236 lines
28 KiB
XML
236 lines
28 KiB
XML
<document id="C632586954C09DF8ACEE79B81D053B12" ID-CLB-Dataset="25209" ID-DOI="10.11646/zootaxa.4686.4.6" ID-GBIF-Dataset="bc39a3b4-60d1-4b17-9240-a757b0d6fba0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3496541" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571291216627" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2019" docId="03AA87D00469F83EFF54CA63FE907880" docLanguage="en" docName="zootaxa.4683.4.6.pdf" docOrigin="Zootaxa 4686 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus spadiceus Shavrin 2019, sp.n." docType="treatment" docVersion="8" lastPageNumber="577" masterDocId="FF93FFA8046CF838FFC3CD72FF977E63" masterDocTitle="The crassipalpis species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="580" masterPageNumber="571" pageNumber="576" updateTime="1698759364262" updateUser="ExternalLinkService">
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<mods:title id="41DB6E688199216CBC753F49276DD7A0">The crassipalpis species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="8A46219F6A78738B2C39CC450138CD53">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="1ED7EB076D1AA799C0FA53AC7A0E8893">Zootaxa</mods:title>
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<mods:part id="AC23C847148213E6627A49649DD37CC1">
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<mods:date id="8582871B5AAD0E3DBE4B935F36FDBACA">2019</mods:date>
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<mods:number id="AB376E2EA41D1FA9F7AAFD60CE67BA59">2019-10-17</mods:number>
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<mods:number id="89F47A8B4B5143F4994E96B21DEBCACF">4686</mods:number>
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<treatment id="03AA87D00469F83EFF54CA63FE907880" ID-DOI="http://doi.org/10.5281/zenodo.5931828" ID-GBIF-Taxon="159739245" ID-Zenodo-Dep="5931828" LSID="urn:lsid:plazi:treatment:03AA87D00469F83EFF54CA63FE907880" httpUri="http://treatment.plazi.org/id/03AA87D00469F83EFF54CA63FE907880" lastPageId="6" lastPageNumber="577" pageId="5" pageNumber="576">
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<subSubSection id="C319654D0469F83DFF54CA63FDFD7948" box="[151,618,1809,1836]" pageId="5" pageNumber="576" type="nomenclature">
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<paragraph id="8BBC36C60469F83DFF54CA63FDFD7948" blockId="5.[151,618,1809,1871]" box="[151,618,1809,1836]" pageId="5" pageNumber="576">
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<heading id="D0F481AA0469F83DFF54CA63FDFD7948" bold="true" box="[151,618,1809,1836]" fontSize="11" level="1" pageId="5" pageNumber="576" reason="1">
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<emphasis id="B977EAD40469F83DFF54CA63FDFD7948" bold="true" box="[151,618,1809,1836]" pageId="5" pageNumber="576">
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<taxonomicName id="4C034D450469F83DFF54CA63FDB57948" ID-CoL="8Y3ZW" authority="Shavrin" authorityName="Shavrin" authorityYear="2019" box="[151,546,1809,1836]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="spadiceus" status="sp. nov.">
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<emphasis id="B977EAD40469F83DFF54CA63FE22794F" bold="true" box="[151,437,1809,1836]" italics="true" pageId="5" pageNumber="576">Geodromicus spadiceus</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24457AF0469F83DFDECCA63FDFD7948" box="[559,618,1809,1835]" pageId="5" pageNumber="576" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C319654D0469F83DFF54CA47FEC0792C" box="[151,343,1845,1871]" pageId="5" pageNumber="576" type="description">
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<paragraph id="8BBC36C60469F83DFF54CA47FEC0792C" blockId="5.[151,618,1809,1871]" box="[151,343,1845,1871]" pageId="5" pageNumber="576">
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(
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<figureCitation id="13382A430469F83DFF5CCA47FF79792C" box="[159,238,1845,1871]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1712,1737]" captionTargetBox="[162,1428,775,1687]" captionTargetId="figure@3.[151,1436,769,1693]" captionTargetPageId="3" captionText="FIGURES 1–3. Habitus of species of Geodromicus crassipalpis group: 1–G. crassipalpis (Uttarakhand), 2–G. amplissimus (holotype), 3–G. spadiceus (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3496543" httpUri="https://zenodo.org/record/3496543/files/figure.png" pageId="5" pageNumber="576">Figs. 3</figureCitation>
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,
