158 lines
18 KiB
XML
158 lines
18 KiB
XML
<document id="9C4B9A1A1FE0AE7FC92C78672AC01C87" ID-CLB-Dataset="46304" ID-DOI="10.5281/zenodo.205904" ID-GBIF-Dataset="62376e09-f30b-4fb0-a635-8f8f43e008fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="205904" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391967158" checkinUser="plazi" docAuthor="Sasakawa, Mitsuhiro" docDate="2011" docId="03A787A1FFD7FFB9FF5EFC31EB3919FF" docLanguage="en" docName="zt03038p028.pdf" docOrigin="Zootaxa 3038" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heteromeringia melanoprotoma Sasakawa, 2011, sp. nov." docType="treatment" docVersion="14" lastPageNumber="21" masterDocId="FF9EFFD9FFC4FFADFFC9FFB5EF5D1878" masterDocTitle="Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="20" updateTime="1698298315006" updateUser="ExternalLinkService">
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<mods:mods id="7F536C5A626A7E42D2717FD53A7B08A8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="88F1334962EEB8B771271A9C0DD61100">
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<mods:title id="0E54C4E2CE78736FC7308AD3F695884B">Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F08CA84C88D55C7AF33AD73013B78EDC" type="personal">
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<mods:roleTerm id="7836EE3FF6B6D6839AC29A0FBF1B5A7F">Author</mods:roleTerm>
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<mods:namePart id="9E6A4DB712401C8C36C2FA8FFD068C2A">Sasakawa, Mitsuhiro</mods:namePart>
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<mods:typeOfResource id="879BE5B0FBAF8D0BD2B4F5C2639B4791">text</mods:typeOfResource>
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<mods:title id="56EBAF7279B83261E68C3916315BE1FA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D34D9805236E3D8EA67F9A5E0613CFB8">
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<mods:date id="1F09F7D8A8D1B31E00971620F216B72A">2011</mods:date>
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<mods:detail id="AC7E06E8145CE10B0553C20B76F52D8B" type="volume">
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<mods:number id="4C05EC0E72916CAC5E5992891315D697">3038</mods:number>
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<mods:start id="56378D101A60733E42B08068A51593C4">1</mods:start>
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<mods:classification id="48345453B2DF32F94B8206696EBE71FC">journal article</mods:classification>
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<mods:identifier id="0F4A12050FB03E8D5FBB5816F74EC717" type="CLB-Dataset">46304</mods:identifier>
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<mods:identifier id="17CBC7299288935AA9DD4086062C3B3B" type="DOI">10.5281/zenodo.205904</mods:identifier>
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<mods:identifier id="6363755C019FC003044D29278ECD2C46" type="GBIF-Dataset">62376e09-f30b-4fb0-a635-8f8f43e008fd</mods:identifier>
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<mods:identifier id="7C5ADEE077C70708D8A4BDC59D2CE5C0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C4AB18AA800180697102646064F0812E" type="Zenodo-Dep">205904</mods:identifier>
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<treatment id="03A787A1FFD7FFB9FF5EFC31EB3919FF" ID-DOI="http://doi.org/10.5281/zenodo.3509707" ID-GBIF-Taxon="119416971" ID-Zenodo-Dep="3509707" LSID="urn:lsid:plazi:treatment:03A787A1FFD7FFB9FF5EFC31EB3919FF" httpUri="http://treatment.plazi.org/id/03A787A1FFD7FFB9FF5EFC31EB3919FF" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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<subSubSection id="C314653CFFD7FFBEFF5EFC31EE7A1BB8" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BB136B7FFD7FFBEFF5EFC31ED311BE5" blockId="19.[151,620,900,960]" box="[151,620,900,925]" pageId="19" pageNumber="20">
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<heading id="D0F981DBFFD7FFBEFF5EFC31ED311BE5" bold="true" box="[151,620,900,925]" fontSize="10" level="2" pageId="19" pageNumber="20" reason="2">
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<emphasis id="B97AEAA5FFD7FFBEFF5EFC31ED311BE5" bold="true" box="[151,620,900,925]" pageId="19" pageNumber="20">
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<taxonomicName id="4C0E4D34FFD7FFBEFF5EFC31ED561BE5" ID-CoL="8CBWQ" box="[151,523,900,925]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanoprotoma" status="sp. nov.">
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<emphasis id="B97AEAA5FFD7FFBEFF5EFC31ED561BE5" bold="true" box="[151,523,900,925]" italics="true" pageId="19" pageNumber="20">Heteromeringia melanoprotoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957DEFFD7FFBEFDDBFC30ED311BE5" box="[530,620,901,925]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF5EFC1DEE7A1BB8" blockId="19.[151,620,900,960]" box="[151,295,936,960]" pageId="19" pageNumber="20">
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(
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<figureCitation id="13352A32FFD7FFBEFF56FC1DEE421BB8" box="[159,287,936,960]" captionStart="FIGURES 25 – 35" captionStartId="18.[151,264,1953,1976]" captionTargetBox="[229,1365,198,1916]" captionTargetId="figure@18.[227,1378,195,1926]" captionTargetPageId="18" captionText="FIGURES 25 – 35. Male genitalia of Heteromeringia abatanensis sp. nov. (25 – 27, paratype), H. melanoprotoma sp. nov. (28 – 31, holotype), and H. paraphalloides sp. nov. (32 – 35, holotype). See Figs 1 – 3 for explanation of parts; 31, 35, ejaculatory apodeme, lateral view; 27, HP, hypophallus. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205909/files/figure.png" pageId="19" pageNumber="20">Figs 28–31</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD7FFBEFF5EFC45EC1A1C54" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="8BB136B7FFD7FFBEFF5EFC45EC1A1C54" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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<emphasis id="B97AEAA5FFD7FFBEFF5EFC45EE1B1C70" bold="true" box="[151,326,1007,1032]" pageId="19" pageNumber="20">
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<typeStatus id="54B58815FFD7FFBEFF5EFC45EF8D1C70" box="[151,208,1008,1032]" pageId="19" pageNumber="20">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54B58815FFD7FFBEFE84FC45EEE81C70" box="[333,437,1008,1032]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED1FAE72FFD7FFBEFDCAFC45ED0E1C70" LSID="urn:lsid:biocol.org:col:34697" box="[515,595,1008,1032]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="19" pageNumber="20">BPBM</collectionCode>
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17249),
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<collectingCountry id="F3197627FFD7FFBEFD71FC44EDAC1C70" box="[696,753,1009,1032]" name="Laos" pageId="19" pageNumber="20">Laos</collectingCountry>
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, Vientiane Prov., Ban Van Eue,
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<date id="FFB01077FFD7FFBEFB95FC45EBAA1C70" box="[1116,1271,1008,1032]" pageId="19" pageNumber="20" value="1968-12-31">31. XII. 1968</date>
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, native collector.
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<typeStatus id="54B58815FFD7FFBEFF0BFBA0EE691C55" box="[194,308,1045,1069]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
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: 2Ƥ, same data as
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<typeStatus id="54B58815FFD7FFBEFDCDFBA1ED3B1C54" box="[516,614,1044,1068]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
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except
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<date id="FFB01077FFD7FFBEFD74FBA1EC1F1C54" box="[701,834,1044,1068]" pageId="19" pageNumber="20" value="1966-02-15">15. II. 1966</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD7FFBEFF0EFB82ECBF1CE0" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="8BB136B7FFD7FFBEFF0EFB82ECBF1CE0" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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<emphasis id="B97AEAA5FFD7FFBEFF0EFB82EE1C1C28" bold="true" box="[199,321,1079,1104]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
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Largely black, but parafacialia, gena, and scutellum brown in male; first antennal flagellomere yellowish, narrowly brown on apex; dc three; fore tarsomeres flattened; surstylus with four or five spinules on inner tip; cerci united; hypophallus short or slender, distiphallus pointed apically.
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</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD7FFBEFF0EFB11ED711FAF" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BB136B7FFD7FFBEFF0EFB11EBDD1DEC" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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<emphasis id="B97AEAA5FFD7FFBEFF0EFB11EE041CC5" bold="true" box="[199,345,1188,1213]" pageId="19" pageNumber="20">Description.</emphasis>
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MALE. Head black except for dark brown orbit, parafacialia, face, and gena; frontalia grayish pruinose, parafrontalia shining; face blackened laterally, strongly shining; peristome black. Antenna brownish yellow, first flagellomere narrowly brown along apical margin, arista brownish yellow with base yellow; palpus black. Thorax black; mesoscutum slightly grayish dusted, weakly shining; pleura strongly shining, anatergite, katatergite, katepimeron, and mediotergite dark brown; scutellum brown. Wing hyaline, slightly infuscated on anteroapical 1/3; calypter yellow, with margin and fringe pale brown; halter yellow. Legs yellow, with all coxae and fore femur slightly brown-tinged, fore tibia and tarsus black. Abdomen shining black. Bristles black.
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</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF0EFA14ED771E78" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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Frons as wide as eye, slightly converging ventrally; ventral part of parafrontalia and parafacialia linearly projecting above eye margin in profile; all or subequal in length; oc 2/3 length of poc; gena 1/5 as high as eye. Antenna similar to that of
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<taxonomicName id="4C0E4D34FFD7FFBEFE91FA5CED7E1E78" box="[344,547,1512,1536]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="paraphalloides">
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<emphasis id="B97AEAA5FFD7FFBEFE91FA5CED7E1E78" box="[344,547,1512,1536]" italics="true" pageId="19" pageNumber="20">H. paraphalloides</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF0EF9B8EC251ECF" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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Mesoscutum with three dc, first dc 1/2 length of second dc, latter 3/4 length of third dc and situated just before level of sa-base, eight rows of acr but five or six sparse rows of longer acr behind level of second dc; scutellum with bsc 1/4 length of asc. Wing
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<quantity id="4CF69B52FFD7FFBEFDC1F9E1ED031E14" box="[520,606,1620,1644]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="20" unit="mm" value="2.8">2.8 mm</quantity>
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long; three costal sections in proportion of 63: 10: 7, r-m beyond middle level of cell dm (14: 10), M1 ratio 3.8-5.0, ultimate section of CuA1 3/5 as long as penultimate section. Fore tarsomeres flattened laterally, almost as wide as distal width of tibia.
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</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF0EF976EB1F1F13" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">
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Surstylus (
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<figureCitation id="13352A32FFD7FFBEFE89F976EE991EA3" box="[320,452,1731,1755]" captionStart="FIGURES 25 – 35" captionStartId="18.[151,264,1953,1976]" captionTargetBox="[229,1365,198,1916]" captionTargetId="figure@18.[227,1378,195,1926]" captionTargetPageId="18" captionText="FIGURES 25 – 35. Male genitalia of Heteromeringia abatanensis sp. nov. (25 – 27, paratype), H. melanoprotoma sp. nov. (28 – 31, holotype), and H. paraphalloides sp. nov. (32 – 35, holotype). See Figs 1 – 3 for explanation of parts; 31, 35, ejaculatory apodeme, lateral view; 27, HP, hypophallus. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205909/files/figure.png" pageId="19" pageNumber="20">Figs 28, 29</figureCitation>
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) narrow, shorter than epandrial height, with four or five spinules on inner apex; cerci separated ventrally; pregonite with three or four strong apical setae, postgonite with one strong apical seta and 4–6 setulae. Phallapodeme 190 µm long; phallus (
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<figureCitation id="13352A32FFD7FFBEFD63F8BEEDA21F5B" box="[682,767,1803,1827]" captionStart="FIGURES 25 – 35" captionStartId="18.[151,264,1953,1976]" captionTargetBox="[229,1365,198,1916]" captionTargetId="figure@18.[227,1378,195,1926]" captionTargetPageId="18" captionText="FIGURES 25 – 35. Male genitalia of Heteromeringia abatanensis sp. nov. (25 – 27, paratype), H. melanoprotoma sp. nov. (28 – 31, holotype), and H. paraphalloides sp. nov. (32 – 35, holotype). See Figs 1 – 3 for explanation of parts; 31, 35, ejaculatory apodeme, lateral view; 27, HP, hypophallus. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205909/files/figure.png" pageId="19" pageNumber="20">Fig. 30</figureCitation>
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) 750 µm long, epiphallus short, mesophallus broadend at middle, hypophallus distinctly short on right side and slender on left side, paraphallus simple, distiphallic tube pointed apically; ejaculatory apodeme (
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<figureCitation id="13352A32FFD7FFBEFD98F8E6EDFF1F13" box="[593,674,1875,1899]" captionStart="FIGURES 25 – 35" captionStartId="18.[151,264,1953,1976]" captionTargetBox="[229,1365,198,1916]" captionTargetId="figure@18.[227,1378,195,1926]" captionTargetPageId="18" captionText="FIGURES 25 – 35. Male genitalia of Heteromeringia abatanensis sp. nov. (25 – 27, paratype), H. melanoprotoma sp. nov. (28 – 31, holotype), and H. paraphalloides sp. nov. (32 – 35, holotype). See Figs 1 – 3 for explanation of parts; 31, 35, ejaculatory apodeme, lateral view; 27, HP, hypophallus. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205909/files/figure.png" pageId="19" pageNumber="20">Fig. 31</figureCitation>
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) narrow, 230 µm long, 90 µm broad.
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</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF0EF8C2EEE81FF7" blockId="19.[151,1437,1007,2007]" box="[199,437,1911,1935]" pageId="19" pageNumber="20">Body length 3.0 mm.</paragraph>
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<paragraph id="8BB136B7FFD7FFBEFF0EF82EED711FAF" blockId="19.[151,1437,1007,2007]" pageId="19" pageNumber="20">FEMALE. Similar to male, but head and scutellum entirely black, fore femur brown on distal 1/3, and fore tarsomeres broader than those of male.</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD0FFB9FF0EFF22EEF818D7" box="[199,421,151,176]" pageId="20" pageNumber="21" type="distribution">
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<paragraph id="8BB136B7FFD0FFB9FF0EFF22EEF818D7" blockId="20.[151,1436,151,392]" box="[199,421,151,176]" pageId="20" pageNumber="21">
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<emphasis id="B97AEAA5FFD0FFB9FF0EFF22EE3D18C8" bold="true" box="[199,352,151,176]" pageId="20" pageNumber="21">Distribution.</emphasis>
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<collectingCountry id="F3197627FFD0FFB9FEAEFF2DEEFC18D7" box="[359,417,152,175]" name="Laos" pageId="20" pageNumber="21">Laos</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD0FFB9FF0EFF09EB0D191C" pageId="20" pageNumber="21" type="discussion">
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<paragraph id="8BB136B7FFD0FFB9FF0EFF09EB0D191C" blockId="20.[151,1436,151,392]" pageId="20" pageNumber="21">
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<emphasis id="B97AEAA5FFD0FFB9FF0EFF09EE6618AC" bold="true" box="[199,315,188,212]" pageId="20" pageNumber="21">Remarks.</emphasis>
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This species is easily separated from
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<taxonomicName id="4C0E4D34FFD0FFB9FCC8FF08EB4018AC" box="[769,1053,188,212]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaena">
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<emphasis id="B97AEAA5FFD0FFB9FCC8FF08EB4018AC" box="[769,1053,188,212]" italics="true" pageId="20" pageNumber="21">Heteromeringia melaena</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0E4D34FFD0FFB9FB95FF08EA7118AC" box="[1116,1324,188,212]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="paraphalloides" status="sp. nov.">
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<emphasis id="B97AEAA5FFD0FFB9FB95FF08EB2D18AC" box="[1116,1136,189,212]" italics="true" pageId="20" pageNumber="21">H</emphasis>
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.
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<emphasis id="B97AEAA5FFD0FFB9FB4BFF09EA7118AC" box="[1154,1324,188,212]" italics="true" pageId="20" pageNumber="21">paraphalloides</emphasis>
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</taxonomicName>
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<emphasis id="B97AEAA5FFD0FFB9FAFEFF09EAC818AC" bold="true" box="[1335,1429,188,212]" pageId="20" pageNumber="21">
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<taxonomicNameLabel id="A24957DEFFD0FFB9FAFEFF09EAC818AC" box="[1335,1429,188,212]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, which also have dark, compressed fore tarsomeres, by its yellow fore femur, male cerci separated ventrally, and presence of an apical seta on the postgonite. In
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<emphasis id="B97AEAA5FFD0FFB9FD6EFEB0EC721964" box="[679,815,260,284]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C0E4D34FFD0FFB9FD6EFEB0EC761964" box="[679,811,260,284]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaena">H. melaena</taxonomicName>
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,
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</emphasis>
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the yellow fore femur is brown-ringed before the apex, the male cerci are entirely separated, and the postgonite is only setulose on the apex; in
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<taxonomicName id="4C0E4D34FFD0FFB9FBA5FE9CEA6A1938" box="[1132,1335,296,320]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="paraphalloides">
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<emphasis id="B97AEAA5FFD0FFB9FBA5FE9CEBDD1938" box="[1132,1152,297,320]" italics="true" pageId="20" pageNumber="21">H</emphasis>
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.
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<emphasis id="B97AEAA5FFD0FFB9FB47FE9DEA6A1938" box="[1166,1335,296,320]" italics="true" pageId="20" pageNumber="21">paraphalloides</emphasis>
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</taxonomicName>
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, the fore femur is black, the cerci are completely fused, and the postgonite is setulose apically.
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</paragraph>
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</subSubSection>
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<subSubSection id="C314653CFFD0FFB9FF0EFEC5EB3919FF" box="[199,1124,367,392]" pageId="20" pageNumber="21" type="etymology">
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<paragraph id="8BB136B7FFD0FFB9FF0EFEC5EB3919FF" blockId="20.[151,1436,151,392]" box="[199,1124,367,392]" pageId="20" pageNumber="21">
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<emphasis id="B97AEAA5FFD0FFB9FF0EFEC5EE1B19F0" bold="true" box="[199,326,368,392]" pageId="20" pageNumber="21">Etymology</emphasis>
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. The specific name refers to the black face (Greek:
|
||
<emphasis id="B97AEAA5FFD0FFB9FC42FEC5ECB519F0" box="[907,1000,368,392]" italics="true" pageId="20" pageNumber="21">melanos</emphasis>
|
||
+
|
||
<emphasis id="B97AEAA5FFD0FFB9FC31FEC4EB0919F0" box="[1016,1108,369,392]" italics="true" pageId="20" pageNumber="21">protome</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |