52F6B779A6F824" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="566">
<emphasis id="B96DEAD8FFE2FFF9FF52F6B77C97F840" bold="true" box="[151,389,1943,1968]" pageId="3" pageNumber="566">FIGURES 1a–1c. a, c:</emphasis>
Habitus of 
<taxonomicName id="4C194D49FFE2FFF9FE3EF6B87E3FF840" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[507,813,1944,1968]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE2FFF9FE3EF6B87E3FF840" box="[507,813,1944,1968]" italics="true" pageId="3" pageNumber="566">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE2FFF9FCF6F6B87E64F840" bold="true" box="[819,886,1944,1968]" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A25E57A3FFE2FFF9FCF6F6B87E64F840" box="[819,886,1944,1968]" pageId="3" pageNumber="566" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(holotype): 
<emphasis id="B96DEAD8FFE2FFF9FC36F6B77902F85F" bold="true" box="[1011,1040,1943,1967]" pageId="3" pageNumber="566">(a)</emphasis>
in dorsal view; 
<emphasis id="B96DEAD8FFE2FFF9FB77F6B779DDF85F" bold="true" box="[1202,1231,1943,1967]" pageId="3" pageNumber="566">(c)</emphasis>
in ventral view (left hind-leg and last abdominal ventrites removed); 
<emphasis id="B96DEAD8FFE2FFF9FD41F69C7F88F824" bold="true" box="[644,666,1980,2004]" pageId="3" pageNumber="566">b:</emphasis>
Habitus of 
<taxonomicName id="4C194D49FFE2FFF9FCD4F69C79A6F824" authority="anatolicus Wewalka." authorityName="Wewalka." box="[785,1204,1980,2004]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="anatolicus">
<emphasis id="B96DEAD8FFE2FFF9FCD4F69C7958F824" box="[785,1098,1980,2004]" italics="true" pageId="3" pageNumber="566">Bidessus anatolicus anatolicus</emphasis>
Wewalka.
</taxonomicName>
</paragraph>
</caption>
<caption id="DF666642FFE5FFFEFF52F7F9785BF901" ID-DOI="http://doi.org/10.5281/zenodo.5453451" ID-Zenodo-Dep="5453451" box="[151,1353,1753,1777]" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="4" pageNumber="567" startId="4.[151,264,1753,1777]" targetBox="[166,1421,195,1712]" targetPageId="4">
<paragraph id="8BA636CAFFE5FFFEFF52F7F9785BF901" blockId="4.[151,1353,1753,1777]" box="[151,1353,1753,1777]" pageId="4" pageNumber="567">
<emphasis id="B96DEAD8FFE5FFFEFF52F7F97C44F901" bold="true" box="[151,342,1753,1777]" pageId="4" pageNumber="567">FIGURES 2a–2b.</emphasis>
Habitus of 
<taxonomicName id="4C194D49FFE5FFFEFE09F7F97FEDF901" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[460,767,1753,1777]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE5FFFEFE09F7F97FEDF901" box="[460,767,1753,1777]" italics="true" pageId="4" pageNumber="567">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE5FFFEFCC0F7F97E5AF901" bold="true" box="[773,840,1753,1777]" pageId="4" pageNumber="567">
<taxonomicNameLabel id="A25E57A3FFE5FFFEFCC0F7F97E5AF901" box="[773,840,1753,1777]" pageId="4" pageNumber="567" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(paratype): 
<emphasis id="B96DEAD8FFE5FFFEFC00F7F97EF1F901" bold="true" box="[965,995,1753,1777]" pageId="4" pageNumber="567">(a)</emphasis>
in dorsal view; 
<emphasis id="B96DEAD8FFE5FFFEFB43F7F979B7F901" bold="true" box="[1158,1189,1753,1777]" pageId="4" pageNumber="567">(b)</emphasis>
in ventral view.
</paragraph>
</caption>
<paragraph id="8BA636CAFFE5FFFEFF02F63F7C59F839" blockId="4.[151,1437,1823,1993]" pageId="4" pageNumber="567">
<emphasis id="B96DEAD8FFE5FFFEFF02F63F7C2EF8C9" bold="true" box="[199,316,1823,1849]" pageId="4" pageNumber="567">Abdomen</emphasis>
(
<figureCitation id="13222A4FFFE5FFFEFE8FF63F7C89F8CA" box="[330,411,1823,1850]" captionStart="FIGURES 2" captionStartId="4.[151,264,1753,1777]" captionTargetBox="[166,1421,195,1712]" captionTargetId="figure-30@4.[151,1436,181,1729]" captionTargetPageId="4" captionText="FIGURES 2a–2b. Habitus of Bidessus anatolicus adiyaman ssp. n. (paratype): (a) in dorsal view; (b) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5453451" httpUri="https://zenodo.org/record/5453451/files/figure.png" pageId="4" pageNumber="567">Fig. 2b</figureCitation>
) with first three ventrites rigidly connected; first and second laterally with coarse punctures, second behind metacoxal processes with several small setiferous punctures; third to fifth without large punctures but with several sparsely distributed small setiferous punctures, this same 
<typeStatus id="54A28868FFE5FFFEFC67F6487EC0F872" box="[930,978,1896,1922]" pageId="4" pageNumber="567">type</typeStatus>
of puncture being denser on last ventrite. Fourth and fifth ventrite centrally with a tuft of long setae (which can easily be rubbed off). Apex of last ventrite weakly pointed.
</paragraph>
<caption id="DF666642FFE4FFFFFF52F6A07913F84D" ID-DOI="http://doi.org/10.5281/zenodo.5453453" ID-Zenodo-Dep="5453453" httpUri="https://zenodo.org/record/5453453/files/figure.png" pageId="5" pageNumber="568" startId="5.[151,264,1920,1944]" targetBox="[166,1421,197,1878]" targetPageId="5">
<paragraph id="8BA636CAFFE4FFFFFF52F6A07913F84D" blockId="5.[151,1436,1920,1981]" pageId="5" pageNumber="568">
<emphasis id="B96DEAD8FFE4FFFFFF52F6A07C45F869" bold="true" box="[151,343,1920,1945]" pageId="5" pageNumber="568">FIGURES 3a–3b.</emphasis>
<taxonomicName id="4C194D49FFE4FFFFFE9BF6A17F81F869" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[350,659,1921,1945]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE4FFFFFE9BF6A17F81F869" box="[350,659,1921,1945]" italics="true" pageId="5" pageNumber="568">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE4FFFFFD5FF6A17FCCF869" bold="true" box="[666,734,1921,1945]" pageId="5" pageNumber="568">
<taxonomicNameLabel id="A25E57A3FFE4FFFFFD5FF6A17FCCF869" box="[666,734,1921,1945]" pageId="5" pageNumber="568" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(paratype): 
<emphasis id="B96DEAD8FFE4FFFFFC98F6A07E69F868" bold="true" box="[861,891,1920,1944]" pageId="5" pageNumber="568">(a)</emphasis>
oblique view on head, pronotum and anterior part of elytra; 
<emphasis id="B96DEAD8FFE4FFFFFF1BF6847DEFF84C" bold="true" box="[222,253,1956,1980]" pageId="5" pageNumber="568">(b)</emphasis>
ventral view on parts of head and prosternum (propleuron and hypomeron).
