286 lines
25 KiB
XML
286 lines
25 KiB
XML
<document ID-DOI="10.11646/zootaxa.5230.4.4" ID-GBIF-Dataset="9c1c539b-e42a-4721-a46c-812f8f3a7569" ID-ISSN="1175-5326" ID-Zenodo-Dep="7564131" ID-ZooBank="F6E85127-C4AC-408C-8059-C1F9B52FB92F" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1674544461123" checkinUser="plazi" docAuthor="Pires-Silva, Carlos M." docDate="2023" docId="03A787F9FFE6FFCB53AE2604FA130634" docLanguage="en" docName="zootaxa.5230.4.4.pdf" docOrigin="Zootaxa 5230 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5230.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euryachen Pires-Silva 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="469" masterDocId="FF9EFF81FFE7FFC953392313FF8E0202" masterDocTitle="Euryachen seuzei a new genus and species of Placusini (Coleoptera: Staphylinidae Aleocharinae) from Brazil" masterLastPageNumber="477" masterPageNumber="467" pageNumber="468" updateTime="1674582142166" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Euryachen seuzei a new genus and species of Placusini (Coleoptera: Staphylinidae Aleocharinae) from Brazil</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pires-Silva, Carlos M.</mods:namePart>
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<mods:affiliation>Programa de pós graduação em Sistemática, Taxonomia animal e Biodiversidade, Museu de Zoologia da Universidade de São Paulo, Laboratório de Coleoptera, São Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5230</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>467</mods:start>
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<mods:end>477</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5230.4.4</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5230.4.4</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9c1c539b-e42a-4721-a46c-812f8f3a7569</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7564131</mods:identifier>
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<mods:identifier type="ZooBank">F6E85127-C4AC-408C-8059-C1F9B52FB92F</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7564127" ID-Zenodo-Dep="7564127" LSID="urn:lsid:plazi:treatment:03A787F9FFE6FFCB53AE2604FA130634" httpUri="http://treatment.plazi.org/id/03A787F9FFE6FFCB53AE2604FA130634" lastPageId="2" lastPageNumber="469" pageId="1" pageNumber="468">
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<subSubSection box="[151,489,1303,1330]" pageId="1" pageNumber="468" type="nomenclature">
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<paragraph blockId="1.[151,489,1303,1330]" box="[151,489,1303,1330]" pageId="1" pageNumber="468">
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<heading bold="true" box="[151,489,1303,1330]" fontSize="11" level="1" pageId="1" pageNumber="468" reason="1">
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<emphasis bold="true" box="[151,489,1303,1330]" pageId="1" pageNumber="468">
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Genus
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<taxonomicName authority="Pires-Silva, 2023" authorityName="Pires-Silva" authorityYear="2023" box="[238,372,1303,1330]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[238,372,1303,1330]" italics="true" pageId="1" pageNumber="468">Euryachen</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[379,489,1303,1329]" pageId="1" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,844,1366,1391]" pageId="1" pageNumber="468" type="type_taxon">
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<paragraph blockId="1.[151,844,1366,1391]" box="[151,844,1366,1391]" pageId="1" pageNumber="468">
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<emphasis bold="true" box="[151,286,1366,1390]" pageId="1" pageNumber="468">
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<typeStatus box="[151,204,1366,1390]" pageId="1" pageNumber="468">Type</typeStatus>
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species
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</emphasis>
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<taxonomicName authority="Pires-Silva, 2023" authorityName="Pires-Silva" authorityYear="2023" box="[291,470,1366,1390]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="seuzei" status="gen. et sp. nov.">
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<emphasis box="[291,470,1366,1390]" italics="true" pageId="1" pageNumber="468">Euryachen seuzei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[476,631,1367,1391]" pageId="1" pageNumber="468" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, present designation.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="469" pageId="1" pageNumber="468" type="diagnosis">
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<paragraph blockId="1.[151,1437,1431,2033]" pageId="1" pageNumber="468">
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<emphasis bold="true" box="[151,276,1431,1457]" pageId="1" pageNumber="468">Diagnosis:</emphasis>
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Head spherical, widest behind eyes; with a broad and distinctive neck occupying the entire basal head width. Posterior angles of head indistinct. Antennomeres VII–X distinctively transverse. Mandibles with dorsal molar area with seven transverse rows of large denticles. Maxillae with galea slightly longer than lacinia; lacinia with about nine large teeth apically. Labium short, 2-articulated. Ligula short and broadly rounded. Pronotum transverse; anterior margin straight with fore angles short and rounded; posterior margin broadly and weakly arcuate, hind angles straight. Tibial spur present, formula 2-1-1. Last tarsomeres of all legs longer than the previous together. Tarsal formula 4-4-5. Abdomen parallel-sided; segments III–VII with two pairs of paratergites each; tergite V of males bearing 7–9 spine-like thicker bristles near middle; tergite VII slightly broader at posterior region; on males with thicker and darker bristles on lateral margins.
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</paragraph>
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<paragraph blockId="1.[151,1437,1431,2033]" pageId="1" pageNumber="468">
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<emphasis bold="true" box="[199,436,1755,1781]" pageId="1" pageNumber="468">Comparative notes:</emphasis>
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The genus
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<taxonomicName authority="Pires-Silva, 2023" authorityName="Pires-Silva" authorityYear="2023" box="[570,691,1755,1781]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[570,691,1755,1781]" italics="true" pageId="1" pageNumber="468">Euryachen</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[700,806,1755,1781]" pageId="1" pageNumber="468">
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<taxonomicNameLabel box="[700,806,1755,1781]" pageId="1" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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, is most similar to
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<taxonomicName authority="Sharp" authorityName="Sharp" authorityYear="1883" box="[1028,1178,1755,1781]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus">
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<emphasis box="[1028,1104,1756,1781]" italics="true" pageId="1" pageNumber="468">Euvira</emphasis>
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Sharp
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</taxonomicName>
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in overall appearance, but is distinguishable in
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[428,531,1791,1817]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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from any other genus by a combination of unique characters: distinct and broad neck, occupying the entire basal head width (unique in the tribe), distinct and short anterior angles of pronotum, and antennomeres VII–X transverse. Moreover, members of
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[826,929,1863,1889]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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may also present secondary sexual characteristics. For instance, males of
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<taxonomicName authority="Erichson" authorityName="Erichson" authorityYear="1837" box="[482,680,1899,1925]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus">
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<emphasis box="[482,572,1899,1925]" italics="true" pageId="1" pageNumber="468">Placusa</emphasis>
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Erichson
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</taxonomicName>
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have modified tergite VIII, forming band of varied sizes spines on posterior margin, while in males of
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<taxonomicName authorityName="Sharp" authorityYear="1883" box="[552,628,1936,1961]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus">
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<emphasis box="[552,628,1936,1961]" italics="true" pageId="1" pageNumber="468">Euvira</emphasis>
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</taxonomicName>
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the tergite VIII is larger and prolonged posteriorly. Males of
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<taxonomicName authorityName="Pires-Silva" authorityYear="2023" box="[1315,1436,1935,1961]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus">
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<emphasis box="[1315,1436,1935,1961]" italics="true" pageId="1" pageNumber="468">Euryachen</emphasis>
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</taxonomicName>
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have thicker spine-like bristles on tergite V, and also bear thick bristles on lateral margins of tergite VII. As far as is known, this kind of sexual dimorphism is unique in
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[735,841,2007,2033]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="468" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="2.[151,1436,151,501]" pageId="2" pageNumber="469">
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The pronotum in the tribe is in mostly transverse. In
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<taxonomicName authorityName="Pires-Silva" authorityYear="2023" box="[798,919,151,177]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[798,919,151,177]" italics="true" pageId="2" pageNumber="469">Euryachen</emphasis>
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</taxonomicName>
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, the pronotum is transverse, similar to that of
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<taxonomicName authorityName="Sharp" authorityYear="1883" box="[151,227,187,212]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,227,187,212]" italics="true" pageId="2" pageNumber="469">Euvira</emphasis>
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</taxonomicName>
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, but is distinguishable by the short anterior angles, contrasting with the broadly rounded angles, and in some cases indistinct, of
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<taxonomicName box="[365,537,223,249]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis box="[365,441,223,248]" italics="true" pageId="2" pageNumber="469">Euvira</emphasis>
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species.
