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<document id="A52C73138C1FFFDB5EC3F72CC7EAE390" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFD8E528531A803B664FFBF5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) hirtus" docType="treatment" docVersion="12" lastPageNumber="143" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="142" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="694742271B017451D802DE1BC3A4A958">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="37CCB8044F1DF17A43EFE2CCFA338E40">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="7CF6663B17D4DA1FF1172D812EA04366">text</mods:typeOfResource>
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<mods:title id="846F21F7BE9E27C9E5EED3A633F95AF4">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="2289D7595D18FC9F95E56538F712C64C">
<mods:date id="C852B3ED5466691886A52B531F0E94A7">2017</mods:date>
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<mods:number id="00B76C5CDF1A03EBEFEF4FC8DE4060F2">2017-06-30</mods:number>
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<mods:number id="501AB861DB39982AE224BBB421BA3FAB">67</mods:number>
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<mods:classification id="47F6E8A21BEBB64554528C2C7638EB8F">journal article</mods:classification>
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<mods:identifier id="A91A7258E685C60575DE0DA6845612E5" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
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<treatment id="03A787BAFFD8E528531A803B664FFBF5" ID-DOI="http://doi.org/10.5281/zenodo.5888469" ID-GBIF-Taxon="192498173" ID-Zenodo-Dep="5888469" LSID="urn:lsid:plazi:treatment:03A787BAFFD8E528531A803B664FFBF5" httpUri="http://treatment.plazi.org/id/03A787BAFFD8E528531A803B664FFBF5" lastPageId="26" lastPageNumber="143" pageId="25" pageNumber="142">
<subSubSection id="C3146527FFD8E52B531A803B67AAF8FA" box="[831,1185,1843,1870]" pageId="25" pageNumber="142" type="nomenclature">
<paragraph id="8BB136ACFFD8E52B531A803B67AAF8FA" blockId="25.[831,1185,1843,1903]" box="[831,1185,1843,1870]" pageId="25" pageNumber="142">
<heading id="D0F981C0FFD8E52B531A803B67AAF8FA" box="[831,1185,1843,1870]" centered="true" fontSize="12" level="2" pageId="25" pageNumber="142" reason="2">
<taxonomicName id="4C0E4D2FFFD8E52B531A803B67AAF8FA" ID-CoL="5DDCZ" authority="(KRAATZ, 1859)" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[831,1185,1843,1870]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="142" phylum="Arthropoda" rank="species" species="hirtus" subGenus="Zyras">
<emphasis id="B97AEABEFFD8E52B531A803B607EF8FA" box="[831,885,1843,1870]" italics="true" pageId="25" pageNumber="142">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD8E52B53A6803B60B2F8FA" box="[899,953,1843,1870]" italics="true" pageId="25" pageNumber="142">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD8E52B53E2803B6709F8FA" box="[967,1026,1843,1870]" italics="true" pageId="25" pageNumber="142">hirtus</emphasis>
(
<bibRefCitation id="EF9F4B5DFFD8E52B5428803B6791F8FA" author="KRAATZ, G." box="[1037,1178,1843,1870]" pageId="25" pageNumber="142" pagination="1 - 193" refId="ref57963" refString="KRAATZ, G. 1859: Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. - Archiv fur Naturgeschichte 25 (1): 1 - 193." type="journal article" year="1859">KRAATZ, 1859</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD8E52B531A80536735F8DB" box="[831,1086,1883,1903]" pageId="25" pageNumber="142" type="description">
<paragraph id="8BB136ACFFD8E52B531A80536735F8DB" blockId="25.[831,1185,1843,1903]" box="[831,1086,1883,1903]" pageId="25" pageNumber="142">
(
<figureCitation id="13352A29FFD8E52B536680536076F8DB" box="[835,893,1883,1903]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="25" pageNumber="142">Figs 21</figureCitation>
,
<figureCitation id="13352A29FFD8E52B53A680536093F8DB" box="[899,920,1883,1903]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="25" pageNumber="142">59</figureCitation>
,
<figureCitation id="13352A29FFD8E52B53BA805360BFF8DB" box="[927,948,1883,1903]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="25" pageNumber="142">95</figureCitation>
,
<figureCitation id="13352A29FFD8E52B539E8053670BF8DB" box="[955,1024,1883,1903]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="25" pageNumber="142">170–173</figureCitation>
,
<figureCitation id="13352A29FFD8E52B542280536732F8DB" box="[1031,1081,1883,1903]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="25" pageNumber="142">Map 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD8E52B533F80836777F815" box="[794,1148,1931,1953]" pageId="25" pageNumber="142" type="reference_group">
<paragraph id="8BB136ACFFD8E52B533F80836777F815" blockId="25.[794,1148,1931,1953]" box="[794,1148,1931,1953]" pageId="25" pageNumber="142">
<treatmentCitationGroup id="AB1E1182FFD8E52B533F80836777F815" box="[794,1148,1931,1953]" pageId="25" pageNumber="142">
<treatmentCitation id="0AAF10BDFFD8E52B533F80836773F815" author="KRAATZ, G." box="[794,1144,1931,1953]" page="25" pageId="25" pageNumber="142" year="1859">
<taxonomicName id="4C0E4D2FFFD8E52B533F80836773F815" authority="KRAATZ, 1859: 25" authorityName="KRAATZ" authorityPageNumber="25" authorityYear="1859" box="[794,1144,1931,1953]" class="Insecta" family="Staphylinidae" genus="Myrmedonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="142" phylum="Arthropoda" rank="species" species="hirta">
<emphasis id="B97AEABEFFD8E52B533F808360CCF814" box="[794,967,1931,1952]" italics="true" pageId="25" pageNumber="142">Myrmedonia hirta</emphasis>
<bibRefCitation id="EF9F4B5DFFD8E52B53E880836773F815" author="KRAATZ, G." box="[973,1144,1931,1953]" pageId="25" pageNumber="142" pagination="1 - 193" refId="ref57963" refString="KRAATZ, G. 1859: Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. - Archiv fur Naturgeschichte 25 (1): 1 - 193." type="journal article" year="1859">KRAATZ, 1859: 25</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD8E52B533F80C167CCF795" pageId="25" pageNumber="142" type="discussion">
<paragraph id="8BB136ACFFD8E52B533F80C167CCF795" blockId="25.[794,1399,1993,2081]" pageId="25" pageNumber="142">
<emphasis id="B97AEABEFFD8E52B533F80C16082F855" bold="true" box="[794,905,1993,2017]" pageId="25" pageNumber="142">Comment</emphasis>
: The female
<typeStatus id="54B5880EFFD8E52B540B80C16787F855" box="[1070,1164,1993,2017]" pageId="25" pageNumber="142" type="holotype">holotype</typeStatus>
from “Ceylon” was revised by
<bibRefCitation id="EF9F4B5DFFD8E52B53B880E16745F7B5" author="ASSING, V." box="[925,1102,2025,2049]" pageId="25" pageNumber="142" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a)</bibRefCitation>
, who reported additional records from
<collectingCountry id="F319763CFFD8E52B538F8F016704F795" box="[938,1039,2057,2081]" name="Sri Lanka" pageId="25" pageNumber="142">Sri Lanka</collectingCountry>
and South
<collectingCountry id="F319763CFFD8E52B54AD8F0167C8F795" box="[1160,1219,2057,2081]" name="India" pageId="25" pageNumber="142">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDBE528509887E16280FEF5" pageId="26" pageNumber="143" type="materials_examined">
<paragraph id="8BB136ACFFDBE528509887E16280FEF5" blockId="26.[189,794,233,321]" pageId="26" pageNumber="143">
<emphasis id="B97AEABEFFDBE528509887E16297FEB5" bold="true" box="[189,412,233,257]" pageId="26" pageNumber="143">Material examined</emphasis>
:
<materialsCitation id="3B663CF1FFDBE528519C87E1628CFEF5" ID-GBIF-Occurrence="3450162336" collectingDate="1972-11-19" collectionCode="MHNG" collectorName="Besuchet &amp; Lobl &amp; Mussard" country="India" elevation="800" location="Madras" pageId="26" pageNumber="143" specimenCount="1">