</paragraph>
</caption>
<paragraph id="8BA636CAFFE7FFFCFF02F1B77E47FF09" blockId="6.[151,1437,151,399]" pageId="6" pageNumber="569">
<emphasis id="B96DEAD8FFE7FFFCFF02F1B77DEDFF41" bold="true" box="[199,255,151,177]" pageId="6" pageNumber="569">Legs</emphasis>
: Protibiae broadened distally; inner margin curved. First two pro- and mesotarsomeres with distally broad- ly enlarged sucking setae (
<figureCitation id="13222A4FFFE7FFFCFE05F19B7F01FF25" box="[448,531,187,213]" captionStart="FIGURES 4" captionStartId="6.[151,264,1998,2022]" captionTargetBox="[166,1421,431,1958]" captionTargetId="figure-136@6.[151,1436,417,1975]" captionTargetPageId="6" captionText="FIGURES 4a–4b. Ventral view on protarsomeres of Bidessus anatolicus adiyaman ssp. n. (paratypes): (a) male; (b) female." figureDoi="http://doi.org/10.5281/zenodo.5453455" httpUri="https://zenodo.org/record/5453455/files/figure.png" pageId="6" pageNumber="569">Fig. 4a</figureCitation>
). Protarsal claws simple. Metatibiae slender and slightly curved. First metatarso- mere slightly shorter than second and third tarsomere together.
</paragraph>
<paragraph id="8BA636CAFFE7FFFCFF02F0237CFCFE79" blockId="6.[151,1437,151,399]" pageId="6" pageNumber="569">
<emphasis id="B96DEAD8FFE7FFFCFF02F0237C64FEED" bold="true" box="[199,374,259,285]" pageId="6" pageNumber="569">Measurements</emphasis>
: TL: 
<quantity id="4CE19B2FFFE7FFFCFE73F0227F59FEED" box="[438,587,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="6" pageNumber="569" unit="mm" value="1.725" valueMax="1.8" valueMin="1.65">1.65–1.8 mm</quantity>
(
<quantity id="4CE19B2FFFE7FFFCFD9FF0227FD1FEED" box="[602,707,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="569" unit="mm" value="1.75">1.75 mm</quantity>
), MW: 1.0– 
<quantity id="4CE19B2FFFE7FFFCFC8FF0237E8CFEED" box="[842,926,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="569" unit="mm" value="1.1">1.1 mm</quantity>
(
<quantity id="4CE19B2FFFE7FFFCFC68F0227904FEED" box="[941,1046,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="569" unit="mm" value="1.05">1.05 mm</quantity>
), MW 
<subScript id="179D348FFFE7FFFCFB99F0357964FED3" attach="left" box="[1116,1142,277,291]" fontSize="6" pageId="6" pageNumber="569">prot</subScript>
: 
<quantity id="4CE19B2FFFE7FFFCFB40F0227808FEED" box="[1157,1306,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="6" pageNumber="569" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.8–0.85 mm</quantity>
(0.85), TL/ MW: 1.62–1.67 (1.67), MW/MW 
<subScript id="179D348FFFE7FFFCFDD7F0197F3EFEB7" attach="left" box="[530,556,313,327]" fontSize="6" pageId="6" pageNumber="569">prot</subScript>
: 1.26–1.34 (1.27); n = 8 (values of the 
<typeStatus id="54A28868FFE7FFFCFC28F007795DFEB1" box="[1005,1103,295,321]" pageId="6" pageNumber="569" type="holotype">holotype</typeStatus>
in brackets). For comparison values of the nominotypical ssp.: TL: 
<quantity id="4CE19B2FFFE7FFFCFD80F06A7FCEFE95" box="[581,732,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.6749999999999998" metricValueMax="1.7" metricValueMin="1.65" pageId="6" pageNumber="569" unit="mm" value="1.6749999999999998" valueMax="1.7" valueMin="1.65">1.65–1.7 mm</quantity>
, MW: 0.95–1.0 mm, MW 
<subScript id="179D348FFFE7FFFCFBC7F07D790EFE9B" attach="right" box="[1026,1052,349,363]" fontSize="6" pageId="6" pageNumber="569">prot</subScript>
: 
<quantity id="4CE19B2FFFE7FFFCFBE9F06A79BFFE95" box="[1068,1197,330,357]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="6" pageNumber="569" unit="m" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.8 m</quantity>
, TL/MW: 1.68–1.76, MW/MW 
<subScript id="179D348FFFE7FFFCFEC3F0A17C32FE7F" attach="left" box="[262,288,385,399]" fontSize="6" pageId="6" pageNumber="569">prot</subScript>
: 1.25–1.31; n = 6.
</paragraph>
<caption id="DF666642FFE7FFFCFF52F6EE7899F817" ID-DOI="http://doi.org/10.5281/zenodo.5453455" ID-Zenodo-Dep="5453455" box="[151,1419,1998,2023]" httpUri="https://zenodo.org/record/5453455/files/figure.png" pageId="6" pageNumber="569" startId="6.[151,264,1998,2022]" targetBox="[166,1421,431,1958]" targetPageId="6">
<paragraph id="8BA636CAFFE7FFFCFF52F6EE7899F817" blockId="6.[151,1419,1998,2023]" box="[151,1419,1998,2023]" pageId="6" pageNumber="569">
<emphasis id="B96DEAD8FFE7FFFCFF52F6EE7C47F817" bold="true" box="[151,341,1998,2023]" pageId="6" pageNumber="569">FIGURES 4a–4b.</emphasis>
Ventral view on protarsomeres of 
<taxonomicName id="4C194D49FFE7FFFCFD71F6EF7EF4F817" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[692,998,1999,2023]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE7FFFCFD71F6EF7EF4F817" box="[692,998,1999,2023]" italics="true" pageId="6" pageNumber="569">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE7FFFCFC28F6EF7922F817" bold="true" box="[1005,1072,1999,2023]" pageId="6" pageNumber="569">
<taxonomicNameLabel id="A25E57A3FFE7FFFCFC28F6EF7922F817" box="[1005,1072,1999,2023]" pageId="6" pageNumber="569" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(paratypes): 
<emphasis id="B96DEAD8FFE7FFFCFB72F6EE79C6F816" bold="true" box="[1207,1236,1998,2022]" pageId="6" pageNumber="569">(a)</emphasis>
male; 
<emphasis id="B96DEAD8FFE7FFFCFADCF6EE782AF816" bold="true" box="[1305,1336,1998,2022]" pageId="6" pageNumber="569">(b)</emphasis>
female.
</paragraph>
</caption>
<caption id="DF666642FFE6FFFDFF52F67E7839F812" ID-DOI="http://doi.org/10.5281/zenodo.5453457" ID-Zenodo-Dep="5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="7" pageNumber="570" startId="7.[151,264,1886,1910]" targetBox="[165,1422,181,1862]" targetPageId="7">
<paragraph id="8BA636CAFFE6FFFDFF52F67E7839F812" blockId="7.[151,1437,1886,2018]" pageId="7" pageNumber="570">
<emphasis id="B96DEAD8FFE6FFFDFF52F67E7C47F886" bold="true" box="[151,341,1886,1910]" pageId="7" pageNumber="570">FIGURES 5a–5h.</emphasis>
Male genitalia of: 
<emphasis id="B96DEAD8FFE6FFFDFDD7F67E7F24F886" bold="true" box="[530,566,1886,1910]" pageId="7" pageNumber="570">a–c</emphasis>
: 
<taxonomicName id="4C194D49FFE6FFFDFD86F67E7E66F886" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[579,884,1886,1910]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE6FFFDFD86F67E7E66F886" box="[579,884,1886,1910]" italics="true" pageId="7" pageNumber="570">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE6FFFDFCBCF67E7EA9F886" bold="true" box="[889,955,1886,1910]" pageId="7" pageNumber="570">
<taxonomicNameLabel id="A25E57A3FFE6FFFDFCBCF67E7EA9F886" box="[889,955,1886,1910]" pageId="7" pageNumber="570" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(holotype): 
<emphasis id="B96DEAD8FFE6FFFDFBF2F67E7946F886" bold="true" box="[1079,1108,1886,1910]" pageId="7" pageNumber="570">(a)</emphasis>
median lobe in ventral view; 
<emphasis id="B96DEAD8FFE6FFFDFAB8F67E788EF886" bold="true" box="[1405,1436,1886,1910]" pageId="7" pageNumber="570">(b)</emphasis>
median lobe in lateral view; 
<emphasis id="B96DEAD8FFE6FFFDFE7DF6A27CC6F86A" bold="true" box="[440,468,1922,1946]" pageId="7" pageNumber="570">(c)</emphasis>
left paramere; 
<emphasis id="B96DEAD8FFE6FFFDFDABF6A27F8DF86A" bold="true" box="[622,671,1922,1946]" pageId="7" pageNumber="570">d–h:</emphasis>
<emphasis id="B96DEAD8FFE6FFFDFD63F6A27E9AF86A" box="[678,904,1922,1946]" italics="true" pageId="7" pageNumber="570">
<taxonomicName id="4C194D49FFE6FFFDFD63F6A27FEDF86A" authorityName="Sharp" authorityYear="1880" box="[678,767,1922,1946]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. 