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</taxonomicName>
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Further, members of
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<taxonomicName authorityName="Sharp" authorityYear="1883" box="[778,854,223,248]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[778,854,223,248]" italics="true" pageId="2" pageNumber="469">Euvira</emphasis>
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</taxonomicName>
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have straight and distinctive posterior angles on the head, while it is indistinct in
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<taxonomicName authorityName="Pires-Silva" authorityYear="2023" box="[475,596,259,285]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[475,596,259,285]" italics="true" pageId="2" pageNumber="469">Euryachen</emphasis>
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</taxonomicName>
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due to the presence of the broad neck.
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</paragraph>
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<paragraph blockId="2.[151,1436,151,501]" pageId="2" pageNumber="469">
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The tergite VIII in
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<taxonomicName authorityName="Pires-Silva" authorityYear="2023" box="[414,535,295,321]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[414,535,295,321]" italics="true" pageId="2" pageNumber="469">Euryachen</emphasis>
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</taxonomicName>
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is a transverse to subquadrate plate in both males and females, contrasting with the modified tergite VIII from males of
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<taxonomicName authorityName="Erichson" authorityYear="1837" box="[600,690,331,357]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[600,690,331,357]" italics="true" pageId="2" pageNumber="469">Placusa</emphasis>
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</taxonomicName>
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(above mentioned); in species of
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<taxonomicName box="[1071,1202,331,357]" class="Insecta" family="Staphylinidae" genus="Placukorna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[1071,1202,331,357]" italics="true" pageId="2" pageNumber="469">Placukorna</emphasis>
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</taxonomicName>
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it is like a conspicuously curved horn. In
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<taxonomicName authorityName="Sharp" authorityYear="1883" box="[398,474,367,392]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[398,474,367,392]" italics="true" pageId="2" pageNumber="469">Euvira</emphasis>
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</taxonomicName>
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, the tergite VIII is broader in males rather than females.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="469" type="etymology">
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<paragraph blockId="2.[151,1436,151,501]" pageId="2" pageNumber="469">
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<emphasis bold="true" box="[199,328,403,429]" pageId="2" pageNumber="469">Etymology</emphasis>
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. The generic name is a junction of two Greek words, ευρύς (
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<emphasis box="[1024,1083,403,429]" italics="true" pageId="2" pageNumber="469">evrýs</emphasis>
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, wide) and the nominative singular noun αὐχήν (
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<emphasis box="[363,443,440,467]" italics="true" pageId="2" pageNumber="469">aukhḗn</emphasis>
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, neck). Thus,
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<taxonomicName authorityName="Pires-Silva" authorityYear="2023" box="[602,723,439,465]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
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<emphasis box="[602,723,439,465]" italics="true" pageId="2" pageNumber="469">Euryachen</emphasis>
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</taxonomicName>
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means “wide neck” in allusion to the large neck distinctively in the tribe. Gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="469" type="key">
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<paragraph blockId="2.[151,837,546,609]" box="[151,640,546,573]" pageId="2" pageNumber="469">
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<heading bold="true" box="[151,640,546,573]" fontSize="11" level="1" pageId="2" pageNumber="469" reason="1">
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<emphasis bold="true" box="[151,640,546,573]" pageId="2" pageNumber="469">
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Updated Key to the genera of
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[526,640,546,572]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[151,837,546,609]" box="[151,837,583,609]" pageId="2" pageNumber="469">
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(modified from
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<bibRefCitation author="Santiago-Jimenez, Q. J. & Santiago-Navarro, R." box="[330,829,583,609]" pageId="2" pageNumber="469" pagination="45 - 57" refId="ref5944" refString="Santiago-Jimenez, Q. J. & Santiago-Navarro, R. (2016) A new genus and species of Placusini from a high mountain in Mexico. ZooKeys, 640, 45 - 57. https: // doi. org / 10.3897 / zookeys. 640.10911" type="journal article" year="2016">Santiago-Jiménez & Santiago-Navarro 2016</bibRefCitation>
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)
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</paragraph>
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<key pageId="2" pageNumber="469">
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<keyStep pageId="2" pageNumber="469">
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<paragraph blockId="2.[151,1437,645,1078]" box="[151,1436,645,667]" pageId="2" pageNumber="469">
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<keyLead box="[151,1436,645,667]" pageId="2" pageNumber="469">1 Head with distinct neck................................................................................ 2</keyLead>
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</paragraph>
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<paragraph blockId="2.[151,1437,645,1078]" box="[151,1436,674,696]" pageId="2" pageNumber="469">
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<keyLead box="[151,1436,674,696]" pageId="2" pageNumber="469">- Head with an indistinct neck............................................................................ 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="2" pageNumber="469">
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<paragraph blockId="2.[151,1437,645,1078]" pageId="2" pageNumber="469">
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<keyLead pageId="2" pageNumber="469">
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2 Neck broad, almost same width of head; head spherical with indistinct posterior angles; anterior angles of head shortly rounded; males with thicker spine-like bristles on the central region of tergite
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<collectionCode box="[881,898,733,754]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="469" type="Museum">V</collectionCode>
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..........................
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<emphasis bold="true" box="[1231,1436,732,754]" pageId="2" pageNumber="469">
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<taxonomicName authority="Pires-Silva, 2023" authorityName="Pires-Silva" authorityYear="2023" box="[1231,1340,732,754]" class="Insecta" family="Staphylinidae" genus="Euryachen" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[1231,1340,732,754]" italics="true" pageId="2" pageNumber="469">Euryachen</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1346,1436,733,754]" pageId="2" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph blockId="2.[151,1437,645,1078]" pageId="2" pageNumber="469">
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<keyLead pageId="2" pageNumber="469">
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- Neck narrower than head. Head quadrate with strong posterior angles; anterior angles of pronotum broadly rounded; tergite
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<collectionCode box="[1419,1436,763,784]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="469" type="Museum">V</collectionCode>
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of males without spine-like bristles....................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="2" pageNumber="469">
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<paragraph blockId="2.[151,1437,645,1078]" pageId="2" pageNumber="469">
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<keyLead pageId="2" pageNumber="469">
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3 Antennomeres 4–10 distinctively transverse; spermatheca rounded, stem somewhat triangular (
|
||
<bibRefCitation author="Pace, R." box="[1137,1240,821,842]" pageId="2" pageNumber="469" pagination="225 - 398" refId="ref5843" refString="Pace, R. (2008) New records of Aleocharinae from Ecuador and Peru, with the description of new species, new subgenera and new genera (Coleoptera, Staphylinidae). Biodiversity of South America, I. Memoirs on Biodiversity, 1, 225 - 398." type="journal article" year="2008">Pace 2008</bibRefCitation>
|
||
, fig 92)..................................................................................................
|
||
<taxonomicName authority="Pace" authorityName="Pace" authorityYear="2008" box="[1313,1436,851,872]" class="Insecta" family="Staphylinidae" genus="Kirtusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1313,1385,851,872]" italics="true" pageId="2" pageNumber="469">Kirtusa</emphasis>
|
||
Pace
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,645,1078]" box="[151,1436,879,902]" pageId="2" pageNumber="469">
|
||
<keyLead box="[151,1436,879,902]" pageId="2" pageNumber="469">
|
||
- Antennomeres 5–10 distinctively transverse; spermatheca not as above................................