<emphasis id="B97AEABEFFDBE528519C87E162F8FEB5" bold="true" box="[441,499,233,257]" pageId="26" pageNumber="143">
<collectingCountry id="F319763CFFDBE528519C87E162F8FEB5" box="[441,499,233,257]" name="India" pageId="26" pageNumber="143">India</collectingCountry>
</emphasis>
:
<specimenCount id="9D08FD25FFDBE528523487E1613DFEB5" box="[529,566,233,257]" pageId="26" pageNumber="143">1 ♂</specimenCount>
,
<location id="8ED16077FFDBE528527187E161ACFEB5" LSID="urn:lsid:plazi:treatment:03A787BAFFD8E528531A803B664FFBF5:8ED16077FFDBE528527187E161ACFEB5" box="[596,679,233,257]" country="India" name="Madras" pageId="26" pageNumber="143">Madras</location>
,
<locationDeviation id="9A032F8EFFDBE52852E787E16215FE95" location="Coonoor" pageId="26" pageNumber="143">
<quantity id="4CF69B49FFDBE52852E787E1600FFEB5" box="[706,772,233,257]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="143" unit="km" value="15.0">15 km</quantity>
E Coonoor
</locationDeviation>
,
<quantity id="4CF69B49FFDBE528511686016273FE95" box="[307,376,265,289]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="143" unit="m" value="800.0">
<elevation id="0023D19FFFDBE528511686016273FE95" box="[307,376,265,289]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="143" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFB0106CFFDBE52851A886016109FE95" box="[397,514,265,289]" pageId="26" pageNumber="143" value="1972-11-19">
<collectingDate id="EFF4E984FFDBE52851A886016109FE95" box="[397,514,265,289]" pageId="26" pageNumber="143" value="1972-11-19">19.XI.1972</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFDBE528526B860161BBFE95" box="[590,688,265,289]" pageId="26" pageNumber="143">Besuchet</collectorName>
,
<collectorName id="26FB537AFFDBE52852E0860161FEFE95" box="[709,757,265,289]" pageId="26" pageNumber="143">Löbl</collectorName>
&amp;
<collectorName id="26FB537AFFDBE528509886216212FEF5" box="[189,281,297,321]" pageId="26" pageNumber="143">Mussard</collectorName>
(
<collectionCode id="ED1FAE69FFDBE528510386216274FEF5" box="[294,383,297,321]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="26" pageNumber="143" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDBE5285098866160ACFC15" pageId="26" pageNumber="143" type="description">
<paragraph id="8BB136ACFFDBE52850988661628EFDD5" blockId="26.[189,794,361,2050]" pageId="26" pageNumber="143">
<emphasis id="B97AEABEFFDBE52850988661625DFE35" bold="true" box="[189,342,361,385]" pageId="26" pageNumber="143">Redescription</emphasis>
: Rather large species: body length
<quantity id="4CF69B49FFDBE528509886816235FE15" box="[189,318,393,417]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.8" metricValueMin="6.4" pageId="26" pageNumber="143" unit="mm" value="7.1" valueMax="7.8" valueMin="6.4">6.4–7.8 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFDBE528523C86816191FE15" box="[537,666,393,417]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="26" pageNumber="143" unit="mm" value="3.1" valueMax="3.5" valueMin="2.7">2.7–3.5 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFDBE52850E186A16218FE75" box="[196,275,425,449]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="26" pageNumber="143">Figs 21</figureCitation>
,
<figureCitation id="13352A29FFDBE528510686A16234FE75" box="[291,319,425,449]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="26" pageNumber="143">59</figureCitation>
,
<figureCitation id="13352A29FFDBE528516B86A16260FE75" box="[334,363,425,449]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="26" pageNumber="143">95</figureCitation>
): body pale-reddish to reddish-brown, rarely with abdominal segment VI somewhat darker; legs yellowish; antennae brown to blackish, with the basal 1–2 and the apical 1–4 antennomeres yellowish to pale reddish; maxillary palpi dark-yellowish with palpomere IV paler yellowish.
</paragraph>
<paragraph id="8BB136ACFFDBE52850988561622FFC75" blockId="26.[189,794,361,2050]" pageId="26" pageNumber="143">
Head (
<figureCitation id="13352A29FFDBE528512B85616252FD35" box="[270,345,617,641]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="26" pageNumber="143">Fig. 59</figureCitation>
) distinctly transverse, median portion extensively impunctate; punctures in lateral portions moderately sparse and moderately coarse; pubescence long and sub-erect to erect. Eyes much longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFDBE52852E285E16019FCB5" box="[711,786,745,769]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="26" pageNumber="143">Fig. 21</figureCitation>
)
<quantity id="4CF69B49FFDBE528509884016237FC95" box="[189,316,777,801]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="26" pageNumber="143" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
long and distinctly incrassate; antennomeres IV weakly oblong to weakly transverse, V as long as broad or weakly transverse, VI–X of increasing width and increasingly transverse, 2.0–2.5 times as broad as long, and XI short, shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFDBE528509884C262FFFB15" blockId="26.[189,794,361,2050]" pageId="26" pageNumber="143">
Pronotum (
<figureCitation id="13352A29FFDBE528511C84C1628FFC55" box="[313,388,969,993]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="26" pageNumber="143">Fig. 59</figureCitation>
) of variable shape, 1.09–1.21 times as broad as long and approximately 1.3 times as broad as head, broadest in anterior half; punctation rather dense, very shallow, and regularly distributed; midline narrowly to broadly impunctate; pubescence suberect and rather long, pale on disc, brown and longer along lateral margins and in antero-lateral portion.
</paragraph>
<paragraph id="8BB136ACFFDBE528509883A162DAFAD5" blockId="26.[189,794,361,2050]" pageId="26" pageNumber="143">
Elytra (
<figureCitation id="13352A29FFDBE528513783A16257FB75" box="[274,348,1193,1217]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="26" pageNumber="143">Fig. 59</figureCitation>
) 0.85–0.90 times as long as pronotum; punctation rather fine, very dense, and shallow, regularly distributed; whole disc with dense and long sub-erect brown pubescence. Hind wings fully developed. Metatarsomere I approximately as long as, or shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFDBE528509882616286F835" blockId="26.[189,794,361,2050]" pageId="26" pageNumber="143">
Abdomen (
<figureCitation id="13352A29FFDBE528511082616289FA35" box="[309,386,1385,1409]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="26" pageNumber="143">Fig. 95</figureCitation>
) approximately as broad as elytra, with rather deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of moderately coarse non-setiferous punctures; tergites III–V with moderately dense and fine setiferous punctation on disc and at posterior margins; tergite VI with a transverse row or band of non-setiferous punctures anteriorly and with moderately dense setiferous punctation on disc and at posterior margin; tergite VII with more or less extensive non-setiferous punctation anteriorly and with moderately sparse to moderately dense setiferous punctation not arranged in distinct rows in posterior portion, posterior margin with palisade fringe; tergite VIII with rather dense and long yellowish setae in posterior half, posterior margin shallowly concave in the middle; all sternites with long brown pubescence.
</paragraph>
<paragraph id="8BB136ACFFDBE5285098808260F7FE95" blockId="26.[189,794,361,2050]" lastBlockId="26.[831,1436,233,353]" pageId="26" pageNumber="143">
♂: posterior margin of sternite VIII convex, in the middle more or less distinctly truncate; median lobe of aedeagus (
<figureCitation id="13352A29FFDBE52850E180C1625FF855" box="[196,340,1993,2017]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="26" pageNumber="143">Figs 170–173</figureCitation>
) 0.88–1.0 mm long; ventral process subapically angled and apically very acute in lateral view; paramere
<quantity id="4CF69B49FFDBE528538A87E16743FEB5" box="[943,1096,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="26" pageNumber="143" unit="mm" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.80 mm</quantity>
long and with very short and broad apical lobe.
</paragraph>
<paragraph id="8BB136ACFFDBE528531A862260B2FED5" blockId="26.[831,1436,233,353]" pageId="26" pageNumber="143">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
<paragraph id="8BB136ACFFDBE528531A8681665FFD35" blockId="26.[831,1437,393,641]" pageId="26" pageNumber="143">
<emphasis id="B97AEABEFFDBE528531A8681673CFE15" bold="true" box="[831,1079,393,417]" pageId="26" pageNumber="143">Intraspecific variation</emphasis>
: The examined specimens from South
<collectingCountry id="F319763CFFDBE52853A686A160B0FE75" box="[899,955,425,449]" name="India" pageId="26" pageNumber="143">India</collectingCountry>
differ from those from
<collectingCountry id="F319763CFFDBE528549486A1661DFE75" box="[1201,1302,425,449]" name="Sri Lanka" pageId="26" pageNumber="143">Sri Lanka</collectingCountry>
by distinctly darker antennae with less transverse antennomeres V–X, a less transverse pronotum, and slightly larger body size. The median lobe of the aedeagus, however, is of nearly identical shape. Therefore, and in view of the rather few specimens examined so far, these differences are interpreted as intra- rather than interspecific variation.
</paragraph>
<paragraph id="8BB136ACFFDBE528531A85A160ACFC15" blockId="26.[831,1437,681,929]" pageId="26" pageNumber="143">
<emphasis id="B97AEABEFFDBE528531A85A1671AFD75" bold="true" box="[831,1041,681,705]" pageId="26" pageNumber="143">Comparative notes</emphasis>
:
<taxonomicName id="4C0E4D2FFFDBE528543885A26796FD75" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1053,1181,681,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="143" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFDBE528543885A26796FD75" box="[1053,1181,681,705]" italics="true" pageId="26" pageNumber="143">Zyras hirtus</emphasis>
</taxonomicName>
is readily distinguished from other species of the
<taxonomicName id="4C0E4D2FFFDBE528547285C267B9FD55" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1111,1202,713,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="143" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFDBE528547285C267B9FD55" box="[1111,1202,713,737]" italics="true" pageId="26" pageNumber="143">Z. hirtus</emphasis>
</taxonomicName>
group recorded from
<collectingCountry id="F319763CFFDBE528531A85E1607CFCB5" box="[831,887,745,769]" name="India" pageId="26" pageNumber="143">India</collectingCountry>
and
<collectingCountry id="F319763CFFDBE528538F85E16705FCB5" box="[938,1038,745,769]" name="Sri Lanka" pageId="26" pageNumber="143">Sri Lanka</collectingCountry>
by the more or less uniformly reddish coloration of the body, by distinctly transverse preapical antennomeres, by the punctation pattern of the abdomen (particularly the rather dense setiferous punctation on all tergites), and by the shape of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFDBE528531A84C1664FFBF5" pageId="26" pageNumber="143" type="distribution">
<paragraph id="8BB136ACFFDBE528531A84C1664FFBF5" blockId="26.[831,1437,969,1090]" pageId="26" pageNumber="143">
<emphasis id="B97AEABEFFDBE528531A84C167AEFC55" bold="true" box="[831,1189,969,993]" pageId="26" pageNumber="143">Distribution and natural history</emphasis>
: The confirmed distribution is confined to
<collectingCountry id="F319763CFFDBE528540684E16782FBB5" box="[1059,1161,1001,1025]" name="Sri Lanka" pageId="26" pageNumber="143">Sri Lanka</collectingCountry>
and South
<collectingCountry id="F319763CFFDBE528552784E16631FBB5" box="[1282,1338,1001,1025]" name="India" pageId="26" pageNumber="143">India</collectingCountry>
(
<figureCitation id="13352A29FFDBE528556D84E16684FBB5" box="[1352,1423,1001,1025]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="26" pageNumber="143">Map 7</figureCitation>
). For additional records see
<bibRefCitation id="EF9F4B5DFFDBE52854788301660CFB95" author="ASSING, V." box="[1117,1287,1033,1057]" pageId="26" pageNumber="143" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a)</bibRefCitation>
. The altitudes of the known localities range from
<quantity id="4CF69B49FFDBE528549683216634FBF5" box="[1203,1343,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="26" pageNumber="143" unit="m" value="600.0" valueMax="900.0" valueMin="300.0">
300 to
<specimenCount id="9D08FD25FFDBE52854DE83216634FBF5" box="[1275,1343,1065,1089]" pageId="26" pageNumber="143" type="generic">900 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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