<taxonomicName id="4C194D49FFE6FFFDFCDBF6A27E9AF86A" authorityName="Wewalka" authorityYear="1972" box="[798,904,1922,1946]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
: 
<emphasis id="B96DEAD8FFE6FFFDFC53F6A27EA7F86A" bold="true" box="[918,949,1922,1946]" pageId="7" pageNumber="570">(d)</emphasis>
median lobe in ventral view; 
<emphasis id="B96DEAD8FFE6FFFDFB26F6A27812F86A" bold="true" box="[1251,1280,1922,1946]" pageId="7" pageNumber="570">(e)</emphasis>
median lobe in lateral view; 
<emphasis id="B96DEAD8FFE6FFFDFEDEF6867C28F84E" bold="true" box="[283,314,1958,1982]" pageId="7" pageNumber="570">(h)</emphasis>
left paramere (all adopted from 
<bibRefCitation id="EF884B3BFFE6FFFDFD47F6867E53F84E" author="Wewalka, G." box="[642,833,1958,1982]" pageId="7" pageNumber="570" pagination="72 - 73" refId="ref5565" refString="Wewalka, G. (1972) Eine neue Art der Gattung Bidessus Sharp aus Kleinasien (Dytiscidae, Col.). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 23 (1971), 72 - 73." type="journal article" year="1972">Wewalka 1972: 73</bibRefCitation>
); 
<emphasis id="B96DEAD8FFE6FFFDFC91F6867E64F84E" bold="true" box="[852,886,1958,1982]" pageId="7" pageNumber="570">f, g</emphasis>
: 
<emphasis id="B96DEAD8FFE6FFFDFC46F6867977F84E" box="[899,1125,1958,1982]" italics="true" pageId="7" pageNumber="570">
<taxonomicName id="4C194D49FFE6FFFDFC46F6867ECEF84E" authorityName="Sharp" authorityYear="1880" box="[899,988,1958,1982]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. 
<taxonomicName id="4C194D49FFE6FFFDFC3EF6867977F84E" authorityName="Wewalka" authorityYear="1972" box="[1019,1125,1958,1982]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
(specimen from Kumluca): 
<emphasis id="B96DEAD8FFE6FFFDFA46F686788EF84E" bold="true" box="[1411,1436,1958,1982]" pageId="7" pageNumber="570">(f)</emphasis>
median lobe in lateral view; 
<emphasis id="B96DEAD8FFE6FFFDFE7DF6EA7CC5F812" bold="true" box="[440,471,1994,2018]" pageId="7" pageNumber="570">(h)</emphasis>
left paramere (arrows indicate direction of view on ventral surface of median lobe).
</paragraph>
</caption>
<paragraph id="8BA636CAFFE9FFF2FF02F1B77821FF41" blockId="8.[151,1437,150,285]" box="[199,1331,150,177]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF02F1B77C1DFF41" bold="true" box="[199,271,151,177]" pageId="8" pageNumber="571">Males</emphasis>
: median lobe in ventral and lateral view as in 
<figureCitation id="13222A4FFFE9FFF2FCD0F1B77E81FF41" box="[789,915,150,177]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="8" pageNumber="571">Figs 5a, 5b</figureCitation>
; parameres two-segmented (
<figureCitation id="13222A4FFFE9FFF2FB11F1B77837FF41" box="[1236,1317,150,177]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="8" pageNumber="571">Fig. 5c</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF02F19B7CA7FEED" blockId="8.[151,1437,150,285]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF02F19B7C3BFF25" bold="true" box="[199,297,187,213]" pageId="8" pageNumber="571">Females</emphasis>
: Gonocoxosternum and gonocoxae as in 
<figureCitation id="13222A4FFFE9FFF2FCCEF19B7E82FF25" box="[779,912,187,213]" captionStart="FIGURES 6" captionStartId="8.[151,264,1060,1084]" captionTargetBox="[151,1436,309,1037]" captionTargetId="figure-86@8.[151,1436,309,1037]" captionTargetPageId="8" captionText="FIGURES 6a–6b. Bidessus anatolicus adiyaman ssp. n. (female paratype): (a) gonocoxosternum; (b) gonocoxae." figureDoi="http://doi.org/10.5281/zenodo.5453459" httpUri="https://zenodo.org/record/5453459/files/figure.png" pageId="8" pageNumber="571">Figs 6a, 6b</figureCitation>
. First three pro- and mesotarsomeres slightly narrower than in males and without distinctly enlarged sucking setae (
<figureCitation id="13222A4FFFE9FFF2FC66F1FF7EE5FF0A" box="[931,1015,223,250]" captionStart="FIGURES 4" captionStartId="6.[151,264,1998,2022]" captionTargetBox="[166,1421,431,1958]" captionTargetId="figure-136@6.[151,1436,417,1975]" captionTargetPageId="6" captionText="FIGURES 4a–4b. Ventral view on protarsomeres of Bidessus anatolicus adiyaman ssp. n. (paratypes): (a) male; (b) female." figureDoi="http://doi.org/10.5281/zenodo.5453455" httpUri="https://zenodo.org/record/5453455/files/figure.png" pageId="8" pageNumber="571">Fig. 4b</figureCitation>
); other external morphological differences to males not found.
</paragraph>
<caption id="DF666642FFE9FFF2FF52F5047833FBCD" ID-DOI="http://doi.org/10.5281/zenodo.5453459" ID-Zenodo-Dep="5453459" box="[151,1313,1060,1085]" httpUri="https://zenodo.org/record/5453459/files/figure.png" pageId="8" pageNumber="571" startId="8.[151,264,1060,1084]" targetBox="[151,1436,309,1037]" targetPageId="8">
<paragraph id="8BA636CAFFE9FFF2FF52F5047833FBCD" blockId="8.[151,1313,1060,1085]" box="[151,1313,1060,1085]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF52F5047C47FBCD" bold="true" box="[151,341,1060,1085]" pageId="8" pageNumber="571">FIGURES 6a–6b.</emphasis>
<taxonomicName id="4C194D49FFE9FFF2FE99F5057F9CFBCD" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[348,654,1061,1085]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE9FFF2FE99F5057F9CFBCD" box="[348,654,1061,1085]" italics="true" pageId="8" pageNumber="571">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE9FFF2FD50F5057FCAFBCD" bold="true" box="[661,728,1061,1085]" pageId="8" pageNumber="571">
<taxonomicNameLabel id="A25E57A3FFE9FFF2FD50F5057FCAFBCD" box="[661,728,1061,1085]" pageId="8" pageNumber="571" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(female paratype): 
<emphasis id="B96DEAD8FFE9FFF2FC65F5047EACFBCC" bold="true" box="[928,958,1060,1084]" pageId="8" pageNumber="571">(a)</emphasis>
gonocoxosternum; 
<emphasis id="B96DEAD8FFE9FFF2FB42F50479B4FBCC" bold="true" box="[1159,1190,1060,1084]" pageId="8" pageNumber="571">(b)</emphasis>
gonocoxae.
</paragraph>
</caption>
<paragraph id="8BA636CAFFE9FFF2FF02F54B794AFB3E" blockId="8.[151,1437,1131,1446]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF02F54B7C55FB75" bold="true" box="[199,327,1131,1157]" pageId="8" pageNumber="571">Variability</emphasis>
: Apart from size the only variation observed was in the elytral pattern: in a few specimens the yel- lowish bands are somewhat more developed than in the 
<typeStatus id="54A28868FFE9FFF2FCDCF5AF7E69FB59" box="[793,891,1167,1193]" pageId="8" pageNumber="571" type="holotype">holotype</typeStatus>
and the first band is connected with the second band by a narrow longitudinal strip (more or less in prolongation of the elytral striae).
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF02F5F77E3BFA55" blockId="8.[151,1437,1131,1446]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF02F5F77CE1FB01" bold="true" box="[199,499,1239,1265]" pageId="8" pageNumber="571">Collecting circumstances</emphasis>
: On 
<date id="FFA7100AFFE9FFF2FDF4F5F77FB3FB02" box="[561,673,1239,1266]" pageId="8" pageNumber="571" value="2017-05-24">24.5.2017</date>
<typeStatus id="54A28868FFE9FFF2FD6EF5F77E04FB01" box="[683,790,1239,1265]" pageId="8" pageNumber="571" type="paratype">paratypes</typeStatus>
of the new taxon were found in small puddles near the İnekli Lake (altitude according to Google Earth ca. 
<quantity id="4CE19B2FFFE9FFF2FD2CF5DB7E21FAE5" box="[745,819,1275,1301]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="8" pageNumber="571" unit="m" value="877.0">877 m</quantity>
) together with 
<specimenCount id="9D1FFD43FFE9FFF2FC21F5DB7988FAE5" box="[996,1178,1275,1301]" pageId="8" pageNumber="571" type="generic">three specimens</specimenCount>
of 
<taxonomicName id="4C194D49FFE9FFF2FB06F5DB7CA5FACA" authority="(Reiche &amp; Saulcy, 1855)" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="exornatus">
<emphasis id="B96DEAD8FFE9FFF2FB06F5DB788EFAE5" box="[1219,1436,1275,1301]" italics="true" pageId="8" pageNumber="571">Bidessus exornatus</emphasis>
(Reiche &amp; Saulcy, 1855)
</taxonomicName>
. The puddles result from peat cutting which leaves holes that fill with water (see 
<figureCitation id="13222A4FFFE9FFF2FAA9F43F7DF2FAAE" captionStart="FIGURES 8" captionStartId="10.[151,264,1491,1515]" captionTargetBox="[164,1423,194,1453]" captionTargetId="figure-30@10.[151,1436,181,1467]" captionTargetPageId="10" captionText="FIGURES 8a–8d. Around the type locality of Bidessus anatolicus adiyaman ssp. n. at the northern part of the İnekli Lake)." figureDoi="http://doi.org/10.5281/zenodo.5453463" httpUri="https://zenodo.org/record/5453463/files/figure.png" pageId="8" pageNumber="571">Figs 8a–8d</figureCitation>
). The other collecting site (near Azaplı Lake; altitude ca. 
<quantity id="4CE19B2FFFE9FFF2FC92F4637E8FFAAD" box="[855,925,1347,1373]" metricMagnitude="2" metricUnit="m" metricValue="8.78" pageId="8" pageNumber="571" unit="m" value="878.0">878 m</quantity>
) is very similar and the distance between both is only about 
<quantity id="4CE19B2FFFE9FFF2FEF7F4477C96FA71" box="[306,388,1383,1409]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="571" unit="km" value="3.8">3.8 km</quantity>
; both sites are connected by a small river slowly running in marshy ground (largely used for peat cutting) and a man-made canal connecting both lakes.
</paragraph>
</subSubSection>
<subSubSection id="C3036541FFE9FFF1FF52F4F37C58FF09" lastPageId="11" lastPageNumber="574" pageId="8" pageNumber="571" type="diagnosis">
<paragraph id="8BA636CAFFE9FFF2FF52F4F37900F9E1" blockId="8.[151,1436,1491,1553]" pageId="8" pageNumber="571">
<emphasis id="B96DEAD8FFE9FFF2FF52F4F37C8DFA1D" bold="true" box="[151,415,1491,1517]" pageId="8" pageNumber="571">Differential diagnosis.</emphasis>
We compared specimens of the new taxon with the following specimens of the other subspecies of 
<taxonomicName id="4C194D49FFE9FFF2FEC8F4D87CB3F9E1" authorityName="Wewalka" authorityYear="1972" box="[269,417,1527,1553]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B96DEAD8FFE9FFF2FEC8F4D87CB3F9E1" box="[269,417,1527,1553]" italics="true" pageId="8" pageNumber="571">B. anatolicus</emphasis>
</taxonomicName>
(all specimens in CHF, except that from Kızılot: CAP):
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF52F71F7E8EF951" blockId="8.[151,1437,1599,1878]" pageId="8" pageNumber="571">
- 
<emphasis id="B96DEAD8FFE9FFF2FF03F71F7CD3F9A9" box="[198,449,1599,1625]" italics="true" pageId="8" pageNumber="571">
<taxonomicName id="4C194D49FFE9FFF2FF03F71F7C3AF9A9" authorityName="Sharp" authorityYear="1880" box="[198,296,1599,1625]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. 
<taxonomicName id="4C194D49FFE9FFF2FE89F71F7CD3F9A9" authorityName="Wewalka" authorityYear="1972" box="[332,449,1599,1625]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
: 
<materialsCitation id="3B713C97FFE9FFF2FE15F71F7E8AF951" collectingDate="1968-05-26" collectingDateMax="2008-07" collectingDateMin="1968-05-26" collectorName="Wewalka &amp; Putz" latitude="36.89" location="Antalya" longLatPrecision="711" longitude="30.71" pageId="8" pageNumber="571" specimenCount="1" stateProvince="Antalya">
6 exs from near Kumluca (
<figureCitation id="13222A4FFFE9FFF2FD3AF71F7E56F9A9" box="[767,836,1599,1625]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
, green circle 1), 
<date id="FFA7100AFFE9FFF2FBC7F71F7966F9A9" box="[1026,1140,1599,1625]" pageId="8" pageNumber="571" value="1983-07-28">
<collectingDate id="EFE3E9E2FFE9FFF2FBC7F71F7966F9A9" box="[1026,1140,1599,1625]" pageId="8" pageNumber="571" value="1983-07-28">28.7.1983</collectingDate>
</date>
, 
<collectorName id="26EC531CFFE9FFF2FB45F71F79F5F9A9" box="[1152,1255,1599,1625]" pageId="8" pageNumber="571">Wewalka</collectorName>
leg.; 
<specimenCount id="9D1FFD43FFE9FFF2FAE2F71F784DF9A9" box="[1319,1375,1599,1625]" pageId="8" pageNumber="571" type="generic">1 ex.</specimenCount>
from near Kızılot (
<figureCitation id="13222A4FFFE9FFF2FE9CF7437C8FF98D" box="[345,413,1635,1661]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
, green circle 3), 
<date id="FFA7100AFFE9FFF2FD92F7437FD5F98D" box="[599,711,1635,1661]" pageId="8" pageNumber="571" value="2008-07">
<collectingDate id="EFE3E9E2FFE9FFF2FD92F7437FD5F98D" box="[599,711,1635,1661]" pageId="8" pageNumber="571" value="2008-07">VII. 2008</collectingDate>
</date>
, 
<collectorName id="26EC531CFFE9FFF2FD14F7437E11F98D" box="[721,771,1635,1661]" pageId="8" pageNumber="571">Pütz</collectorName>
leg.; the 
<typeStatus id="54A28868FFE9FFF2FCAFF7437E88F98D" box="[874,922,1635,1661]" pageId="8" pageNumber="571">type</typeStatus>
locality of this subspecies is near 
<collectingRegion id="49DDF828FFE9FFF2FAD3F7437863F98D" box="[1302,1393,1635,1661]" country="Turkey" name="Antalya" pageId="8" pageNumber="571">Antalya</collectingRegion>
, ca. 
<geoCoordinate id="EE2D500DFFE9FFF2FF03F7A77C0BF951" box="[198,281,1671,1697]" degrees="36.89" direction="north" orientation="latitude" pageId="8" pageNumber="571" precision="555" value="36.89">36.89N</geoCoordinate>
<geoCoordinate id="EE2D500DFFE9FFF2FEE5F7A77C61F951" box="[288,371,1671,1697]" degrees="30.71" direction="east" orientation="longitude" pageId="8" pageNumber="571" precision="555" value="30.71">30.71E</geoCoordinate>
, 
<date id="FFA7100AFFE9FFF2FEBBF7A77CE2F952" box="[382,496,1671,1698]" pageId="8" pageNumber="571" value="1968-05-26">
<collectingDate id="EFE3E9E2FFE9FFF2FEBBF7A77CE2F952" box="[382,496,1671,1698]" pageId="8" pageNumber="571" value="1968-05-26">26.5.1968</collectingDate>
</date>
, 
<collectorName id="26EC531CFFE9FFF2FE39F7A77F71F951" box="[508,611,1671,1697]" pageId="8" pageNumber="571">Wewalka</collectorName>
leg. (
<figureCitation id="13222A4FFFE9FFF2FD66F7A77FF5F951" box="[675,743,1671,1697]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
, green circle 2)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF52F78B7F0BF8FD" blockId="8.[151,1437,1599,1878]" pageId="8" pageNumber="571">
- 
<emphasis id="B96DEAD8FFE9FFF2FF03F78B7C8CF935" box="[198,414,1707,1733]" italics="true" pageId="8" pageNumber="571">
<taxonomicName id="4C194D49FFE9FFF2FF03F78B7C3AF935" authorityName="Sharp" authorityYear="1880" box="[198,296,1707,1733]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. kadmos
</emphasis>
: 2 exs from Park Ha’ Yarden, 
<collectingCountry id="F30E765AFFE9FFF2FD2CF78B7E38F935" box="[745,810,1707,1733]" name="Israel" pageId="8" pageNumber="571">Israel</collectingCountry>
, 
<date id="FFA7100AFFE9FFF2FCF6F78B7EB4F935" box="[819,934,1707,1733]" pageId="8" pageNumber="571" value="1986-12-04">12.4.1986</date>
, 
<materialsCitation id="3B713C97FFE9FFF2FC75F78B7C2AF91A" collectingDate="1985-08-27" collectorName="Jach" country="Jordan" location="En Jezer" pageId="8" pageNumber="571" specimenCount="1">
<collectorName id="26EC531CFFE9FFF2FC75F78B7EF0F935" box="[944,994,1707,1733]" pageId="8" pageNumber="571">Jäch</collectorName>
leg.; 
<specimenCount id="9D1FFD43FFE9FFF2FBDBF78B7946F935" box="[1054,1108,1707,1733]" pageId="8" pageNumber="571" type="generic">1 ex.</specimenCount>
<location id="8EC66011FFE9FFF2FB9CF78B79ABF935" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE9FFF2FB9CF78B79ABF935" box="[1113,1209,1707,1733]" country="Jordan" name="En Jezer" pageId="8" pageNumber="571">En Jezer</location>
, 
<collectingCountry id="F30E765AFFE9FFF2FB06F78B781CF935" box="[1219,1294,1707,1733]" name="Jordan" pageId="8" pageNumber="571">Jordan</collectingCountry>
delta, 
<collectingCountry id="F30E765AFFE9FFF2FA92F78B788AF935" box="[1367,1432,1707,1733]" name="Israel" pageId="8" pageNumber="571">Israel</collectingCountry>
, 
<date id="FFA7100AFFE9FFF2FF03F7EF7C2AF91A" box="[198,312,1743,1770]" pageId="8" pageNumber="571" value="1985-08-27">
<collectingDate id="EFE3E9E2FFE9FFF2FF03F7EF7C2AF91A" box="[198,312,1743,1770]" pageId="8" pageNumber="571" value="1985-08-27">27.8.1985</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B713C97FFE9FFF2FE80F7EF7980F91A" box="[325,1170,1743,1770]" collectingDate="1985-07-16" collectorName="Jach" country="Israel" location="En Jalabina" municipality="Lake Tiberias" pageId="8" pageNumber="571" specimenCount="1">
<collectorName id="26EC531CFFE9FFF2FE80F7EF7C65F919" box="[325,375,1743,1769]" pageId="8" pageNumber="571">Jäch</collectorName>
leg.; 
<specimenCount id="9D1FFD43FFE9FFF2FE7DF7EF7CFDF919" box="[440,495,1743,1769]" pageId="8" pageNumber="571" type="generic">1 ex.</specimenCount>
from 
<location id="8EC66011FFE9FFF2FDF0F7EF7FA9F919" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE9FFF2FDF0F7EF7FA9F919" box="[565,699,1743,1769]" country="Israel" municipality="Lake Tiberias" name="En Jalabina" pageId="8" pageNumber="571">En Jalabina</location>
, north of 
<collectingMunicipality id="6BC2ACB0FFE9FFF2FCEFF7EF7ED5F919" box="[810,967,1743,1769]" pageId="8" pageNumber="571">Lake Tiberias</collectingMunicipality>
, 
<collectingCountry id="F30E765AFFE9FFF2FC16F7EF7906F919" box="[979,1044,1743,1769]" name="Israel" pageId="8" pageNumber="571">Israel</collectingCountry>
, 
<date id="FFA7100AFFE9FFF2FBE5F7EF7980F91A" box="[1056,1170,1743,1770]" pageId="8" pageNumber="571" value="1985-07-16">
<collectingDate id="EFE3E9E2FFE9FFF2FBE5F7EF7980F91A" box="[1056,1170,1743,1770]" pageId="8" pageNumber="571" value="1985-07-16">16.7.1985</collectingDate>
</date>
</materialsCitation>
, Jäch leg. (all localities near yellow circle 
<quantity id="4CE19B2FFFE9FFF2FE50F7D37CD2F8FD" box="[405,448,1779,1805]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="571" unit="in" value="4.0">4 in</quantity>
<figureCitation id="13222A4FFFE9FFF2FE02F7D37F19F8FD" box="[455,523,1779,1805]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF52F6377906F8A6" blockId="8.[151,1437,1599,1878]" pageId="8" pageNumber="571">
- 
<emphasis id="B96DEAD8FFE9FFF2FF03F6377CB1F8C1" box="[198,419,1815,1841]" italics="true" pageId="8" pageNumber="571">
<taxonomicName id="4C194D49FFE9FFF2FF03F6377C3AF8C1" authorityName="Sharp" authorityYear="1880" box="[198,296,1815,1841]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. phoenix
</emphasis>
: 
<materialsCitation id="3B713C97FFE9FFF2FE74F637780FF8C2" box="[433,1309,1815,1842]" collectingDate="1985-08-05" collectorName="Or Aguira, S &amp; Jach" country="Israel" location="Haifa" pageId="8" pageNumber="571" specimenCount="1" stateProvince="Haifa">
<specimenCount id="9D1FFD43FFE9FFF2FE74F6377CFAF8C1" box="[433,488,1815,1841]" pageId="8" pageNumber="571" type="generic">1 ex.</specimenCount>
from 
<collectorName id="26EC531CFFE9FFF2FDEEF6377FABF8C1" box="[555,697,1815,1841]" pageId="8" pageNumber="571">Or Aguira, S</collectorName>
<collectingRegion id="49DDF828FFE9FFF2FD05F6377E13F8C1" box="[704,769,1815,1841]" country="Israel" name="Haifa" pageId="8" pageNumber="571">Haifa</collectingRegion>
, 
<collectingCountry id="F30E765AFFE9FFF2FCC9F6377E5FF8C1" box="[780,845,1815,1841]" name="Israel" pageId="8" pageNumber="571">Israel</collectingCountry>
, 
<date id="FFA7100AFFE9FFF2FC92F6377EAEF8C2" box="[855,956,1815,1842]" pageId="8" pageNumber="571" value="1985-08-05">
<collectingDate id="EFE3E9E2FFE9FFF2FC92F6377EAEF8C2" box="[855,956,1815,1842]" pageId="8" pageNumber="571" value="1985-08-05">5.8.1985</collectingDate>
</date>
, 
<collectorName id="26EC531CFFE9FFF2FC02F6377EEBF8C1" box="[967,1017,1815,1841]" pageId="8" pageNumber="571">Jäch</collectorName>
leg. (
<figureCitation id="13222A4FFFE9FFF2FBFDF637796EF8C1" box="[1080,1148,1815,1841]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
, blue circle 5)
</materialsCitation>
; 
<materialsCitation id="3B713C97FFE9FFF2FAECF6377902F8A6" collectingDate="1985-04-24" collectorName="Jach" country="Israel" location="En Tamar" municipality="Dead Sea area" pageId="8" pageNumber="571" specimenCount="1">
<specimenCount id="9D1FFD43FFE9FFF2FAECF637784DF8C1" box="[1321,1375,1815,1841]" pageId="8" pageNumber="571" type="generic">1 ex.</specimenCount>
from 
<location id="8EC66011FFE9FFF2FF03F61B7C26F8A5" LSID="urn:lsid:plazi:treatment:03B087DCFFE0FFF1FF52F53B7C58FF09:8EC66011FFE9FFF2FF03F61B7C26F8A5" box="[198,308,1851,1877]" country="Israel" municipality="Dead Sea area" name="En Tamar" pageId="8" pageNumber="571">En Tamar</location>
, 
<collectingMunicipality id="6BC2ACB0FFE9FFF2FEFAF61B7CF0F8A5" box="[319,482,1851,1877]" pageId="8" pageNumber="571">Dead Sea area</collectingMunicipality>
, 
<collectingCountry id="F30E765AFFE9FFF2FE2BF61B7F3DF8A5" box="[494,559,1851,1877]" name="Israel" pageId="8" pageNumber="571">Israel</collectingCountry>
, 
<date id="FFA7100AFFE9FFF2FDFFF61B7FBFF8A6" box="[570,685,1851,1878]" pageId="8" pageNumber="571" value="1985-04-24">
<collectingDate id="EFE3E9E2FFE9FFF2FDFFF61B7FBFF8A6" box="[570,685,1851,1878]" pageId="8" pageNumber="571" value="1985-04-24">24.4.1985</collectingDate>
</date>
, 
<collectorName id="26EC531CFFE9FFF2FD7DF61B7FF8F8A5" box="[696,746,1851,1877]" pageId="8" pageNumber="571">Jäch</collectorName>
leg. (
<figureCitation id="13222A4FFFE9FFF2FCECF61B7E7FF8A5" box="[809,877,1851,1877]" captionStart="FIGURE 7" captionStartId="9.[151,250,1956,1980]" captionTargetBox="[151,1436,525,1933]" captionTargetId="figure-203@9.[151,1436,525,1933]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Bidessus anatolicus adiyaman ssp. n. (red star), Bidessus a. anatolicus (green circles), Bidessus a. kadmos (yellow circle) and Bidessus a. phoenix (blue circles) (map adopted from Microsoft Encarta World Atlas 2000)." figureDoi="http://doi.org/10.5281/zenodo.5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="8" pageNumber="571">Fig. 7</figureCitation>
, blue circle 6)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636CAFFE9FFF2FF02F6A37E87F815" blockId="8.[151,1436,1923,2021]" pageId="8" pageNumber="571">
Differences between the new subspecies and 
<emphasis id="B96DEAD8FFE9FFF2FD02F6A47E48F86D" box="[711,858,1923,1949]" italics="true" pageId="8" pageNumber="571">B. a. kadmos</emphasis>
and 
<emphasis id="B96DEAD8FFE9FFF2FC57F6A4793AF86D" box="[914,1064,1923,1949]" italics="true" pageId="8" pageNumber="571">B. a. phoenix</emphasis>
are easily recognisable—the latter two have a distinctly less broadened habitus and are even more slender than 
<emphasis id="B96DEAD8FFE9FFF2FBD8F68879DCF831" box="[1053,1230,1959,1985]" italics="true" pageId="8" pageNumber="571">
B. a. 
<taxonomicName id="4C194D49FFE9FFF2FB9CF68779DCF831" authorityName="Wewalka" authorityYear="1972" box="[1113,1230,1959,1985]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
(see figs 2a–2c in 
<bibRefCitation id="EF884B3BFFE9FFF2FF52F6EB7C50F815" author="Wewalka, G." box="[151,322,1995,2021]" pageId="8" pageNumber="571" pagination="143 - 152" refId="ref5603" refString="Wewalka, G. (1989) Systematic and faunistic notes on Noteridae and Dytiscidae of the Near East (Coleoptera). Koleopterolo- gische Rundschau, 59, 143 - 152." type="journal article" year="1989">Wewalka 1989</bibRefCitation>
); they need not to be dealt with here in further detail.
</paragraph>
<paragraph id="8BA636CAFFE8FFF3FF02F1B77C7DFE05" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="572">
Differences between the new 
<taxonomicName id="4C194D49FFE8FFF3FDD4F1B77F78FF41" box="[529,618,151,177]" pageId="9" pageNumber="572" rank="subSpecies" subSpecies="and">ssp. and</taxonomicName>
the nominotypical 
<taxonomicName id="4C194D49FFE8FFF3FC87F1B77E84FF41" box="[834,918,151,177]" pageId="9" pageNumber="572" rank="subSpecies" subSpecies="are">ssp. are</taxonomicName>
present, but are less obvious. Both can be best separated by the body shape which is considerably broader in the new 
<taxonomicName id="4C194D49FFE8FFF3FC70F19B7902FF25" box="[949,1040,187,213]" pageId="9" pageNumber="572" rank="subSpecies" subSpecies="and">ssp. and</taxonomicName>
the sides of elytra centrally appear much more inflated (compare 
<figureCitation id="13222A4FFFE8FFF3FE35F1FF7F8EFF0A" box="[496,668,223,250]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[170,1413,182,1919]" captionTargetId="figure-17@3.[170,1417,181,1920]" captionTargetPageId="3" captionText="FIGURES 1a–1c. a, c: Habitus of Bidessus anatolicus adiyaman ssp. n. (holotype): (a) in dorsal view; (c) in ventral view (left hind-leg and last abdominal ventrites removed); b: Habitus of Bidessus anatolicus anatolicus Wewalka." figureDoi="http://doi.org/10.5281/zenodo.5453449" httpUri="https://zenodo.org/record/5453449/files/figure.png" pageId="9" pageNumber="572">Figs 1b and 1b</figureCitation>
). This is well reflected by the ratios “total length/maximum width” which differ considerably: 
<quantity id="4CE19B2FFFE8FFF3FE00F0237F40FEED" box="[453,594,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.1783" metricValueMax="4.2418" metricValueMin="4.1148" pageId="9" pageNumber="572" unit="in" value="1.645" valueMax="1.67" valueMin="1.62">1.62–1.67 in</quantity>
the new 
<taxonomicName id="4C194D49FFE8FFF3FD7DF0237E00FEED" box="[696,786,259,285]" pageId="9" pageNumber="572" rank="subSpecies" subSpecies="and">ssp. and</taxonomicName>
<quantity id="4CE19B2FFFE8FFF3FCDFF0237EB5FEED" box="[794,935,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.3434" metricValueMax="4.4196" metricValueMin="4.2672" pageId="9" pageNumber="572" unit="in" value="1.71" valueMax="1.74" valueMin="1.68">1.68–1.74 in</quantity>
the nominotypical 
<taxonomicName id="4C194D49FFE8FFF3FB45F02379CFFEED" box="[1152,1245,259,285]" pageId="9" pageNumber="573" rank="subSpecies" subSpecies="the">ssp. The</taxonomicName>
maximum width of body is in both 
<taxonomicName id="4C194D49FFE8FFF3FEAAF0077CE8FEB1" box="[367,506,295,321]" pageId="9" pageNumber="572" rank="subSpecies" subSpecies="situated">ssp. situated</taxonomicName>
before the midlength of elytra, but the outline of the body is here more strongly curved in the new ssp. In 
<emphasis id="B96DEAD8FFE8FFF3FE73F06B7F74FE95" box="[438,614,331,357]" italics="true" pageId="9" pageNumber="572">
B. a. 
<taxonomicName id="4C194D49FFE8FFF3FE34F06B7F74FE95" authorityName="Wewalka" authorityYear="1972" box="[497,614,331,357]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
the sutural lines are less impressed, the punctures on the disc (mainly between sutural line and stria) are less coarse (in 
<emphasis id="B96DEAD8FFE8FFF3FD5DF04F7E34FE79" box="[664,806,367,393]" italics="true" pageId="9" pageNumber="572">B. a. kadmos</emphasis>
and 
<emphasis id="B96DEAD8FFE8FFF3FC9FF04F7EF9FE79" box="[858,1003,367,393]" italics="true" pageId="9" pageNumber="572">B. a. phoenix</emphasis>
lines still less impressed and punctures less coarse). In 
<emphasis id="B96DEAD8FFE8FFF3FE82F0B37CEBFE5D" box="[327,505,403,429]" italics="true" pageId="9" pageNumber="572">
B. a. 
<taxonomicName id="4C194D49FFE8FFF3FE41F0B37CEBFE5D" authorityName="Wewalka" authorityYear="1972" box="[388,505,403,429]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
the sides of of the pronotum are less converging forwards and form with the sides of the elytra an almost straight line and, thus, the discontinuity of the body outline at the junction of pronotum and elytra is negligible.
</paragraph>
<caption id="DF666642FFE8FFF3FF52F6847840F810" ID-DOI="http://doi.org/10.5281/zenodo.5453461" ID-Zenodo-Dep="5453461" httpUri="https://zenodo.org/record/5453461/files/figure.png" pageId="9" pageNumber="572" startId="9.[151,250,1956,1980]" targetBox="[151,1436,525,1933]" targetPageId="9">
<paragraph id="8BA636CAFFE8FFF3FF52F6847840F810" blockId="9.[151,1437,1956,2016]" pageId="9" pageNumber="572">
<emphasis id="B96DEAD8FFE8FFF3FF52F6847C01F84C" bold="true" box="[151,275,1956,1980]" pageId="9" pageNumber="572">FIGURE 7.</emphasis>
Distribution of 
<taxonomicName id="4C194D49FFE8FFF3FE73F6847FF8F84C" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[438,746,1956,1980]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFE8FFF3FE73F6847FF8F84C" box="[438,746,1956,1980]" italics="true" pageId="9" pageNumber="572">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFE8FFF3FD35F6847E21F84C" bold="true" box="[752,819,1956,1980]" pageId="9" pageNumber="572">
<taxonomicNameLabel id="A25E57A3FFE8FFF3FD35F6847E21F84C" box="[752,819,1956,1980]" pageId="9" pageNumber="572" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(red star), 
<emphasis id="B96DEAD8FFE8FFF3FC66F6847997F84C" box="[931,1157,1956,1980]" italics="true" pageId="9" pageNumber="572">
<taxonomicName id="4C194D49FFE8FFF3FC66F6847EEEF84C" authorityName="Sharp" authorityYear="1880" box="[931,1020,1956,1980]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. 
<taxonomicName id="4C194D49FFE8FFF3FBDEF6847997F84C" authorityName="Wewalka" authorityYear="1972" box="[1051,1157,1956,1980]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
(green circles), 
<emphasis id="B96DEAD8FFE8FFF3FAEFF6847DF6F810" italics="true" pageId="9" pageNumber="572">
<taxonomicName id="4C194D49FFE8FFF3FAEFF6847891F84C" authorityName="Sharp" authorityYear="1880" box="[1322,1411,1956,1980]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. kadmos
</emphasis>
(yellow circle) and 
<emphasis id="B96DEAD8FFE8FFF3FE74F6E87F6BF810" box="[433,633,1992,2016]" italics="true" pageId="9" pageNumber="572">
<taxonomicName id="4C194D49FFE8FFF3FE74F6E87F18F810" authorityName="Sharp" authorityYear="1880" box="[433,522,1992,2016]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
a. phoenix
</emphasis>
(blue circles) (map adopted from Microsoft Encarta World Atlas 2000).
</paragraph>
</caption>
<caption id="DF666642FFEBFFF0FF52F4F37891FA1B" ID-DOI="http://doi.org/10.5281/zenodo.5453463" ID-Zenodo-Dep="5453463" box="[151,1411,1491,1518]" httpUri="https://zenodo.org/record/5453463/files/figure.png" pageId="10" pageNumber="573" startId="10.[151,264,1491,1515]" targetBox="[164,1423,194,1453]" targetPageId="10">
<paragraph id="8BA636CAFFEBFFF0FF52F4F37891FA1B" blockId="10.[151,1411,1491,1518]" box="[151,1411,1491,1518]" pageId="10" pageNumber="573">
<emphasis id="B96DEAD8FFEBFFF0FF52F4F37C47FA1B" bold="true" box="[151,341,1491,1515]" pageId="10" pageNumber="573">FIGURES 8a–8d.</emphasis>
Around the type locality of 
<taxonomicName id="4C194D49FFEBFFF0FDB1F4F37EB5FA1B" authority="Aykut &amp; Taşar &amp; Fery, 2021" authorityName="Aykut &amp; Taşar &amp; Fery" authorityYear="2021" box="[628,935,1491,1515]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="adiyaman">
<emphasis id="B96DEAD8FFEBFFF0FDB1F4F37EB5FA1B" box="[628,935,1491,1515]" italics="true" pageId="10" pageNumber="573">Bidessus anatolicus adiyaman</emphasis>
</taxonomicName>
<emphasis id="B96DEAD8FFEBFFF0FC68F4F37EE2FA1B" bold="true" box="[941,1008,1491,1515]" pageId="10" pageNumber="573">
<taxonomicNameLabel id="A25E57A3FFEBFFF0FC68F4F37EE2FA1B" box="[941,1008,1491,1515]" pageId="10" pageNumber="573" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
at the northern part of the İnekli Lake).
</paragraph>
</caption>
<paragraph id="8BA636CAFFEBFFF0FF02F7397E23F814" blockId="10.[151,1437,1561,2020]" pageId="10" pageNumber="573">
The male genitalia also show differences. Firstly, however, some notes on the figures of the median lobe in We- walka (1972 and 1989) are necessary. The figures of the ventral view of the median lobe of the new ssp. (
<figureCitation id="13222A4FFFEBFFF0FAD2F71D7874F9A8" box="[1303,1382,1597,1624]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5a</figureCitation>
) and the nominotypical ssp. (
<figureCitation id="13222A4FFFEBFFF0FE60F7417CEAF98C" box="[421,504,1633,1660]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5d</figureCitation>
; adopted from fig. 
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<bibRefCitation id="EF884B3BFFEBFFF0FCC6F7417EB9F98C" author="Wewalka, G." box="[771,939,1633,1660]" pageId="10" pageNumber="573" pagination="72 - 73" refId="ref5565" refString="Wewalka, G. (1972) Eine neue Art der Gattung Bidessus Sharp aus Kleinasien (Dytiscidae, Col.). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 23 (1971), 72 - 73." type="journal article" year="1972">Wewalka 1972</bibRefCitation>
) show considerable differences in the shape of the apex (see also the median lobe in ventral view of 
<emphasis id="B96DEAD8FFEBFFF0FCC6F7A67E84F96F" box="[771,918,1669,1695]" italics="true" pageId="10" pageNumber="573">B. a. phoenix</emphasis>
in 
<bibRefCitation id="EF884B3BFFEBFFF0FC7DF7A5798AF950" author="Wewalka, G." box="[952,1176,1669,1696]" pageId="10" pageNumber="573" pagination="143 - 152" refId="ref5603" refString="Wewalka, G. (1989) Systematic and faunistic notes on Noteridae and Dytiscidae of the Near East (Coleoptera). Koleopterolo- gische Rundschau, 59, 143 - 152." type="journal article" year="1989">Wewalka 1989: 150</bibRefCitation>
, fig. 3a). However, this discrepancy can be easily resolved: if the median lobe is oriented as in 
<figureCitation id="13222A4FFFEBFFF0FC7EF789791EF934" box="[955,1036,1705,1732]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5b</figureCitation>
and is observed from the right side (arrow) then a ventral view results which is given in 
<figureCitation id="13222A4FFFEBFFF0FD2FF7ED7E2EF918" box="[746,828,1741,1768]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5a</figureCitation>
. If, however, the median lobe is oriented as in 
<figureCitation id="13222A4FFFEBFFF0FA8FF7ED788EF918" box="[1354,1436,1741,1768]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Figs 5e</figureCitation>
or 5f and is observed from the right side (arrow) then the apex of the lobe is hidden and it results a broadly rounded apex as given in 
<figureCitation id="13222A4FFFEBFFF0FE97F6357CB6F8C0" box="[338,420,1813,1840]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5d</figureCitation>
. Notes: 
<figureCitation id="13222A4FFFEBFFF0FE3BF6357F8DF8C0" box="[510,671,1813,1840]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">
Figs 5f and 
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belong to a specimen of 
<emphasis id="B96DEAD8FFEBFFF0FC70F6367977F8DF" box="[949,1125,1813,1839]" italics="true" pageId="10" pageNumber="573">
B. a. 
<taxonomicName id="4C194D49FFEBFFF0FC35F6357977F8DF" authorityName="Wewalka" authorityYear="1972" box="[1008,1125,1813,1839]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
from Kumluca (see above). Having noted these special circumstances, we can give that in the new 
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dorsal outline of the median lobe in lateral view is almost evenly curved at the base and distinctly less bent (
<figureCitation id="13222A4FFFEBFFF0FC20F67D792BF888" box="[997,1081,1885,1912]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5b</figureCitation>
). Additionally, the lobe is more curved in distal half and the apex slightly bent dorsally. Differences in ventral view are less obvious. The paramere (
<figureCitation id="13222A4FFFEBFFF0FF65F6857DFCF830" box="[160,238,1957,1984]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">Fig. 5c</figureCitation>
) is somewhat broader in the new ssp. (for 
<emphasis id="B96DEAD8FFEBFFF0FD04F6867E7DF84F" box="[705,879,1957,1983]" italics="true" pageId="10" pageNumber="573">B. a. anotolicus</emphasis>
see 
<figureCitation id="13222A4FFFEBFFF0FC5AF6857957F830" box="[927,1093,1957,1984]" captionStart="FIGURES 5" captionStartId="7.[151,264,1886,1910]" captionTargetBox="[165,1422,181,1862]" captionTargetId="figure-17@7.[165,1422,181,1862]" captionTargetPageId="7" captionText="FIGURES 5a–5h. Male genitalia of: a–c: Bidessus anatolicus adiyaman ssp. n. (holotype): (a) median lobe in ventral view; (b) median lobe in lateral view; (c) left paramere; d–h: Bidessus a. anatolicus: (d) median lobe in ventral view; (e) median lobe in lateral view; (h) left paramere (all adopted from Wewalka 1972: 73); f, g: Bidessus a. anatolicus (specimen from Kumluca): (f) median lobe in lateral view; (h) left paramere (arrows indicate direction of view on ventral surface of median lobe)." figureDoi="http://doi.org/10.5281/zenodo.5453457" httpUri="https://zenodo.org/record/5453457/files/figure.png" pageId="10" pageNumber="573">
Figs 
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and 5h
</figureCitation>
); however, we suspect that this feature cannot be used for a safe separation of the two taxa.
</paragraph>
<paragraph id="8BA636CAFFEAFFF1FF02F1B77C58FF09" blockId="11.[151,1436,151,249]" pageId="11" pageNumber="574">
Finally, our present knowledge of the distribution of the two 
<taxonomicName id="4C194D49FFEAFFF1FCA3F1B77EE9FF41" box="[870,1019,151,177]" pageId="11" pageNumber="574" rank="subSpecies" subSpecies="indicates">ssp. indicates</taxonomicName>
that 
<emphasis id="B96DEAD8FFEAFFF1FBF5F1B779CCFF41" box="[1072,1246,151,177]" italics="true" pageId="11" pageNumber="574">
B. a. 
<taxonomicName id="4C194D49FFEAFFF1FBACF1B779CCFF41" authorityName="Wewalka" authorityYear="1972" box="[1129,1246,151,177]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="574" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
is a more coastal taxon (altitudes less than 
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) whilst the new 
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found far away from the coast and at much greater altitudes (almost 
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).
</paragraph>
</subSubSection>
</treatment>
</document>