|
||
<taxonomicName authority="Sharp" authorityName="Sharp" authorityYear="1883" box="[1309,1436,880,902]" class="Insecta" family="Staphylinidae" genus="Euvira" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1309,1374,880,901]" italics="true" pageId="2" pageNumber="469">Euvira</emphasis>
|
||
Sharp
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="469">
|
||
<paragraph blockId="2.[151,1437,645,1078]" box="[151,1436,909,931]" pageId="2" pageNumber="469">
|
||
<keyLead box="[151,1436,909,931]" pageId="2" pageNumber="469">
|
||
4 Head rounded or transversally sub-orbicular; tergite VIII transverse, not modified...
|
||
<taxonomicName authority="Pace" authorityName="Pace" authorityYear="2013" box="[1061,1257,909,930]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1061,1206,909,930]" italics="true" pageId="2" pageNumber="469">Speiraphallusa</emphasis>
|
||
Pace
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Pace, R." box="[1270,1374,909,930]" pageId="2" pageNumber="469" pagination="33 - 63" refId="ref5894" refString="Pace, R. (2013) New distributional data, new species and three new genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26, 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
|
||
; fig 1)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,645,1078]" box="[151,1436,938,960]" pageId="2" pageNumber="469">
|
||
<keyLead box="[151,1436,938,960]" pageId="2" pageNumber="469">- Head transverse; tergite VIII not transverse, modified, bearing spines or as a curved horn............................ 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="469">
|
||
<paragraph blockId="2.[151,1437,645,1078]" pageId="2" pageNumber="469">
|
||
<keyLead pageId="2" pageNumber="469">
|
||
<specimenCount box="[151,275,967,990]" pageId="2" pageNumber="469" type="male">5 Male</specimenCount>
|
||
tergite VIII with a variable number of small to large spines (
|
||
<bibRefCitation author="Klimaszewski, J. & Pelletier, G. & Germain, C. & Hebert, C. & Humble, L. M. & Winchester, N. N." box="[857,1106,968,990]" pageId="2" pageNumber="469" pagination="1 - 47" refId="ref5764" refString="Klimaszewski, J., Pelletier, G., Germain, C., Hebert, C., Humble, L. M. & Winchester, N. N. (2001) Diversity of Placusa (Coleoptera: Staphylinidae, Aleocharinae) in Canada, with descriptions of two new species. The Canadian Entomologist, 133, 1 - 47. https: // doi. org / 10.4039 / Ent 1331 - 1" type="journal article" year="2001">
|
||
Klimaszewski
|
||
<emphasis box="[1001,1043,968,989]" italics="true" pageId="2" pageNumber="469">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
; figs 33–49).......................................................................................................
|
||
<taxonomicName authority="Erichson" authorityName="Erichson" authorityYear="1837" box="[1268,1436,997,1019]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1268,1345,997,1018]" italics="true" pageId="2" pageNumber="469">Placusa</emphasis>
|
||
Erichson
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,645,1078]" pageId="2" pageNumber="469">
|
||
<keyLead pageId="2" pageNumber="469">
|
||
- Male and female tergite VIII without spines, forming a conspicuously curved horn (
|
||
<bibRefCitation author="Santiago-Jimenez, Q. J. & Santiago-Navarro, R." pageId="2" pageNumber="469" pagination="45 - 57" refId="ref5944" refString="Santiago-Jimenez, Q. J. & Santiago-Navarro, R. (2016) A new genus and species of Placusini from a high mountain in Mexico. ZooKeys, 640, 45 - 57. https: // doi. org / 10.3897 / zookeys. 640.10911" type="journal article" year="2016">Santiago-Jiménez & Santiago-Navarro 2016</bibRefCitation>
|
||
; figs 2, 11)...............................................................
|
||
<taxonomicName authority="Santiago-Jimenez" box="[1148,1437,1056,1078]" class="Insecta" family="Staphylinidae" genus="Placukorna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1148,1260,1056,1077]" italics="true" pageId="2" pageNumber="469">Placukorna</emphasis>
|
||
Santiago-Jiménez
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |