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<document id="7C2E3E95DE6E16016D323B4DBC12EFB4" ID-CLB-Dataset="21683" ID-DOI="10.11646/zootaxa.4798.1.1" ID-GBIF-Dataset="d8c0dac0-6b03-43f4-be6d-1dc3d3d9cfa3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3896551" ID-ZooBank="4830ED2F-DB55-468E-97D4-2A278CBA6CE0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592308062015" checkinUser="plazi" docAuthor="Antić, Dragan &amp; Akkari, Nesrine" docDate="2020" docId="03A90C3B1515FFBAFF247571FB32F970" docLanguage="en" docName="zootaxa.4798.1.1.pdf" docOrigin="Zootaxa 4798 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Haasea vidinensis" docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="FF907443153BFF88FFB3730DFFB5FF93" masterDocTitle="Haasea Verhoeff, 1895 - a genus of tumultuous history and chaotic recordsredefinition, revision of taxonomy and geographic distributions, with descriptions of two new species from Austria and Serbia (Diplopoda, Chordeumatida, Haaseidae)" masterLastPageNumber="77" masterPageNumber="1" pageNumber="47" updateTime="1698842500395" updateUser="ExternalLinkService">
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<mods:title id="56008FFD1FBB6860A2397357B7A1681F">Haasea Verhoeff, 1895 - a genus of tumultuous history and chaotic recordsredefinition, revision of taxonomy and geographic distributions, with descriptions of two new species from Austria and Serbia (Diplopoda, Chordeumatida, Haaseidae)</mods:title>
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<mods:namePart id="1F93252E87EAC2D8DED86A9AE4327B9F">Antić, Dragan</mods:namePart>
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<mods:namePart id="38631AE90F8D74B0EF1C7004DC53155C">Akkari, Nesrine</mods:namePart>
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<treatment id="03A90C3B1515FFBAFF247571FB32F970" ID-GBIF-Taxon="164612307" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970" httpUri="http://treatment.plazi.org/id/03A90C3B1515FFBAFF247571FB32F970" lastPageId="50" lastPageNumber="51" pageId="46" pageNumber="47">
<subSubSection id="C31AEEA61515FFA6FF247571FDFCF906" box="[151,585,1659,1686]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BBFBD2D1515FFA6FF247571FDFCF906" blockId="46.[151,585,1659,1722]" box="[151,585,1659,1686]" pageId="46" pageNumber="47">
<heading id="D0F70A411515FFA6FF247571FDFCF906" bold="true" box="[151,585,1659,1686]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<taxonomicName id="4C00C6AE1515FFA6FF247571FDFCF906" ID-CoL="3J3JM" authority="(Strasser, 1973 a)" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[151,585,1659,1686]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1515FFA6FF247571FDFCF906" bold="true" box="[151,585,1659,1686]" pageId="46" pageNumber="47">
<emphasis id="B974613F1515FFA6FF247571FEC6F905" bold="true" box="[151,371,1659,1686]" italics="true" pageId="46" pageNumber="47">Haasea vidinensis</emphasis>
(
<bibRefCitation id="EF91C0DC1515FFA6FE317576FDF4F906" author="Strasser, K." box="[386,577,1659,1685]" pageId="46" pageNumber="47" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser, 1973a</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61515FFA6FF2475ADFE27F929" box="[151,402,1695,1722]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BBFBD2D1515FFA6FF2475ADFE27F929" blockId="46.[151,585,1659,1722]" box="[151,402,1695,1722]" pageId="46" pageNumber="47">
<figureCitation id="133BA1A81515FFA6FF2475ADFF59F929" box="[151,236,1696,1722]" captionStart="FIGURE 1" captionStartId="7.[151,250,1858,1883]" captionTargetBox="[212,1375,181,1833]" captionTargetId="figure@7.[212,1375,181,1833]" captionTargetPageId="7" captionText="FIGURE 1. Living Haasea Verhoeff, 1895. A. H. germanica (Verhoeff, 1901), female from Reiflding, Germany. B. H. ger- manica (Verhoeff, 1901), mating pair from Riedl, Germany. C. H. flavescens (Latzel, 1884), male from Schwaighofen, Austria. D. H. flavescens (Latzel, 1884), mating pair from Gaisberg, Austria. E. H. hungarica (Verhoeff, 1928), male from Fruška gora Mt., Serbia. F. H. vidinensis (Strasser, 1973a), female from the Ponor Radovanske reke Cave, Serbia. Photos credit: AD. Jörg Spelda, E. Dragiša Savić (photo in lab), F. Nikola Vesović (photo in lab)." figureDoi="http://doi.org/10.5281/zenodo.3896553" httpUri="https://zenodo.org/record/3896553/files/figure.png" pageId="46" pageNumber="47">Figs 1F</figureCitation>
,
<figureCitation id="133BA1A81515FFA6FF4B7592FEA3F929" box="[248,278,1695,1722]" captionStart="FIGURE 25" captionStartId="48.[151,250,1762,1787]" captionTargetBox="[264,1324,181,1737]" captionTargetId="figure@48.[264,1324,181,1737]" captionTargetPageId="48" captionText="FIGURE 25. Haasea vidinensis (Strasser, 1973a). A. Allotype female of H. guidononveilleri Makarov, 2008 syn. nov. (IZB), habitus, lateral view. B. Male from Kučaj Mts, Serbia, right anterior gonopod, mesal view. C. Male from Kučaj Mts, Serbia, right anterior gonopod, lateral view. D. Male from Kalafat Mt, Serbia, left anterior gonopod, distomesal view. E. Male from Kalafat Mt, Serbia, right anterior gonopod, detail of distal part, posterodistal view. F. Male from Kalafat Mt, Serbia, right anterior gonopod, detail with lateral processes, lateral view. Abbreviations: a—angiocoxite; alp—anterior lateral process; dp—distal part; f—furrow; mdl—mesodistal lobe; ml—“Opuntia-like” membranous lobe; mp—mesal process; msp—medial sternal process; pc—papillated cone; pl—posterior lamella; plp—posterior lateral process; so—seminal opening. Scale bars: 1 mm (A), 0.1 mm (BE), 0.01 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3896601" httpUri="https://zenodo.org/record/3896601/files/figure.png" pageId="46" pageNumber="47">25</figureCitation>
,
<figureCitation id="133BA1A81515FFA6FE917592FEF5F92A" box="[290,320,1695,1721]" captionStart="FIGURE 26" captionStartId="49.[151,250,1698,1723]" captionTargetBox="[151,1435,181,1673]" captionTargetId="figure@49.[151,1435,181,1673]" captionTargetPageId="49" captionText="FIGURE 26. Haasea vidinensis (Strasser, 1973a). A. Lectotype male (NMNHS10807), right anterior gonopod, lateral view. B. Paralectotype male (NMNHS10808), left anterior gonopod, lateral view. C. H. guidononveilleri Makarov, 2008 syn. nov., holotype male (IZB), left anterior gonopod, lateral view. D. Male from Rtanj Mt, Serbia, right anterior gonopod, lateral view. E. Male from Kučaj Mts, Serbia, left anterior gonopod, lateral view. F. Male from Kučaj Mts, Serbia, left anterior gonopod, mesal view. G. Lectotype male (NMNHS10807), right posterior gonopod, anteromesal view. H. Male from Kučaj Mts, Serbia, right posterior gonopod, mesal view. I. H. guidononveilleri Makarov, 2008 syn. nov., holotype male (IZB), posterior gonopods, posterior view. J. Male from Rtanj, Mt, Serbia, posterior gonopods, posteror view. Abbreviations: ab—anterior branch; cp—coxal process; pb—posterior branch; plp—posterior lateral process; s—sternum t—telopodite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3896603" httpUri="https://zenodo.org/record/3896603/files/figure.png" pageId="46" pageNumber="47">26</figureCitation>
,
<figureCitation id="133BA1A81515FFA6FEFF75ADFEDFF929" box="[332,362,1696,1722]" captionStart="FIGURE 38" captionStartId="67.[151,250,1157,1182]" captionTargetBox="[151,1435,417,1133]" captionTargetId="figure@67.[151,1435,417,1133]" captionTargetPageId="67" captionText="FIGURE 38. Distribution of the genus Haasea." figureDoi="http://doi.org/10.5281/zenodo.3896629" httpUri="https://zenodo.org/record/3896629/files/figure.png" pageId="46" pageNumber="47">38</figureCitation>
,
<figureCitation id="133BA1A81515FFA6FEC575ADFE27F929" box="[374,402,1696,1722]" captionStart="FIGURE 42" captionStartId="70.[151,250,1979,2004]" captionTargetBox="[188,1399,1135,1953]" captionTargetId="figure@70.[188,1399,1135,1953]" captionTargetPageId="70" captionText="FIGURE 42. Distribution of H. intermedia Mršić, 1985, H. lacusnigri (Gulička, 1968), H. makarovi sp. nov., H. microcorna (Strasser, 1971), H. plasana (Verhoeff, 1899) and H. vidinensis (Strasser, 1973a)." figureDoi="http://doi.org/10.5281/zenodo.3896637" httpUri="https://zenodo.org/record/3896637/files/figure.png" pageId="46" pageNumber="47">42</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61515FFA6FF2475E9FD20F888" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8BBFBD2D1515FFA6FF2475E9FD74F96F" blockId="46.[151,705,1764,1819]" box="[151,705,1764,1788]" pageId="46" pageNumber="47">
<taxonomicName id="4C00C6AE1515FFA6FF2475E9FD74F96F" ID-CoL="6T7RT" authority="Strasser, 1973 a" authorityName="Strasser" authorityYear="1973" box="[151,705,1764,1788]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1515FFA6FF2475E9FD94F96F" box="[151,545,1764,1788]" italics="true" pageId="46" pageNumber="47">Orobainosoma (Histrosoma) vidinense</emphasis>
<bibRefCitation id="EF91C0DC1515FFA6FD9475E9FD74F96F" author="Strasser, K." box="[551,705,1764,1788]" pageId="46" pageNumber="47" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser, 1973a</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BBFBD2D1515FFA6FF24740EFD20F888" blockId="46.[151,705,1764,1819]" box="[151,661,1794,1819]" pageId="46" pageNumber="47">
<taxonomicName id="4C00C6AE1515FFA6FF24740EFD87F888" ID-CoL="84938" authority="Makarov, 2008" authorityName=", Makarov" authorityYear="2008" box="[151,562,1794,1819]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1515FFA6FF24740EFE27F888" box="[151,402,1795,1819]" italics="true" pageId="46" pageNumber="47">Haasea guidononveilleri</emphasis>
<bibRefCitation id="EF91C0DC1515FFA6FE2A740FFD87F888" author="Makarov, S. E." box="[409,562,1794,1819]" pageId="46" pageNumber="47" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov, 2008</bibRefCitation>
</taxonomicName>
<emphasis id="B974613F1515FFA6FD8B740EFD20F888" bold="true" box="[568,661,1795,1819]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A247DC441515FFA6FD8B740EFD20F888" box="[568,661,1795,1819]" pageId="46" pageNumber="47">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61515FFA6FF247445FE77F838" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BBFBD2D1515FFA6FF247445FE77F838" blockId="46.[151,1437,1864,2035]" pageId="46" pageNumber="47">
<emphasis id="B974613F1515FFA6FF247445FEA7F8F0" bold="true" box="[151,274,1864,1891]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
Differs from the other congeners by the presence of a strongly developed, knife-like mesal process, parallel and closely fitting to angiocoxite with basal tooth as well as by the presence of rudimentary anterior and posterior lateral processes.
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61515FFBAFF7474B8FDB5FE3C" lastPageId="50" lastPageNumber="51" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BBFBD2D1515FFA7FF7474B8FEDDFDF0" blockId="46.[151,1437,1864,2035]" lastBlockId="47.[151,1437,153,611]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B974613F1515FFA6FF7474B8FE44F85C" bold="true" box="[199,497,1972,1999]" pageId="46" pageNumber="47">Type material examined:</emphasis>
NMNHS:
<typeStatus id="54BB038F1515FFA6FDDF74B8FD69F85C" box="[620,732,1973,1999]" pageId="46" pageNumber="47" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D0676A41515FFA6FD5074B8FD4DF85C" box="[739,760,1973,1999]" pageId="46" pageNumber="47" type="male"></specimenCount>
(
<collectionCode id="ED1125E81515FFA6FCB574B8FCD8F85C" box="[774,877,1973,1999]" pageId="46" pageNumber="47">NMNHS</collectionCode>
10 807), here designated,
<emphasis id="B974613F1515FFA6FB2674B8FF6EF860" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4C00C6AE1515FFA6FB2674B8FA89F85C" authorityName="Verhoeff" authorityYear="1897" box="[1173,1340,1973,1999]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Orobainosoma</taxonomicName>
(Histro- soma)
</emphasis>
<taxonomicName id="4C00C6AE1515FFA6FF5174D4FE47F860" authority="Strasser, 1971" authorityName="Strasser" authorityYear="1971" box="[226,498,2009,2035]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="vidinense">
<emphasis id="B974613F1515FFA6FF5174D4FEF9F860" box="[226,332,2009,2035]" italics="true" pageId="46" pageNumber="47">vidinense</emphasis>
<bibRefCitation id="EF91C0DC1515FFA6FEE074D4FE47F860" author="Strasser, K." box="[339,498,2009,2035]" pageId="46" pageNumber="47" pagination="313 - 345" refId="ref46776" refString="Strasser, K. (1971) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 52 (3 - 5), 313 - 345." type="journal article" year="1971">Strasser, 1971</bibRefCitation>
</taxonomicName>
, alcohol material: body in three pieces:
<typeStatus id="54BB038F1515FFA6FC1D74D4FBAAF860" box="[942,1055,2009,2035]" pageId="46" pageNumber="47" type="holotype">holotypus</typeStatus>
, Bulg., Distr. Vidin, v. Bela, gouf- fre Džamiite,
<date id="FFBE9BED1514FFA7FE817394FE1EFF20" box="[306,427,153,179]" pageId="47" pageNumber="48" value="1969-10-19">19.X.1969</date>
, P. Beron leg., one microslide (
<collectionCode id="ED1125E81514FFA7FCBA7394FCC5FF20" box="[777,880,153,179]" pageId="47" pageNumber="48">NMNHS</collectionCode>
10 807):
<typeStatus id="54BB038F1514FFA7FC6E7394FBFBFF20" box="[989,1102,153,179]" pageId="47" pageNumber="48" type="holotype">holotypus</typeStatus>
,
<taxonomicName id="4C00C6AE1514FFA7FBE97394FE86FF44" authority="Str." authorityName="Strasser" authorityYear="1973" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1514FFA7FBE97394FEB4FF44" italics="true" pageId="47" pageNumber="48">Orobainosoma (Histrosoma) vidinense</emphasis>
Str.
</taxonomicName>
, 19.10.69. g. Džamiite, Bela, Vidin, NW Bulg. (1386), Gon. v. h., 8. 9. Bp. (anterior and posterior gonopods, leg-pairs 10 and 11).
<typeStatus id="54BB038F1514FFA7FDBB73ECFD1EFF68" box="[520,683,225,251]" pageId="47" pageNumber="48" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="ED1125E81514FFA7FD0F73ECFC96FF68" box="[700,803,225,251]" pageId="47" pageNumber="48">NMNHS</collectionCode>
10 808):
<specimenCount id="9D0676A41514FFA7FC2773ECFC61FF68" box="[916,980,225,251]" pageId="47" pageNumber="48" type="male">2 ♂♂</specimenCount>
(one without gonopods),
<specimenCount id="9D0676A41514FFA7FB4F73ECFA88FF68" box="[1276,1341,225,251]" pageId="47" pageNumber="48" type="female">4 ♀♀</specimenCount>
, 2 juve- niles. MHNG:
<typeStatus id="54BB038F1514FFA7FE8C7208FE57FE8C" box="[319,482,261,287]" pageId="47" pageNumber="48" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="ED1125E81514FFA7FE417208FDFAFE8C" box="[498,591,261,287]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="47" pageNumber="48" type="Museum">MHNG</collectionCode>
):
<specimenCount id="9D0676A41514FFA7FDD27208FD3BFE8C" box="[609,654,261,287]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0676A41514FFA7FD297208FD70FE8C" box="[666,709,261,287]" pageId="47" pageNumber="48" type="female">1 ♀</specimenCount>
, alcohol material,
<taxonomicName id="4C00C6AE1514FFA7FC247208FAA3FE8C" authority="Str." authorityName="Strasser" authorityYear="1973" box="[919,1302,261,287]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1514FFA7FC247208FB53FE8C" box="[919,1254,261,287]" italics="true" pageId="47" pageNumber="48">Orob. (Histrosoma) vidinense</emphasis>
Str.
</taxonomicName>
, NW Bulg.;
<specimenCount id="9D0676A41514FFA7FF247224FF72FED0" box="[151,199,297,323]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
, body in two parts, mounted on cardboard; two microslides: 1)
<taxonomicName id="4C00C6AE1514FFA7FC017224FA28FED0" authority="Str." authorityName="Strasser" authorityYear="1973" box="[946,1437,297,323]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1514FFA7FC017224FAD9FED0" box="[946,1388,297,323]" italics="true" pageId="47" pageNumber="48">Orobainosoma (Histrosoma) vidinense</emphasis>
Str.
</taxonomicName>
<date id="FFBE9BED1514FFA7FF247240FEACFEF4" box="[151,281,333,359]" pageId="47" pageNumber="48" value="1969-10-19">19.10.1969</date>
. Gr. Džamiite. Bela (Vidin). NW Bulg., anterior and posterior gonopods, pleurotergite 7; 2)
<taxonomicName id="4C00C6AE1514FFA7FA947240FDBEFE18" authority="Str." authorityName="Strasser" authorityYear="1973" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1514FFA7FA947240FE6BFE18" italics="true" pageId="47" pageNumber="48">Orobainosoma (Histrosoma) vidinense</emphasis>
Str.
</taxonomicName>
<date id="FFBE9BED1514FFA7FDA3727CFD27FE18" box="[528,658,369,395]" pageId="47" pageNumber="48" value="1969-10-19">19.10.1969</date>
. Gr. Džamiite. Bela (Vidin). NW Bulg., leg-pairs 17, 10 and 11;
<specimenCount id="9D0676A41514FFA7FADF727CFA22FE18" box="[1388,1431,369,395]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
, microslide:
<taxonomicName id="4C00C6AE1514FFA7FEAF7298FD49FE3C" authority="Str." authorityName="Strasser" authorityYear="1973" box="[284,764,405,431]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vidinense" subGenus="Histrosoma">
<emphasis id="B974613F1514FFA7FEAF7298FD7AFE3C" box="[284,719,405,431]" italics="true" pageId="47" pageNumber="48">Orobainosoma (Histrosoma) vidinense</emphasis>
Str.
</taxonomicName>
<date id="FFBE9BED1514FFA7FCB17298FC31FE3C" box="[770,900,405,431]" pageId="47" pageNumber="48" value="1969-10-19">19.10.1969</date>
. Gr. Džamiite. Bela (Vidin). NW Bulg., anterior and posterior gonopods, leg-pairs 10 and 11. Other examined types: IZB: One vial labelled as
<emphasis id="B974613F1514FFA7FB7872B7FF6EFE64" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="4C00C6AE1514FFA7FB7872B7FA94FE40" authorityName="Verhoeff" authorityYear="1895" box="[1227,1313,442,467]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">Haasea</taxonomicName>
guidonon- veielli
</emphasis>
(sic!) Makarov, 2005,
<typeStatus id="54BB038F1514FFA7FE6E72D0FD8AFE64" box="[477,575,477,503]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
<specimenCount id="9D0676A41514FFA7FDF572D0FDEEFE64" box="[582,603,477,503]" pageId="47" pageNumber="48" type="male"></specimenCount>
(body in three pieces),
<specimenCount id="9D0676A41514FFA7FCD672D0FC45FE64" box="[869,1008,477,503]" pageId="47" pageNumber="48" type="female" typeStatus="allotype">1 allotype ♀</specimenCount>
(habitus picture),
<specimenCount id="9D0676A41514FFA7FB0D72D0FA2DFE64" box="[1214,1432,477,503]" pageId="47" pageNumber="48" type="juvenile" typeStatus="paratype">1 paratype juvenile</specimenCount>
,
<collectingCountry id="F317FDBD1514FFA7FF24710CFF68FD88" box="[151,221,513,539]" name="Serbia" pageId="47" pageNumber="48">Srbija</collectingCountry>
,
<collectingRegion id="49C473CF1514FFA7FF5B710CFE8BFD88" box="[232,318,513,539]" country="Serbia" name="Zajecar" pageId="47" pageNumber="48">Zaječar</collectingRegion>
, pl. Tupižnica, jama Ledenica,
<quantity id="4CF810C81514FFA7FD2B710CFD5BFD88" box="[664,750,513,539]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FFBE9BED1514FFA7FD49710DFCCBFD88" box="[762,894,512,539]" pageId="47" pageNumber="48" value="2004-05-14">14.05.2004</date>
., leg. S. Ognjenović (
<collectionCode id="ED1125E81514FFA7FBD9710CFB2EFD88" box="[1130,1179,513,539]" pageId="47" pageNumber="48">IZB</collectionCode>
), one microslide labels as
<taxonomicName id="4C00C6AE1514FFA7FF06712BFDF6FDAC" authority="Makarov, 2005" authorityName="Makarov" authorityYear="2005" box="[181,579,548,575]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1514FFA7FF06712BFE25FDAC" box="[181,400,549,575]" italics="true" pageId="47" pageNumber="48">H. guidononveilleri</emphasis>
Makarov, 2005
</taxonomicName>
,
<typeStatus id="54BB038F1514FFA7FDFE7128FD1AFDAC" box="[589,687,549,575]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
<specimenCount id="9D0676A41514FFA7FD057128FD7BFDAC" box="[694,718,549,575]" pageId="47" pageNumber="48" type="male"></specimenCount>
: gonopods, X legs,
<collectingRegion id="49C473CF1514FFA7FC1A7128FC4AFDAC" box="[937,1023,549,575]" country="Serbia" name="Zajecar" pageId="47" pageNumber="48">Zaječar</collectingRegion>
, Tupižnica, Ledenica,
<date id="FFBE9BED1514FFA7FB497129FACEFDAC" box="[1274,1403,548,575]" pageId="47" pageNumber="48" value="2004-05-14">14.05.2004</date>
, S. Ognjenović (
<collectionCode id="ED1125E81514FFA7FE9A7144FEEEFDF0" box="[297,347,585,611]" pageId="47" pageNumber="48">IZB</collectionCode>
).
</paragraph>
<caption id="DF7FEDA51514FFA7FF247402FC2DF8E3" ID-DOI="http://doi.org/10.5281/zenodo.3896599" ID-Zenodo-Dep="3896599" httpUri="https://zenodo.org/record/3896599/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1807,1832]" targetBox="[151,1435,633,1781]" targetPageId="47">
<paragraph id="8BBFBD2D1514FFA7FF247402FC2DF8E3" blockId="47.[151,1437,1807,1904]" pageId="47" pageNumber="48">
<emphasis id="B974613F1514FFA7FF247402FE95F8BB" bold="true" box="[151,288,1807,1832]" pageId="47" pageNumber="48">FIGURE 24.</emphasis>
<taxonomicName id="4C00C6AE1514FFA7FE95741DFDC6F8BB" authority="(Strasser, 1966)" baseAuthorityName="Strasser" baseAuthorityYear="1966" box="[294,627,1807,1832]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pretneri">
<emphasis id="B974613F1514FFA7FE95741DFE7EF8B4" box="[294,459,1808,1831]" italics="true" pageId="47" pageNumber="48">Haasea pretneri</emphasis>
(
<bibRefCitation id="EF91C0DC1514FFA7FE6B7402FDD9F8BB" author="Strasser, K." box="[472,620,1807,1832]" pageId="47" pageNumber="48" pagination="159 - 220" refId="ref46749" refString="Strasser, K. (1966) Die Diplopoden Sloweniens-Kacice (Diplopoda) Slovenije. Porocila (Acta carsologica), 4, 159 - 220." type="journal article" year="1966">Strasser, 1966</bibRefCitation>
)
</taxonomicName>
, lectotype male (MHNG).
<emphasis id="B974613F1514FFA7FC307402FC0EF8BB" bold="true" box="[899,955,1807,1832]" pageId="47" pageNumber="48">AC.</emphasis>
Habitus.
<emphasis id="B974613F1514FFA7FBAE7402FB83F8B4" bold="true" box="[1053,1078,1807,1831]" pageId="47" pageNumber="48">D.</emphasis>
Microslide with dissected parts.
<emphasis id="B974613F1514FFA7FA367402FA29F8B4" bold="true" box="[1413,1436,1807,1831]" pageId="47" pageNumber="48">E.</emphasis>
Right anterior gonopod, lateral view.
<emphasis id="B974613F1514FFA7FDA6743EFD9DF8D8" bold="true" box="[533,552,1843,1867]" pageId="47" pageNumber="48">F.</emphasis>
Left anterior gonopod, mesal view.
<emphasis id="B974613F1514FFA7FC28743EFC00F8DF" bold="true" box="[923,949,1843,1868]" pageId="47" pageNumber="48">G.</emphasis>
Telopodite of the posterior gonopod.
<emphasis id="B974613F1514FFA7FA8B743EFAE7F8D8" bold="true" box="[1336,1362,1843,1867]" pageId="47" pageNumber="48">H.</emphasis>
Coxite of the posterior gonopod. Without scale. Photos credit: Peter Schwendinger.
</paragraph>
</caption>
<caption id="DF7FEDA5150BFFB8FF2475EFFE0BF864" ID-DOI="http://doi.org/10.5281/zenodo.3896601" ID-Zenodo-Dep="3896601" httpUri="https://zenodo.org/record/3896601/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1762,1787]" targetBox="[264,1324,181,1737]" targetPageId="48">
<paragraph id="8BBFBD2D150BFFB8FF2475EFFE0BF864" blockId="48.[151,1436,1762,2039]" pageId="48" pageNumber="49">
<emphasis id="B974613F150BFFB8FF2475EFFEAFF968" bold="true" box="[151,282,1762,1787]" pageId="48" pageNumber="49">FIGURE 25</emphasis>
.
<taxonomicName id="4C00C6AE150BFFB8FE9B75E9FD22F968" authority="(Strasser, 1973 a)" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[296,663,1763,1787]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F150BFFB8FE9B75E9FE54F968" box="[296,481,1763,1787]" italics="true" pageId="48" pageNumber="49">Haasea vidinensis</emphasis>
(
<bibRefCitation id="EF91C0DC150BFFB8FE5C75EEFD3AF968" author="Strasser, K." box="[495,655,1763,1787]" pageId="48" pageNumber="49" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser, 1973a</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B974613F150BFFB8FD1275EFFD0FF968" bold="true" box="[673,698,1762,1787]" pageId="48" pageNumber="49">A.</emphasis>
Allotype female of
<taxonomicName id="4C00C6AE150BFFB8FC3475E9FB45F968" authority="Makarov, 2008" authorityName=", Makarov" authorityYear="2008" box="[903,1264,1762,1787]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F150BFFB8FC3475E9FBFBF968" box="[903,1102,1763,1787]" italics="true" pageId="48" pageNumber="49">H. guidononveilleri</emphasis>
<bibRefCitation id="EF91C0DC150BFFB8FBE575EFFB45F968" author="Makarov, S. E." box="[1110,1264,1762,1787]" pageId="48" pageNumber="49" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov, 2008</bibRefCitation>
</taxonomicName>
<emphasis id="B974613F150BFFB8FB4475EEFAE0F968" bold="true" box="[1271,1365,1763,1787]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A247DC44150BFFB8FB4475EEFAE0F968" box="[1271,1365,1763,1787]" pageId="48" pageNumber="49">syn. nov.</taxonomicNameLabel>
</emphasis>
(IZB), habitus, lateral view.
<emphasis id="B974613F150BFFB8FEC0740AFE3EF88C" bold="true" box="[371,395,1799,1823]" pageId="48" pageNumber="49">B.</emphasis>
Male from Kučaj Mts, Serbia, right anterior gonopod, mesal view.
<emphasis id="B974613F150BFFB8FBF5740BFBEBF88C" bold="true" box="[1094,1118,1798,1823]" pageId="48" pageNumber="49">C.</emphasis>
Male from Kučaj Mts, Serbia, right anterior gonopod, lateral view.
<emphasis id="B974613F150BFFB8FDB57426FDAAF8D0" bold="true" box="[518,543,1835,1859]" pageId="48" pageNumber="49">D.</emphasis>
Male from Kalafat Mt, Serbia, left anterior gonopod, distomesal view.
<emphasis id="B974613F150BFFB8FB5A7426FAB5F8D0" bold="true" box="[1257,1280,1835,1859]" pageId="48" pageNumber="49">E.</emphasis>
Male from Kalafat Mt, Serbia, right anterior gonopod, detail of distal part, posterodistal view.
<emphasis id="B974613F150BFFB8FC707442FC63F8F4" bold="true" box="[963,982,1871,1895]" pageId="48" pageNumber="49">F.</emphasis>
Male from Kalafat Mt, Serbia, right anterior gonopod, detail with lateral processes, lateral view.
<emphasis id="B974613F150BFFB8FD18747FFCD5F818" bold="true" box="[683,864,1906,1931]" pageId="48" pageNumber="49">Abbreviations: a</emphasis>
—angiocoxite;
<emphasis id="B974613F150BFFB8FC4D747EFB95F818" bold="true" box="[1022,1056,1907,1931]" pageId="48" pageNumber="49">alp</emphasis>
—anterior lateral process;
<emphasis id="B974613F150BFFB8FA83747EFAF9F818" bold="true" box="[1328,1356,1907,1931]" pageId="48" pageNumber="49">dp</emphasis>
—distal part;
<emphasis id="B974613F150BFFB8FF78749BFF66F83C" bold="true" box="[203,211,1942,1967]" pageId="48" pageNumber="49">f</emphasis>
—furrow;
<emphasis id="B974613F150BFFB8FE8C749AFEDCF83C" bold="true" box="[319,361,1943,1967]" pageId="48" pageNumber="49">mdl</emphasis>
—mesodistal lobe;
<emphasis id="B974613F150BFFB8FD9E749AFDFCF83C" bold="true" box="[557,585,1943,1967]" pageId="48" pageNumber="49">ml</emphasis>
—“
<taxonomicName id="4C00C6AE150BFFB8FDDD749AFD74F83D" box="[622,705,1943,1966]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="genus">
<emphasis id="B974613F150BFFB8FDDD749AFD74F83D" box="[622,705,1943,1966]" italics="true" pageId="48" pageNumber="49">Opuntia</emphasis>
</taxonomicName>
-like” membranous lobe;
<emphasis id="B974613F150BFFB8FC72749AFC51F83C" bold="true" box="[961,996,1943,1967]" pageId="48" pageNumber="49">mp</emphasis>
—mesal process;
<emphasis id="B974613F150BFFB8FB2B749AFB70F83C" bold="true" box="[1176,1221,1943,1967]" pageId="48" pageNumber="49">msp</emphasis>
—medial sternal process;
<emphasis id="B974613F150BFFB8FF7C74B6FF5DF840" bold="true" box="[207,232,1979,2003]" pageId="48" pageNumber="49">pc</emphasis>
—papillated cone;
<emphasis id="B974613F150BFFB8FE1974B6FE0AF840" bold="true" box="[426,447,1979,2003]" pageId="48" pageNumber="49">pl</emphasis>
—posterior lamella;
<emphasis id="B974613F150BFFB8FD2174B6FD00F840" bold="true" box="[658,693,1979,2003]" pageId="48" pageNumber="49">plp</emphasis>
—posterior lateral process;
<emphasis id="B974613F150BFFB8FC7C74B6FC53F840" bold="true" box="[975,998,1979,2003]" pageId="48" pageNumber="49">so</emphasis>
—seminal opening.
<emphasis id="B974613F150BFFB8FB0074B7FA9EF840" bold="true" box="[1203,1323,1978,2003]" pageId="48" pageNumber="49">Scale bars:</emphasis>
1 mm (
<emphasis id="B974613F150BFFB8FAC874B7FA38F840" bold="true" box="[1403,1421,1978,2003]" pageId="48" pageNumber="49">A</emphasis>
), 0.1 mm (
<emphasis id="B974613F150BFFB8FF4074D2FE94F864" bold="true" box="[243,289,2015,2039]" pageId="48" pageNumber="49">BE</emphasis>
), 0.01 mm (
<emphasis id="B974613F150BFFB8FE1374D2FE1AF864" bold="true" box="[416,431,2015,2039]" pageId="48" pageNumber="49">F</emphasis>
).
</paragraph>
</caption>
<caption id="DF7FEDA5150AFFB9FF2475AFFB40F825" ID-DOI="http://doi.org/10.5281/zenodo.3896603" ID-Zenodo-Dep="3896603" httpUri="https://zenodo.org/record/3896603/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,250,1698,1723]" targetBox="[151,1435,181,1673]" targetPageId="49">
<paragraph id="8BBFBD2D150AFFB9FF2475AFFB40F825" blockId="49.[151,1437,1698,1975]" pageId="49" pageNumber="50">
<emphasis id="B974613F150AFFB9FF2475AFFEAEF928" bold="true" box="[151,283,1698,1723]" pageId="49" pageNumber="50">FIGURE 26</emphasis>
.
<taxonomicName id="4C00C6AE150AFFB9FE9975A9FD29F928" authority="(Strasser, 1973 a)" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[298,668,1699,1723]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F150AFFB9FE9975A9FE51F928" box="[298,484,1699,1723]" italics="true" pageId="49" pageNumber="50">Haasea vidinensis</emphasis>
(
<bibRefCitation id="EF91C0DC150AFFB9FE4075AEFD21F928" author="Strasser, K." box="[499,660,1699,1723]" pageId="49" pageNumber="50" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser, 1973a</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B974613F150AFFB9FD1475AFFD75F928" bold="true" box="[679,704,1698,1723]" pageId="49" pageNumber="50">A.</emphasis>
Lectotype male (NMNHS10807), right anterior gonopod, lateral view.
<emphasis id="B974613F150AFFB9FF2475CAFF1BF94C" bold="true" box="[151,174,1735,1759]" pageId="49" pageNumber="50">B.</emphasis>
Paralectotype male (NMNHS10808), left anterior gonopod, lateral view.
<emphasis id="B974613F150AFFB9FC1675CBFC0BF94C" bold="true" box="[933,958,1734,1759]" pageId="49" pageNumber="50">C.</emphasis>
<taxonomicName id="4C00C6AE150AFFB9FC7675C5FA85F94C" authority="Makarov, 2008" authorityName=", Makarov" authorityYear="2008" box="[965,1328,1734,1759]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F150AFFB9FC7675C5FB38F94C" box="[965,1165,1735,1759]" italics="true" pageId="49" pageNumber="50">H. guidononveilleri</emphasis>
<bibRefCitation id="EF91C0DC150AFFB9FB2675CBFA85F94C" author="Makarov, S. E." box="[1173,1328,1734,1759]" pageId="49" pageNumber="50" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov, 2008</bibRefCitation>
</taxonomicName>
<emphasis id="B974613F150AFFB9FA8475CAFA20F94C" bold="true" box="[1335,1429,1735,1759]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A247DC44150AFFB9FA8475CAFA20F94C" box="[1335,1429,1735,1759]" pageId="49" pageNumber="50">syn. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZB), left anterior gonopod, lateral view.
<emphasis id="B974613F150AFFB9FD6775E6FD59F890" bold="true" box="[724,748,1771,1795]" pageId="49" pageNumber="50">D.</emphasis>
Male from Rtanj Mt, Serbia, right anterior gonopod, lateral view.
<emphasis id="B974613F150AFFB9FA3675E6FA29F890" bold="true" box="[1413,1436,1771,1795]" pageId="49" pageNumber="50">E.</emphasis>
Male from Kučaj Mts, Serbia, left anterior gonopod, lateral view.
<emphasis id="B974613F150AFFB9FC837402FCF6F8B4" bold="true" box="[816,835,1807,1831]" pageId="49" pageNumber="50">F.</emphasis>
Male from Kučaj Mts, Serbia, left anterior gonopod, mesal view.
<emphasis id="B974613F150AFFB9FF67743FFF5BF8D8" bold="true" box="[212,238,1842,1867]" pageId="49" pageNumber="50">G.</emphasis>
Lectotype male (NMNHS10807), right posterior gonopod, anteromesal view.
<emphasis id="B974613F150AFFB9FBA2743EFB9EF8D8" bold="true" box="[1041,1067,1843,1867]" pageId="49" pageNumber="50">H.</emphasis>
Male from Kučaj Mts, Serbia, right posterior gonopod, mesal view.
<emphasis id="B974613F150AFFB9FE69745AFE5FF8FC" bold="true" box="[474,490,1879,1903]" pageId="49" pageNumber="50">I.</emphasis>
<taxonomicName id="4C00C6AE150AFFB9FE437455FCE3F8FC" authority="Makarov, 2008" authorityName=", Makarov" authorityYear="2008" box="[496,854,1878,1903]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F150AFFB9FE437455FD03F8FC" box="[496,694,1879,1903]" italics="true" pageId="49" pageNumber="50">H. guidononveilleri</emphasis>
<bibRefCitation id="EF91C0DC150AFFB9FD0F745BFCE3F8FC" author="Makarov, S. E." box="[700,854,1878,1903]" pageId="49" pageNumber="50" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov, 2008</bibRefCitation>
</taxonomicName>
<emphasis id="B974613F150AFFB9FCE8745AFC0DF8FC" bold="true" box="[859,952,1879,1903]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A247DC44150AFFB9FCE8745AFC0DF8FC" box="[859,952,1879,1903]" pageId="49" pageNumber="50">syn. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZB), posterior gonopods, posterior view.
<emphasis id="B974613F150AFFB9FEBC7476FE97F800" bold="true" box="[271,290,1915,1939]" pageId="49" pageNumber="50">J.</emphasis>
Male from Rtanj, Mt, Serbia, posterior gonopods, posteror view.
<emphasis id="B974613F150AFFB9FC017477FBC1F800" bold="true" box="[946,1140,1914,1939]" pageId="49" pageNumber="50">Abbreviations: ab</emphasis>
—anterior branch;
<emphasis id="B974613F150AFFB9FA807476FAF9F800" bold="true" box="[1331,1356,1915,1939]" pageId="49" pageNumber="50">cp</emphasis>
—coxal process;
<emphasis id="B974613F150AFFB9FF427492FEB8F824" bold="true" box="[241,269,1951,1975]" pageId="49" pageNumber="50">pb</emphasis>
—posterior branch;
<emphasis id="B974613F150AFFB9FE6A7492FE49F824" bold="true" box="[473,508,1951,1975]" pageId="49" pageNumber="50">plp</emphasis>
—posterior lateral process;
<emphasis id="B974613F150AFFB9FCA67492FCAAF824" bold="true" box="[789,799,1951,1975]" pageId="49" pageNumber="50">s</emphasis>
—sternum
<emphasis id="B974613F150AFFB9FC237492FC2DF824" bold="true" box="[912,920,1951,1975]" pageId="49" pageNumber="50">t</emphasis>
—telopodite.
<emphasis id="B974613F150AFFB9FB977493FB2EF824" bold="true" box="[1060,1179,1950,1975]" pageId="49" pageNumber="50">Scale bars:</emphasis>
0.1 mm.
</paragraph>
</caption>
<paragraph id="8BBFBD2D1509FFBAFF747395FDB5FE3C" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
<emphasis id="B974613F1509FFBAFF747395FDB5FF20" bold="true" box="[199,512,152,179]" pageId="50" pageNumber="51">Other material examined:</emphasis>
<materialsCitation id="3B68B7701509FFBAFDBB7394FBE7FF44" ID-GBIF-Occurrence="2636224573" collectingDate="2016-09-11" collectionCode="IZB" collectorName="S. Curcic" country="Serbia" county="Radovanska" location="Kucaj Mts" municipality="Cave" pageId="50" pageNumber="51" specimenCount="4" specimenCount-male="4" stateProvince="Boljevac">
<collectingCountry id="F317FDBD1509FFBAFDBB7394FDC5FF20" box="[520,624,153,179]" name="Serbia" pageId="50" pageNumber="51">SERBIA</collectingCountry>
:
<specimenCount id="9D0676A41509FFBAFDCF7394FD09FF20" box="[636,700,153,179]" pageId="50" pageNumber="51" type="male">4 ♂♂</specimenCount>
(one for SEM),
<collectingCounty id="62DEC5A11509FFBAFCCB7394FBB4FF20" box="[888,1025,153,179]" pageId="50" pageNumber="51">Radovanska</collectingCounty>
vrelska
<collectingMunicipality id="6BDB27571509FFBAFBD17394FB2BFF20" box="[1122,1182,153,179]" pageId="50" pageNumber="51">Cave</collectingMunicipality>
,
<location id="8EDFEBF61509FFBAFB187394FA91FF20" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFB187394FA91FF20" box="[1195,1316,153,179]" country="Serbia" county="Radovanska" municipality="Cave" name="Kucaj Mts" pageId="50" pageNumber="51" stateProvince="Boljevac">Kučaj Mts</location>
, village of
<location id="8EDFEBF61509FFBAFF2473B0FEB7FF44" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFF2473B0FEB7FF44" box="[151,258,189,215]" country="Serbia" county="Radovanska" municipality="Cave" name="Jablanica" pageId="50" pageNumber="51" stateProvince="Boljevac">Jablanica</location>
, ner
<collectingRegion id="49C473CF1509FFBAFE8E73B0FE16FF44" box="[317,419,189,215]" country="Serbia" name="Boljevac" pageId="50" pageNumber="51">Boljevac</collectingRegion>
,
<collectingMethod id="5241C53A1509FFBAFE0273B0FD88FF44" box="[433,573,189,215]" pageId="50" pageNumber="51">pit-fall traps</collectingMethod>
collected on
<date id="FFBE9BED1509FFBAFD6873B0FCEFFF44" box="[731,858,189,215]" pageId="50" pageNumber="51" value="2016-09-11">
<collectingDate id="EFFA62051509FFBAFD6873B0FCEFFF44" box="[731,858,189,215]" pageId="50" pageNumber="51" value="2016-09-11">09.11.2016</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFCDB73B0FC66FF44" box="[872,979,188,215]" pageId="50" pageNumber="51">S. Ćurčić</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFBAB73B0FBFFFF44" box="[1048,1098,189,215]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFBD373B0FCA9FF68" ID-GBIF-Occurrence="2636224498" collectingDate="2008-04-08" collectionCode="IZB" collectorName="D. Antic &amp; D. Stojanovic" country="Serbia" county="Radovanska" location="Kucaj Mts" municipality="Cave" pageId="50" pageNumber="51" specimenCount="4" specimenCount-juvenile="3" specimenCount-male="1" stateProvince="Boljevac">
<specimenCount id="9D0676A41509FFBAFBD373B0FB3AFF44" box="[1120,1167,189,215]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0676A41509FFBAFB2E73B0FAA8FF44" box="[1181,1309,189,215]" pageId="50" pageNumber="51" type="juvenile">3 juveniles</specimenCount>
, same data except
<date id="FFBE9BED1509FFBAFF5973ECFEDEFF68" box="[234,363,225,251]" pageId="50" pageNumber="51" value="2008-04-08">
<collectingDate id="EFFA62051509FFBAFF5973ECFEDEFF68" box="[234,363,225,251]" pageId="50" pageNumber="51" value="2008-04-08">04.08.2008</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFECB73ECFE6CFF68" box="[376,473,225,251]" pageId="50" pageNumber="51">D. Antić</collectorName>
&amp;
<collectorName id="26F5D8FB1509FFBAFDB173ECFD2BFF68" box="[514,670,225,251]" pageId="50" pageNumber="51">D. Stojanović</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFD5173ECFCA1FF68" box="[738,788,225,251]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFC9873ECFF6FFE8C" ID-GBIF-Occurrence="2636224521" collectingDate="2010-07-07" collectionCode="IZB" collectorName="D. Antic" country="Serbia" county="Radovanska" location="Kucaj Mts" municipality="Cave" pageId="50" pageNumber="51" specimenCount="1" specimenCount-juvenile="1" stateProvince="Boljevac">
<specimenCount id="9D0676A41509FFBAFC9873ECFC2AFF68" box="[811,927,225,251]" pageId="50" pageNumber="51" type="juvenile">1 juvenile</specimenCount>
, same data except
<date id="FFBE9BED1509FFBAFBC973ECFB4EFF68" box="[1146,1275,225,251]" pageId="50" pageNumber="51" value="2010-07-07">
<collectingDate id="EFFA62051509FFBAFBC973ECFB4EFF68" box="[1146,1275,225,251]" pageId="50" pageNumber="51" value="2010-07-07">07.07.2010</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFABB73ECFADCFF68" box="[1288,1385,225,251]" pageId="50" pageNumber="51">D. Antić</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFF137208FF67FE8C" box="[160,210,261,287]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFF547208FA22FEB3" ID-GBIF-Occurrence="2636224373" box="[231,1431,261,288]" collectionCode="IZB" collectorName="D. Antic" country="Serbia" county="Ponor Radovanske" location="Jablanica" municipality="Cave" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Boljevac">
<specimenCount id="9D0676A41509FFBAFF547208FEA4FE8C" box="[231,273,261,287]" pageId="50" pageNumber="51" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62DEC5A11509FFBAFE937208FE41FE8C" box="[288,500,261,287]" pageId="50" pageNumber="51">Ponor Radovanske</collectingCounty>
reke
<collectingMunicipality id="6BDB27571509FFBAFD877208FDC5FE8C" box="[564,624,261,287]" pageId="50" pageNumber="51">Cave</collectingMunicipality>
, village
<location id="8EDFEBF61509FFBAFD637208FC8EFE8C" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFD637208FC8EFE8C" box="[720,827,261,287]" country="Serbia" county="Ponor Radovanske" municipality="Cave" name="Jablanica" pageId="50" pageNumber="51" stateProvince="Boljevac">Jablanica</location>
, near
<collectingRegion id="49C473CF1509FFBAFCCC7208FC50FE8C" box="[895,997,261,287]" country="Serbia" name="Boljevac" pageId="50" pageNumber="51">Boljevac</collectingRegion>
,
<location id="8EDFEBF61509FFBAFC427208FBDFFE8C" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFC427208FBDFFE8C" box="[1009,1130,261,287]" country="Serbia" county="Ponor Radovanske" municipality="Cave" name="Kucaj Mts" pageId="50" pageNumber="51" stateProvince="Boljevac">Kučaj Mts</location>
, 2016.,
<collectorName id="26F5D8FB1509FFBAFB777208FA96FE8C" box="[1220,1315,261,287]" pageId="50" pageNumber="51">D. Antić</collectorName>
leg.(
<collectionCode id="ED1125E81509FFBAFAEE7208FA3AFEB3" box="[1373,1423,261,288]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFF247224FD4EFED0" ID-GBIF-Occurrence="2636224348" box="[151,763,297,323]" collectingDate="2019-02-10" collectionCode="IZB" collectorName="D. Antic" country="Serbia" county="Ponor Radovanske" location="Jablanica" municipality="Cave" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Boljevac">
<specimenCount id="9D0676A41509FFBAFF247224FF74FED0" box="[151,193,297,323]" pageId="50" pageNumber="51" type="female">1 ♀</specimenCount>
, same data except
<date id="FFBE9BED1509FFBAFE257224FDA2FED0" box="[406,535,297,323]" pageId="50" pageNumber="51" value="2019-02-10">
<collectingDate id="EFFA62051509FFBAFE257224FDA2FED0" box="[406,535,297,323]" pageId="50" pageNumber="51" value="2019-02-10">02.10.2019</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFD917224FD34FED0" box="[546,641,297,323]" pageId="50" pageNumber="51">D. Antić</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFD727224FD46FED0" box="[705,755,297,323]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFCB47224FD60FEF4" ID-GBIF-Occurrence="2636224500" collectingDate="2007-11-14" collectionCode="IZB" collectorName="S. Ognjenovic" country="Serbia" location="Mt. Rtanj" municipality="Zvecani Propast Pit" pageId="50" pageNumber="51" specimenCount="22" specimenCount-female="2" specimenCount-male="20">
<specimenCount id="9D0676A41509FFBAFCB47224FCE3FED0" box="[775,854,297,323]" pageId="50" pageNumber="51" type="male">20 ♂♂</specimenCount>
,
<specimenCount id="9D0676A41509FFBAFCD17224FC17FED0" box="[866,930,297,323]" pageId="50" pageNumber="51" type="female">2 ♀♀</specimenCount>
,
<collectingMunicipality id="6BDB27571509FFBAFC1C7224FB3BFED0" box="[943,1166,297,323]" pageId="50" pageNumber="51">Zvečani Propast Pit</collectingMunicipality>
,
<location id="8EDFEBF61509FFBAFB2A7224FABDFED0" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFB2A7224FABDFED0" box="[1177,1288,297,323]" country="Serbia" municipality="Zvecani Propast Pit" name="Mt. Rtanj" pageId="50" pageNumber="51">Mt. Rtanj</location>
,
<collectingMethod id="5241C53A1509FFBAFAA17224FA29FED0" box="[1298,1436,297,323]" pageId="50" pageNumber="51">pit-fall traps</collectingMethod>
collected on
<date id="FFBE9BED1509FFBAFE9A7240FE1CFEF4" box="[297,425,333,359]" pageId="50" pageNumber="51" value="2007-11-14">
<collectingDate id="EFFA62051509FFBAFE9A7240FE1CFEF4" box="[297,425,333,359]" pageId="50" pageNumber="51" value="2007-11-14">14.11.2007</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFE057240FDEDFEF4" box="[438,600,333,359]" pageId="50" pageNumber="51">S. Ognjenović</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFD287240FD78FEF4" box="[667,717,333,359]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D0676A41509FFBAFD507240FC96FEF4" box="[739,803,333,359]" pageId="50" pageNumber="51" type="male">3 ♂♂</specimenCount>
(one for SEM),
<materialsCitation id="3B68B7701509FFBAFC517240FBF1FE18" ID-GBIF-Occurrence="2636224399" collectingDate="2012-09-23" collectionCode="IZB" collectorName="D. Antic" country="Serbia" county="Samar-Veliki Pester Cave" location="Kalafat Mt." municipality="Kopajkosara" pageId="50" pageNumber="51" specimenCount="4" specimenCount-female="1" specimenCount-juvenile="3">
<specimenCount id="9D0676A41509FFBAFC517240FBBBFEF4" box="[994,1038,333,359]" pageId="50" pageNumber="51" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D0676A41509FFBAFBAE7240FB29FEF4" box="[1053,1180,333,359]" pageId="50" pageNumber="51" type="juvenile">3 juveniles</specimenCount>
,
<collectingCounty id="62DEC5A11509FFBAFB1A7240FF64FE18" pageId="50" pageNumber="51">Samar—Veliki Pešter Cave</collectingCounty>
System, village of
<collectingMunicipality id="6BDB27571509FFBAFE1D727CFD8AFE18" box="[430,575,369,395]" pageId="50" pageNumber="51">Kopajkošara</collectingMunicipality>
,
<location id="8EDFEBF61509FFBAFDF9727CFD7AFE18" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFDF9727CFD7AFE18" box="[586,719,369,395]" country="Serbia" county="Samar-Veliki Pester Cave" municipality="Kopajkosara" name="Kalafat Mt." pageId="50" pageNumber="51">Kalafat Mt.</location>
,
<date id="FFBE9BED1509FFBAFD68727CFCE9FE18" box="[731,860,369,395]" pageId="50" pageNumber="51" value="2012-09-23">
<collectingDate id="EFFA62051509FFBAFD68727CFCE9FE18" box="[731,860,369,395]" pageId="50" pageNumber="51" value="2012-09-23">23.09.2012</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFCDB727CFC7DFE18" box="[872,968,369,395]" pageId="50" pageNumber="51">D. Antić</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFBB9727CFB89FE18" box="[1034,1084,369,395]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B68B7701509FFBAFBE2727CFE49FE3C" ID-GBIF-Occurrence="2636224451" collectingDate="2012-06-26" collectionCode="IZB" collectorName="D. Antic" country="Serbia" county="Samar-Veliki Pester Cave" location="Kalafat Mt." municipality="Kopajkosara" pageId="50" pageNumber="51" specimenCount="2" specimenCount-juvenile="2">
<specimenCount id="9D0676A41509FFBAFBE2727CFB7AFE18" box="[1105,1231,369,395]" pageId="50" pageNumber="51" type="juvenile">2 juveniles</specimenCount>
, same data except
<date id="FFBE9BED1509FFBAFF247298FEADFE3C" box="[151,280,405,431]" pageId="50" pageNumber="51" value="2012-06-26">
<collectingDate id="EFFA62051509FFBAFF247298FEADFE3C" box="[151,280,405,431]" pageId="50" pageNumber="51" value="2012-06-26">26.06.2012</collectingDate>
</date>
,
<collectorName id="26F5D8FB1509FFBAFE907298FE37FE3C" box="[291,386,405,431]" pageId="50" pageNumber="51">D. Antić</collectorName>
leg. (
<collectionCode id="ED1125E81509FFBAFE717298FE41FE3C" box="[450,500,405,431]" pageId="50" pageNumber="51">IZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61509FFBAFF7472B5FC3BFDD3" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BBFBD2D1509FFBAFF7472B5FC3BFDD3" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
<emphasis id="B974613F1509FFBAFF7472B5FEECFE40" bold="true" box="[199,345,440,467]" pageId="50" pageNumber="51">Distribution</emphasis>
(
<figureCitation id="133BA1A81509FFBAFEDB72B4FE0BFE40" box="[360,446,441,467]" captionStart="FIGURE 38" captionStartId="67.[151,250,1157,1182]" captionTargetBox="[151,1435,417,1133]" captionTargetId="figure@67.[151,1435,417,1133]" captionTargetPageId="67" captionText="FIGURE 38. Distribution of the genus Haasea." figureDoi="http://doi.org/10.5281/zenodo.3896629" httpUri="https://zenodo.org/record/3896629/files/figure.png" pageId="50" pageNumber="51">Figs 38</figureCitation>
,
<figureCitation id="133BA1A81509FFBAFE7A72B4FE5CFE40" box="[457,489,441,467]" captionStart="FIGURE 42" captionStartId="70.[151,250,1979,2004]" captionTargetBox="[188,1399,1135,1953]" captionTargetId="figure@70.[188,1399,1135,1953]" captionTargetPageId="70" captionText="FIGURE 42. Distribution of H. intermedia Mršić, 1985, H. lacusnigri (Gulička, 1968), H. makarovi sp. nov., H. microcorna (Strasser, 1971), H. plasana (Verhoeff, 1899) and H. vidinensis (Strasser, 1973a)." figureDoi="http://doi.org/10.5281/zenodo.3896637" httpUri="https://zenodo.org/record/3896637/files/figure.png" pageId="50" pageNumber="51">42</figureCitation>
).
<collectingCountry id="F317FDBD1509FFBAFE4F72B4FDE9FE40" box="[508,604,441,467]" name="Bulgaria" pageId="50" pageNumber="51">Bulgaria</collectingCountry>
: Džamiite Pot hole, near
<collectingRegion id="49C473CF1509FFBAFCC472B4FC00FE40" box="[887,949,441,467]" country="Bulgaria" name="Vidin" pageId="50" pageNumber="51">Vidin</collectingRegion>
(
<bibRefCitation id="EF91C0DC1509FFBAFC7072B4FBD5FE40" author="Strasser, K." box="[963,1120,441,467]" pageId="50" pageNumber="51" pagination="313 - 345" refId="ref46776" refString="Strasser, K. (1971) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 52 (3 - 5), 313 - 345." type="journal article" year="1971">Strasser 1971</bibRefCitation>
);
<collectingCountry id="F317FDBD1509FFBAFBC072B4FB0EFE40" box="[1139,1211,441,467]" name="Serbia" pageId="50" pageNumber="51">Serbia</collectingCountry>
: Jama Ledenica Pit, Mt. Tupižnica (
<bibRefCitation id="EF91C0DC1509FFBAFEF272D0FE5EFE64" author="Makarov, S. E." box="[321,491,477,503]" pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov 2008</bibRefCitation>
), Jezava Cave System, Mt. Kalafat, near Kopajkošara (
<bibRefCitation id="EF91C0DC1509FFBAFBFE72D0FB43FE64" author="Makarov, S. E." box="[1101,1270,477,503]" pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov 2008</bibRefCitation>
), Samar-Veliki Pešter Cave System, Mt. Kalafat, near Kopajkošara (
<bibRefCitation id="EF91C0DC1509FFBAFD55710CFC3AFD88" author="Makarov, S. E." box="[742,911,513,539]" pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov 2008</bibRefCitation>
; present study); Zvečani Propast Pit, Mt. Rtanj (present study); Radovanska reka Cave, Kučaj Mts. (present study).
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61509FFBAFF747144FCF0FDF0" box="[199,837,584,611]" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8BBFBD2D1509FFBAFF747144FCF0FDF0" blockId="50.[151,1437,152,1763]" box="[199,837,584,611]" pageId="50" pageNumber="51">
<materialsCitation id="3B68B7701509FFBAFF747144FCF4FDF0" ID-GBIF-Occurrence="2636224617" box="[199,833,584,611]" country="Bulgaria" location="Dzamiite Pot" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Vidin" typeStatus="holotype">
<emphasis id="B974613F1509FFBAFF747144FED3FDF0" bold="true" box="[199,358,584,611]" pageId="50" pageNumber="51">
<typeStatus id="54BB038F1509FFBAFF747144FEB7FDF0" box="[199,258,585,611]" pageId="50" pageNumber="51">Type</typeStatus>
locality.
</emphasis>
<location id="8EDFEBF61509FFBAFEDF7144FE4AFDF0" LSID="urn:lsid:plazi:treatment:03A90C3B1515FFBAFF247571FB32F970:8EDFEBF61509FFBAFEDF7144FE4AFDF0" box="[364,511,585,611]" country="Bulgaria" name="Dzamiite Pot" pageId="50" pageNumber="51" stateProvince="Vidin">Džamiite Pot</location>
hole, near
<collectingRegion id="49C473CF1509FFBAFDC87144FD0CFDF0" box="[635,697,585,611]" country="Bulgaria" name="Vidin" pageId="50" pageNumber="51">Vidin</collectingRegion>
in
<collectingCountry id="F317FDBD1509FFBAFD6E7144FCF4FDF0" box="[733,833,585,611]" name="Bulgaria" pageId="50" pageNumber="51">Bulgaria</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31AEEA61509FFBAFF747161FB32F970" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8BBFBD2D1509FFBAFF747161FC71FB54" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
<emphasis id="B974613F1509FFBAFF747161FEFAFD14" bold="true" box="[199,335,620,647]" pageId="50" pageNumber="51">Comments.</emphasis>
<bibRefCitation id="EF91C0DC1509FFBAFEE57160FDB8FD14" author="Strasser, K." box="[342,525,621,647]" pageId="50" pageNumber="51" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser (1973a)</bibRefCitation>
established the new subgenus
<taxonomicName id="4C00C6AE1509FFBAFCD77163FBF3FD14" authority="Strasser" authorityName="Strasser" authorityYear="1973" box="[868,1094,621,647]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Histrosoma">
<emphasis id="B974613F1509FFBAFCD77163FC53FD14" box="[868,998,622,647]" italics="true" pageId="50" pageNumber="51">Histrosoma</emphasis>
Strasser
</taxonomicName>
for his
<taxonomicName id="4C00C6AE1509FFBAFB287163FA98FD14" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[1179,1325,621,647]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFB287163FA98FD14" box="[1179,1325,621,647]" italics="true" pageId="50" pageNumber="51">H. vidinensis</emphasis>
</taxonomicName>
and characterized it by the absence of “Pugio” and “Epithema”. Later, in his description of
<taxonomicName id="4C00C6AE1509FFBAFBF8719FFF57FD5C" authority=", Makarov (2008)" authorityName=", Makarov" authorityYear="2008" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1509FFBAFBF8719FFA9BFD38" box="[1099,1326,657,683]" italics="true" pageId="50" pageNumber="51">H. guidononveilleri,</emphasis>
<bibRefCitation id="EF91C0DC1509FFBAFA84719CFF57FD5C" author="Makarov, S. E." pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov (2008)</bibRefCitation>
</taxonomicName>
stated that it is closely related to
<taxonomicName id="4C00C6AE1509FFBAFDDB71BBFD49FD5C" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[616,764,693,719]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFDDB71BBFD49FD5C" box="[616,764,693,719]" italics="true" pageId="50" pageNumber="51">H. vidinensis</emphasis>
</taxonomicName>
and gave some differences between the two species. Also, he (
<bibRefCitation id="EF91C0DC1509FFBAFF7271D4FEDFFD60" author="Makarov, S. E." box="[193,362,729,755]" pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov 2008</bibRefCitation>
) disagreed with Strasser (1973) about the absence of “Epithema” in
<emphasis id="B974613F1509FFBAFBC571D4FB58FD60" box="[1142,1261,729,755]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="4C00C6AE1509FFBAFBC571D4FB5CFD60" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[1142,1257,729,755]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">vidinensis</taxonomicName>
,
</emphasis>
confirming the presence of this structure in both taxa as a reduced process and disagreed with the validity of
<taxonomicName id="4C00C6AE1509FFBAFB1B71F3FA9FFC84" authorityName="Strasser" authorityYear="1973" box="[1192,1322,766,791]" class="Diplopoda" family="Haaseidae" genus="Orobainosoma" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Histrosoma">
<emphasis id="B974613F1509FFBAFB1B71F3FA9FFC84" box="[1192,1322,766,791]" italics="true" pageId="50" pageNumber="51">Histrosoma</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF91C0DC1509FFBAFA8471F0FF57FCA8" author="Makarov, S. E." pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov (2008)</bibRefCitation>
compared his
<taxonomicName id="4C00C6AE1509FFBAFE30702CFD89FCA8" authorityName=", Makarov" authorityYear="2008" box="[387,572,801,827]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1509FFBAFE30702CFD89FCA8" box="[387,572,801,827]" italics="true" pageId="50" pageNumber="51">guidononveilleri</emphasis>
</taxonomicName>
with
<emphasis id="B974613F1509FFBAFDC8702CFD44FCA8" box="[635,753,801,827]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="4C00C6AE1509FFBAFDC8702CFD5BFCA8" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[635,750,801,827]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">vidinensis</taxonomicName>
,
</emphasis>
stating, among others, the presence of both rudimentary “Pugio” and “Cornicle” in
<taxonomicName id="4C00C6AE1509FFBAFE257048FDFAFCCC" authorityName=", Makarov" authorityYear="2008" box="[406,591,837,863]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1509FFBAFE257048FDFAFCCC" box="[406,591,837,863]" italics="true" pageId="50" pageNumber="51">guidononveilleri</emphasis>
</taxonomicName>
, structures considered absent by
<bibRefCitation id="EF91C0DC1509FFBAFC0F7048FBC1FCCC" author="Strasser, K." box="[956,1140,837,863]" pageId="50" pageNumber="51" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser (1973a)</bibRefCitation>
. After examination of both
<typeStatus id="54BB038F1509FFBAFF247064FF60FC10" box="[151,213,873,899]" pageId="50" pageNumber="51">types</typeStatus>
, as well as a new material from
<collectingCountry id="F317FDBD1509FFBAFD8C7064FD3CFC10" box="[575,649,873,899]" name="Serbia" pageId="50" pageNumber="51">Serbia</collectingCountry>
, we noticed that posterior lamella (
<emphasis id="B974613F1509FFBAFBA67064FB99FC10" bold="true" box="[1045,1068,873,899]" pageId="50" pageNumber="51">pl</emphasis>
) (“Epithema”) is present in both taxa. Mesal process (
<emphasis id="B974613F1509FFBAFE3B7080FE1AFC34" bold="true" box="[392,431,909,935]" pageId="50" pageNumber="51">mp</emphasis>
) (“Keratit”) is strongly developed, parallel and closely fitting to angiocoxite, knife-like, with basal posterior tooth. The size of the basal tooth on
<emphasis id="B974613F1509FFBAFCA070BCFC8FFC58" bold="true" box="[787,826,945,971]" pageId="50" pageNumber="51">mp</emphasis>
is variable, from short, triangular to somewhat longer, thorn-like. Also, in
<typeStatus id="54BB038F1509FFBAFEC070D8FE16FC7C" box="[371,419,981,1007]" pageId="50" pageNumber="51">type</typeStatus>
specimens of
<taxonomicName id="4C00C6AE1509FFBAFDF670D8FD4BFC7C" authorityName=", Makarov" authorityYear="2008" box="[581,766,981,1007]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1509FFBAFDF670D8FD4BFC7C" box="[581,766,981,1007]" italics="true" pageId="50" pageNumber="51">guidononveilleri</emphasis>
</taxonomicName>
, mesal process on the right gonopod is without basal tooth. Both species are characterized by the presence of a reduced, small and triangular anterior lateral process (
<emphasis id="B974613F1509FFBAFA9870F4FAE5FB80" bold="true" box="[1323,1360,1017,1043]" pageId="50" pageNumber="51">alp</emphasis>
) (“Pugio”) with additional mesal sharp projection. Also, both are characterized by the presence of the posterior lateral process (
<emphasis id="B974613F1509FFBAFF48774CFE97FBC8" bold="true" box="[251,290,1089,1115]" pageId="50" pageNumber="51">plp</emphasis>
) (“Cornicle”), variable from short triangular to somewhat longer, acuminate. It is obvious that
<bibRefCitation id="EF91C0DC1509FFBAFAF7774CFF5BFBEC" author="Strasser, K." pageId="50" pageNumber="51" pagination="411 - 470" refId="ref46801" refString="Strasser, K. (1973 a) Uber Diplopoden Bulgariens, III. Annales zoologici, 30 (15), 411 - 470." type="journal article" year="1973">Strasser (1973a)</bibRefCitation>
overlooked some structures in his
<taxonomicName id="4C00C6AE1509FFBAFDC57768FD53FBEC" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[630,742,1125,1151]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFDC57768FD53FBEC" box="[630,742,1125,1151]" italics="true" pageId="50" pageNumber="51">vidinensis</emphasis>
</taxonomicName>
. On the other hand,
<bibRefCitation id="EF91C0DC1509FFBAFC7F7768FB31FBEC" author="Makarov, S. E." box="[972,1156,1125,1151]" pageId="50" pageNumber="51" pagination="85 - 97" refId="ref45407" refString="Makarov, S. E. (2008). Haasea guidononveilleri n. sp., a new diplopod from Serbia and revision of the &quot; lacusnigri &quot; group of species (Diplopoda, Haaseidae). In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 85 - 97." type="book chapter" year="2008">Makarov (2008)</bibRefCitation>
made drawings from the left gonopod, which was somewhat flattened during the preparation of microscopic slides (
<figureCitation id="133BA1A81509FFBAFB227784FB43FB37" box="[1169,1270,1161,1188]" captionStart="FIGURE 26" captionStartId="49.[151,250,1698,1723]" captionTargetBox="[151,1435,181,1673]" captionTargetId="figure@49.[151,1435,181,1673]" captionTargetPageId="49" captionText="FIGURE 26. Haasea vidinensis (Strasser, 1973a). A. Lectotype male (NMNHS10807), right anterior gonopod, lateral view. B. Paralectotype male (NMNHS10808), left anterior gonopod, lateral view. C. H. guidononveilleri Makarov, 2008 syn. nov., holotype male (IZB), left anterior gonopod, lateral view. D. Male from Rtanj Mt, Serbia, right anterior gonopod, lateral view. E. Male from Kučaj Mts, Serbia, left anterior gonopod, lateral view. F. Male from Kučaj Mts, Serbia, left anterior gonopod, mesal view. G. Lectotype male (NMNHS10807), right posterior gonopod, anteromesal view. H. Male from Kučaj Mts, Serbia, right posterior gonopod, mesal view. I. H. guidononveilleri Makarov, 2008 syn. nov., holotype male (IZB), posterior gonopods, posterior view. J. Male from Rtanj, Mt, Serbia, posterior gonopods, posteror view. Abbreviations: ab—anterior branch; cp—coxal process; pb—posterior branch; plp—posterior lateral process; s—sternum t—telopodite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3896603" httpUri="https://zenodo.org/record/3896603/files/figure.png" pageId="50" pageNumber="51">Fig. 26C</figureCitation>
) whereas right anterior gonopod remained “normal” and is exactly matching
<taxonomicName id="4C00C6AE1509FFBAFCFE77A0FC08FB54" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[845,957,1197,1223]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFCFE77A0FC08FB54" box="[845,957,1197,1223]" italics="true" pageId="50" pageNumber="51">vidinensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBFBD2D1509FFBAFF7477DCFE20FAA0" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
Considering the absence of consistent differences between the two species and that both taxa inhabit the Carpatho-Balkan caves, near the Serbo-Bulgarian border, we consider
<taxonomicName id="4C00C6AE1509FFBAFC3C77FBFBDEFA9C" authorityName=", Makarov" authorityYear="2008" box="[911,1131,1269,1295]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="guidononveilleri">
<emphasis id="B974613F1509FFBAFC3C77FBFBDEFA9C" box="[911,1131,1269,1295]" italics="true" pageId="50" pageNumber="51">H. guidononveilleri</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="4C00C6AE1509FFBAFF4F7617FE3AFAA0" baseAuthorityName="Strasser" baseAuthorityYear="1973" box="[252,399,1305,1331]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFF4F7617FE3AFAA0" box="[252,399,1305,1331]" italics="true" pageId="50" pageNumber="51">H. vidinensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBFBD2D1509FFBAFF747630FBECF9E4" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
In the vial and on the microslide of
<typeStatus id="54BB038F1509FFBAFDD27630FD7DFAC4" box="[609,712,1341,1367]" pageId="50" pageNumber="51" type="lectotype">lectotype</typeStatus>
male of
<taxonomicName id="4C00C6AE1509FFBAFC9D7633FBD0FAC4" authority=", Strasser" authorityName="Strasser" box="[814,1125,1341,1367]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFC9D7633FC49FAC4" box="[814,1020,1341,1367]" italics="true" pageId="50" pageNumber="51">Haasea vidinensis</emphasis>
, Strasser
</taxonomicName>
clearly indicated on the label “
<typeStatus id="54BB038F1509FFBAFF78766CFEF4FAE8" box="[203,321,1377,1403]" pageId="50" pageNumber="51" type="holotype">holotypus</typeStatus>
”, as well as
<typeStatus id="54BB038F1509FFBAFE7F766CFD82FAE8" box="[460,567,1377,1403]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
for the rest, but he did not designate a
<typeStatus id="54BB038F1509FFBAFC41766CFBE1FAE8" box="[1010,1108,1377,1403]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
and
<typeStatus id="54BB038F1509FFBAFB38766CFB43FAE8" box="[1163,1270,1377,1403]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
in the original description of the species (Strasser 1973). Later,
<bibRefCitation id="EF91C0DC1509FFBAFD777688FBADFA33" author="Stoev, P. &amp; Beron, P." box="[708,1048,1413,1440]" pageId="50" pageNumber="51" pagination="91 - 102" refId="ref46622" refString="Stoev, P. &amp; Beron, P. (2001) On type specimens of Myriapoda (Chilopoda, Diplopoda) in the collection of the National Museum of Natural History in Sofia. Arthropoda Selecta, 9 (2), 91 - 102." type="journal article" year="2001">Stoev &amp; Beron (2001: 9697)</bibRefCitation>
repeated what is on the Strassers label. However, according to the ICZN 72.4.7: “The mere citation of “Type” or equivalent expression, in a published work other than that in which the nominal species-group taxon is established, or in an unpublished catalogue of a museum, or on a label, is not necessarily evidence that a specimen is or is fixed as any of the kinds of types referred to in this Chapter”. The designation of
<typeStatus id="54BB038F1509FFBAFDFC7518FD04F9BC" box="[591,689,1557,1583]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
and
<typeStatus id="54BB038F1509FFBAFD5B7518FCE6F9BC" box="[744,851,1557,1583]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
is only valid if it appears in the original description of the species. Since this is not a case with
<taxonomicName id="4C00C6AE1509FFBAFDC57537FCF7F9C0" box="[630,834,1593,1619]" class="Diplopoda" family="Haaseidae" genus="Haasea" kingdom="Animalia" order="Chordeumatida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vidinensis">
<emphasis id="B974613F1509FFBAFDC57537FCF7F9C0" box="[630,834,1593,1619]" italics="true" pageId="50" pageNumber="51">Haasea vidinensis</emphasis>
</taxonomicName>
, all the type specimens of this species in NMNHS are considered as
<typeStatus id="54BB038F1509FFBAFE857550FE2DF9E4" box="[310,408,1629,1655]" pageId="50" pageNumber="51" type="syntype">syntypes</typeStatus>
and we here designate a
<typeStatus id="54BB038F1509FFBAFD027550FCADF9E4" box="[689,792,1629,1655]" pageId="50" pageNumber="51" type="lectotype">lectotype</typeStatus>
to fix the species taxonomy.
</paragraph>
<paragraph id="8BBFBD2D1509FFBAFF74758CFB32F970" blockId="50.[151,1437,152,1763]" pageId="50" pageNumber="51">
In his original description, Strasser (1973) stated he examined five males, five females and two juveniles. All specimens of the
<typeStatus id="54BB038F1509FFBAFEEC75A8FE3AF92C" box="[351,399,1701,1727]" pageId="50" pageNumber="51">type</typeStatus>
series were found in NMNHS and MHNG, with note that one male in NMNHS is without gonopods and leg-pairs 10 and 11, these structures were found on a microslide in MHNG.
</paragraph>
</subSubSection>
</treatment>
</document>