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<figureCitation id="13382A430469F83DFF38CA47FEB0792C" box="[251,295,1845,1871]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1944,1969]" captionTargetBox="[198,1388,174,1918]" captionTargetId="figure@7.[183,1403,166,1924]" captionTargetPageId="7" captionText="FIGURES 4–10. Aedeagus of species of Geodromicus crassipalpis group (4–5: G. crassipalpis (Uttarakhand); 6–7: G. spadiceus (paratype); 8–9: G. amplissimus (paratype; Yunnan); 10: G. sp. (Gansu)): 4, 6, 8, 10–parameral view, 5, 7, 9–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3496545" httpUri="https://zenodo.org/record/3496545/files/figure.png" pageId="5" pageNumber="576">6–7</figureCitation>
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,
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<figureCitation id="13382A430469F83DFEF0CA47FED9792C" box="[307,334,1845,1871]" captionStart="FIGURE 11" captionStartId="8.[151,250,1097,1122]" captionTargetBox="[179,1407,378,1069]" captionTargetId="figure@8.[179,1407,378,1069]" captionTargetPageId="8" captionText="FIGURE 11. Distribution of Geodromicus crassipalpis (black circles) and G. spadiceus (black square) in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.3496547" httpUri="https://zenodo.org/record/3496547/files/figure.png" pageId="5" pageNumber="576">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C319654D0469F83EFF54CA0FFB277E98" lastPageId="6" lastPageNumber="577" pageId="5" pageNumber="576" type="materials_examined">
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<paragraph id="8BBC36C60469F83DFF54CA0FFD0979BC" blockId="5.[151,1437,1916,2015]" pageId="5" pageNumber="576">
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<emphasis id="B977EAD40469F83DFF54CA0FFE5479F4" bold="true" box="[151,451,1916,1943]" pageId="5" pageNumber="576">Type material examined:</emphasis>
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<materialsCitation id="3B6B3C9B0469F83DFE08CA0FFD0579D8" ID-GBIF-Occurrence="2427264232" collectingDate="2006-06-01" collectorName="J. Schmidt'" country="Nepal" county="Sanam Khola" elevation="3900" location="Manaslu Mts." municipality="Samdo" pageId="5" pageNumber="576" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54B888640469F83DFE08CA0FFDA479F4" box="[459,563,1917,1943]" pageId="5" pageNumber="576" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D05FD4F0469F83DFDF8CA0FFDC479F4" box="[571,595,1917,1943]" pageId="5" pageNumber="576" type="male">♂</specimenCount>
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: ‘
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<collectingCountry id="F31476560469F83DFDA7CA0FFD3B79F4" box="[612,684,1917,1943]" name="Nepal" pageId="5" pageNumber="576">Nepal</collectingCountry>
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<location id="8EDC601D0469F83DFD77CA0FFCD879F4" LSID="urn:lsid:plazi:treatment:03AA87D00469F83EFF54CA63FE907880:8EDC601D0469F83DFD77CA0FFCD879F4" box="[692,847,1917,1943]" country="Nepal" county="Sanam Khola" municipality="Samdo" name="Manaslu Mts." pageId="5" pageNumber="576">Manaslu Mts.</location>
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| N-slope
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<collectingMunicipality id="6BD8ACBC0469F83DFC07CA0FFB8579F4" box="[964,1042,1917,1943]" pageId="5" pageNumber="576">Samdo</collectingMunicipality>
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<collectingCounty id="62DD4E4A0469F83DFBE4CA0FFB2879F4" box="[1063,1215,1917,1943]" pageId="5" pageNumber="576">Sanam Khola</collectingCounty>
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<quantity id="4CFB9B230469F83DFB16CA0FFAB479F4" box="[1237,1315,1917,1943]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="576" unit="m" value="3900.0">
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<elevation id="002ED1F50469F83DFB16CA0FFAB479F4" box="[1237,1315,1917,1943]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="576" unit="m" value="3900.0">3900m</elevation>
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</quantity>
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<date id="FFBD10060469F83DFAE8CA0FFA0B79F4" box="[1323,1436,1917,1943]" pageId="5" pageNumber="576" value="2006-06-01">
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<collectingDate id="EFF9E9EE0469F83DFAE8CA0FFA0B79F4" box="[1323,1436,1917,1943]" pageId="5" pageNumber="576" value="2006-06-01">1.VI.2006</collectingDate>
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</date>
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| leg.
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<collectorName id="26F653100469F83DFF1BCAD3FECF79D8" box="[216,344,1953,1979]" pageId="5" pageNumber="576">J. Schmidt’</collectorName>
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<rectangular label, printed
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</materialsCitation>
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>, ‘
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<materialsCitation id="3B6B3C9B0469F83DFD71CAD3FD0D79BC" ID-GBIF-Occurrence="2427264223" collectionCode="NME" collectorName="Shavrin A. V." country="Nepal" pageId="5" pageNumber="576" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B888640469F83DFD71CAD3FCDC79D8" box="[690,843,1953,1979]" pageId="5" pageNumber="576" type="holotype">HOLOTYPE</typeStatus>
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|
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<taxonomicName id="4C034D450469F83DFCA0CAD3FC6D79D8" authorityName="L. Redtenbacher" authorityYear="1857" box="[867,1018,1953,1979]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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| spadiceus
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<taxonomicNameLabel id="A24457AF0469F83DFB4BCAD3FB2A79D8" box="[1160,1213,1953,1979]" pageId="5" pageNumber="576" rank="species">sp.n.</taxonomicNameLabel>
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||
|
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<collectorName id="26F653100469F83DFB16CAD3FAF079D8" box="[1237,1383,1953,1979]" pageId="5" pageNumber="576">Shavrin A.V.</collectorName>
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des. 2019’ <red rectangulat label, printed> (
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<collectionCode id="ED12AE030469F83DFD8DCAB7FD0579BC" box="[590,658,1989,2015]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="5" pageNumber="576">NME</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BBC36C6046AF83EFF04CDEBFB277E98" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
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<typeStatus id="54B88864046AF83EFF04CDEBFEA37ED0" box="[199,308,153,179]" pageId="6" pageNumber="577" type="paratype">Paratypes</typeStatus>
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(
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<specimenCount id="9D05FD4F046AF83EFE86CDEBFE417ED7" box="[325,470,153,180]" pageId="6" pageNumber="577" type="generic">7 specimens</specimenCount>
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):
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<specimenCount id="9D05FD4F046AF83EFE2ACDEBFDBE7ED0" box="[489,553,153,179]" pageId="6" pageNumber="577" type="male">3 ♂♂</specimenCount>
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[two specimens dissected],
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<specimenCount id="9D05FD4F046AF83EFCA4CDEBFC307ED0" box="[871,935,153,179]" pageId="6" pageNumber="577" type="female">4 ♀♀</specimenCount>
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<materialsCitation id="3B6B3C9B046AF83EFC6CCDEBFB697EB4" ID-GBIF-Occurrence="2427264231" collectingDate="2011-09-07" collectingDateMax="2011-09-11" collectingDateMin="2011-09-07" collectionCode="NME" collectorName="Shavrin A. V." country="Nepal" pageId="6" pageNumber="577" specimenCount="1" typeStatus="paratype">
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[two specimens without right antennomeres
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<date id="FFBD1006046AF83EFF54CDCFFEAC7EB4" box="[151,315,189,215]" pageId="6" pageNumber="577" value="2011-09-11" valueMax="2011-09-07" valueMin="2011-09-11">
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<collectingDate id="EFF9E9EE046AF83EFF54CDCFFEAC7EB4" box="[151,315,189,215]" pageId="6" pageNumber="577" value="2011-09-11" valueMax="2011-09-07" valueMin="2011-09-11">9–11 and 7–11</collectingDate>
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</date>
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respectively]: same data as the holotype, with additional red rectangular printed label
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</materialsCitation>
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: ‘
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<materialsCitation id="3B6B3C9B046AF83EFACFCDCFFB3B7E98" ID-GBIF-Occurrence="2427264237" collectionCode="CS, NME" collectorName="Shavrin A. V." country="Nepal" pageId="6" pageNumber="577" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
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<typeStatus id="54B88864046AF83EFACFCDCFFA0B7EB4" box="[1292,1436,189,215]" pageId="6" pageNumber="577" type="paratype">PARATYPE</typeStatus>
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|
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<taxonomicName id="4C034D45046AF83EFF67CD93FEAC7E98" authorityName="L. Redtenbacher" authorityYear="1857" box="[164,315,225,251]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="580" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
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| spadiceus
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<taxonomicNameLabel id="A24457AF046AF83EFE01CD93FE607E98" box="[450,503,225,251]" pageId="6" pageNumber="577" rank="species">sp.n.</taxonomicNameLabel>
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||
|
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||
<collectorName id="26F65310046AF83EFDC8CD93FD0D7E98" box="[523,666,225,251]" pageId="6" pageNumber="577">Shavrin A.V.</collectorName>
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||
des. 2019’ (
|
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<specimenCount id="9D05FD4F046AF83EFCE5CD93FCC77E98" box="[806,848,225,251]" pageId="6" pageNumber="577" type="male">1 ♂</specimenCount>
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||
,
|
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<specimenCount id="9D05FD4F046AF83EFC9FCD93FC1F7E98" box="[860,904,225,251]" pageId="6" pageNumber="577" type="female">1 ♀</specimenCount>
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||
:
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<collectionCode id="ED12AE03046AF83EFC56CD93FC2D7E98" box="[917,954,225,251]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="6" pageNumber="577">CS</collectionCode>
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||
;
|
||
<specimenCount id="9D05FD4F046AF83EFC05CD93FB907E98" box="[966,1031,225,251]" pageId="6" pageNumber="577" type="male">2 ♂♂</specimenCount>
|
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,
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<specimenCount id="9D05FD4F046AF83EFBD0CD93FBC37E98" box="[1043,1108,225,251]" pageId="6" pageNumber="577" type="female">3 ♀♀</specimenCount>
|
||
:
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||
<collectionCode id="ED12AE03046AF83EFBA3CD93FB347E98" box="[1120,1187,225,251]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="6" pageNumber="577">NME</collectionCode>
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||
)
|
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</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319654D046AF83EFF04CC76FBB37868" pageId="6" pageNumber="577" type="description">
|
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<paragraph id="8BBC36C6046AF83EFF04CC76FE8F7FEF" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
<emphasis id="B977EAD4046AF83EFF04CC76FECE7F7C" bold="true" box="[199,345,260,287]" pageId="6" pageNumber="577">Description.</emphasis>
|
||
Measurements (n=8): HW: 1.13–1.26; HL: 0.76–0.84; OL: 0.27–0.28; TL: 0.08; AL (
|
||
<typeStatus id="54B88864046AF83EFAEBCC77FA067F43" box="[1320,1425,261,288]" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
|
||
): 3.87; PL: 0.94–0.97; PWmax: 1.44–1.66; PWmin: 1.32–1.47; ESL: 2.09–2.27; EW: 2.31–2.72; MTbL (
|
||
<typeStatus id="54B88864046AF83EFAEBCC5BFA067F27" box="[1320,1425,297,324]" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
|
||
): 1.65; MTrL (
|
||
<typeStatus id="54B88864046AF83EFEE8CC3FFE037F0B" box="[299,404,333,360]" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
|
||
): 0.77 (MTrL 1–4: 0.45; MTrL 5: 0.32); AW: 2.13–2.68; AedL: 1.35; BL: 5.87–7.30 (
|
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<typeStatus id="54B88864046AF83EFAA3CC3FFF5C7FE8" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
|
||
: 6.90).
|
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</paragraph>
|
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<paragraph id="8BBC36C6046AF83EFF04CCE7FCD87C00" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
Head, antennae, mouthparts, pronotum, femora and abdomen brown; elytra reddish-brown, usually with darker prescutellar area; apex of apical portion of maxillary palpi, sometimes apical antennomeres and tibia yellow-brown; tarsi yellow. Median portion of head with or without indistinct, irregular transverse meshes, denser between ocelli; infraorbital ridges with indistinct fine to distinct isodiametric microsculpture; pronotum with fine transverse microreticulation, more distinct in anterio-lateral and markedly sparser in posterio-median portions, sometimes with small impunctate area in posterio-median third. Habitus as in
|
||
<figureCitation id="13382A43046AF83EFCC5CF3BFCDE7C00" box="[774,841,585,611]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1712,1737]" captionTargetBox="[162,1428,775,1687]" captionTargetId="figure@3.[151,1436,769,1693]" captionTargetPageId="3" captionText="FIGURES 1–3. Habitus of species of Geodromicus crassipalpis group: 1–G. crassipalpis (Uttarakhand), 2–G. amplissimus (holotype), 3–G. spadiceus (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3496543" httpUri="https://zenodo.org/record/3496543/files/figure.png" pageId="6" pageNumber="577">Fig. 3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04CF1FFC707D3C" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
Head 1.4–1.5 times as wide as long, with distinctly convex infraorbital ridges; anteromedian depression deep and wide, gradually narrowed basad and extending toward very wide and moderately deep interocellar depression. Ocelli very small, sometimes indistinct; distance between ocelli markedly longer than distance between ocellus and posterior margin of eye. Punctation irregular, moderately coarse, smooth, markedly denser on infraorbital ridges. Apical maxillary palpomere 1.7 times as long as preapical segment. Antennae very long, almost reaching apical margin of elytra when reclined; lengths × widths of antennomeres (
|
||
<typeStatus id="54B88864046AF83EFC41CE53FC7C7D5F" box="[898,1003,801,828]" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
|
||
): 1: 0.37 × 0.15; 2: 0.27 × 0.10; 3: 0.37 × 0.12; 4–5: 0.32 × 0.12; 6–9: 0.35 × 0.12; 10: 0.32 × 0.12; 11: 0.50 × 0.12.
|
||
</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04CE1BFDC87A70" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">Pronotum 1.5–1.7 times as wide as long, 1.2–1.3 times as wide as head, widest slightly anterior to middle, more narrowed posteriad than anteriad, with anterior angles rounded, slightly protruded anteriad; median portion with or without moderately short and deep longitudinal impression in antero-median portion; postero-lateral portions distinctly impressed and reflexed. Punctation dense, coarse, and deep, sparser and finer along middle, sometimes with impunctate area in postero-median third.</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C96FFBDC7A38" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">Elytra slightly wider than long, slightly widened posteriad, with straight or rounded hind margins. Punctation similar to that of pronotum, sometimes sparser and somewhat finer in apical portion.</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C917FD7D7A1C" blockId="6.[151,1437,152,1763]" box="[199,746,1125,1151]" pageId="6" pageNumber="577">Metatarsus more than twice as long as metatibia.</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C9FBFCC67AA4" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">Abdomen with two transverse, large tomentose spots in the middle of tergite IV, and slightly smaller spots in the middle of tergite V. Punctation distinct, dense, rather deep.</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C9A3FBB47B34" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
Male. Posterior margin of tergite VIII rounded. Posterior margin of sternite VIII sinuate. Aedeagus (
|
||
<figureCitation id="13382A43046AF83EFAD9C9A3FACB7A8F" box="[1306,1372,1233,1260]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1944,1969]" captionTargetBox="[198,1388,174,1918]" captionTargetId="figure@7.[183,1403,166,1924]" captionTargetPageId="7" captionText="FIGURES 4–10. Aedeagus of species of Geodromicus crassipalpis group (4–5: G. crassipalpis (Uttarakhand); 6–7: G. spadiceus (paratype); 8–9: G. amplissimus (paratype; Yunnan); 10: G. sp. (Gansu)): 4, 6, 8, 10–parameral view, 5, 7, 9–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3496545" httpUri="https://zenodo.org/record/3496545/files/figure.png" pageId="6" pageNumber="577">Fig. 6</figureCitation>
|
||
) with moderately wide median lobe, in apical portion gradually narrowed toward acute apex; parameres slightly extending beyond apex of median lobe, with three moderately short apical setae, medial margins without small setae; internal sac with long flagellum, slightly widened basally. Aedeagus laterally as in
|
||
<figureCitation id="13382A43046AF83EFC18C84FFB887B34" box="[987,1055,1341,1367]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1944,1969]" captionTargetBox="[198,1388,174,1918]" captionTargetId="figure@7.[183,1403,166,1924]" captionTargetPageId="7" captionText="FIGURES 4–10. Aedeagus of species of Geodromicus crassipalpis group (4–5: G. crassipalpis (Uttarakhand); 6–7: G. spadiceus (paratype); 8–9: G. amplissimus (paratype; Yunnan); 10: G. sp. (Gansu)): 4, 6, 8, 10–parameral view, 5, 7, 9–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3496545" httpUri="https://zenodo.org/record/3496545/files/figure.png" pageId="6" pageNumber="577">Fig. 7</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C813FC2D7B18" blockId="6.[151,1437,152,1763]" box="[199,954,1377,1403]" pageId="6" pageNumber="577">Female. Posterior margins of tergite VIII and sternite VIII rounded.</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04C8F6FBB37868" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
<emphasis id="B977EAD4046AF83EFF04C8F6FE247BFC" bold="true" box="[199,435,1412,1439]" pageId="6" pageNumber="577">Comparative notes.</emphasis>
|
||
Regarding the wide median lobe of the aedeagus,
|
||
<taxonomicName id="4C034D45046AF83EFBC1C8F7FB6D7BFC" authority="Shavrin" authorityName="Shavrin" authorityYear="2019" box="[1026,1274,1413,1439]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="spadiceus" status="sp. nov.">
|
||
<emphasis id="B977EAD4046AF83EFBC1C8F7FB817BFC" box="[1026,1046,1413,1439]" italics="true" pageId="6" pageNumber="577">G</emphasis>
|
||
.
|
||
<emphasis id="B977EAD4046AF83EFBE4C8F7FB027BFC" box="[1063,1173,1413,1439]" italics="true" pageId="6" pageNumber="577">spadiceus</emphasis>
|
||
Shavrin
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B977EAD4046AF83EFAC4C8F7FAA87BFC" bold="true" box="[1287,1343,1413,1439]" pageId="6" pageNumber="577">
|
||
<taxonomicNameLabel id="A24457AF046AF83EFAC4C8F7FAA87BFC" box="[1287,1343,1413,1439]" pageId="6" pageNumber="577" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="4C034D45046AF83EFEC9C8DBFE2F7BA0" authorityName="Shavrin" authorityYear="2019" box="[266,440,1449,1475]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="amplissimus">
|
||
<emphasis id="B977EAD4046AF83EFEC9C8DBFE2F7BA0" box="[266,440,1449,1475]" italics="true" pageId="6" pageNumber="577">G. amplissimus</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs by the narrower apical portion of the aedeagus, by the absence of additional setae on the medial edges of the narrower parameres, and by a slightly wider flagellum. The new species differs from all the species of the
|
||
<taxonomicName id="4C034D45046AF83EFDD3C883FD007868" authorityName="Champion" authorityYear="1920" box="[528,663,1521,1547]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="crassipalpis">
|
||
<emphasis id="B977EAD4046AF83EFDD3C883FD007868" box="[528,663,1521,1547]" italics="true" pageId="6" pageNumber="577">crassipalpis</emphasis>
|
||
</taxonomicName>
|
||
group by the reddish-brown elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319654D046AF83EFF04CB67FE9F7830" pageId="6" pageNumber="577" type="etymology">
|
||
<paragraph id="8BBC36C6046AF83EFF04CB67FE9F7830" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
<emphasis id="B977EAD4046AF83EFF04CB67FED9784C" bold="true" box="[199,334,1557,1583]" pageId="6" pageNumber="577">Etymology.</emphasis>
|
||
The specific epithet is the Latin adjective
|
||
<taxonomicName id="4C034D45046AF83EFCEFCB67FC0D784C" authorityName="Shavrin" authorityYear="2019" box="[812,922,1557,1583]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="spadiceus">
|
||
<emphasis id="B977EAD4046AF83EFCEFCB67FC0D784C" box="[812,922,1557,1583]" italics="true" pageId="6" pageNumber="577">spadiceus</emphasis>
|
||
</taxonomicName>
|
||
(reddish-brown). It refers to the coloration of the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319654D046AF83EFF04CB2EFE907880" pageId="6" pageNumber="577" type="distribution">
|
||
<paragraph id="8BBC36C6046AF83EFF04CB2EFF6578F8" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
<emphasis id="B977EAD4046AF83EFF04CB2EFEF67814" bold="true" box="[199,353,1628,1655]" pageId="6" pageNumber="577">Distribution.</emphasis>
|
||
The species is at present known only from the
|
||
<typeStatus id="54B88864046AF83EFCB0CB2FFC347814" box="[883,931,1629,1655]" pageId="6" pageNumber="577">type</typeStatus>
|
||
locality in the Manaslu range, Central
|
||
<collectingCountry id="F3147656046AF83EFA9ACB2FFA0B7814" box="[1369,1436,1629,1655]" name="Nepal" pageId="6" pageNumber="577">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="13382A43046AF83EFF63CBF3FF7378F8" box="[160,228,1665,1691]" captionStart="FIGURES 4–10" captionStartId="7.[151,264,1944,1969]" captionTargetBox="[198,1388,174,1918]" captionTargetId="figure@7.[183,1403,166,1924]" captionTargetPageId="7" captionText="FIGURES 4–10. Aedeagus of species of Geodromicus crassipalpis group (4–5: G. crassipalpis (Uttarakhand); 6–7: G. spadiceus (paratype); 8–9: G. amplissimus (paratype; Yunnan); 10: G. sp. (Gansu)): 4, 6, 8, 10–parameral view, 5, 7, 9–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3496545" httpUri="https://zenodo.org/record/3496545/files/figure.png" pageId="6" pageNumber="577">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC36C6046AF83EFF04CBD7FE907880" blockId="6.[151,1437,152,1763]" pageId="6" pageNumber="577">
|
||
<emphasis id="B977EAD4046AF83EFF04CBD7FE1A78DC" bold="true" box="[199,397,1700,1727]" pageId="6" pageNumber="577">Natural History.</emphasis>
|
||
The specimens were collected at an elevation of
|
||
<quantity id="4CFB9B23046AF83EFC00CBD7FB8D78DC" box="[963,1050,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="577" unit="m" value="3900.0">3900 m</quantity>
|
||
a.s.l. Detailed ecological data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |