315 lines
37 KiB
XML
315 lines
37 KiB
XML
<document id="F3C6E1EA0E50564B0639266733E9F40C" ID-CLB-Dataset="46781" ID-DOI="10.11646/zootaxa.4078.1.29" ID-GBIF-Dataset="c59a6000-9262-47b9-a672-6fd625f486da" ID-ISSN="1175-5326" ID-Zenodo-Dep="262814" ID-ZooBank="8377D13F-7C34-4CD6-9749-D5B9F804F25F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460362571044" checkinUser="plazi" docAuthor="Holzenthal, Ralph W., Blahnik, Roger J. & Calor, Adolfo R." docDate="2016" docId="03A6460CFFB1FFF1FF7FBC89FC22A6DF" docLanguage="en" docName="zootaxa.4078.1.29.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helicopsyche (Feropsyche) angeloi Holzenthal, Blahnik & Calor, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="346" masterDocId="FF9F3E74FFB3FFF3FFE8BD3FFFE3A166" masterDocTitle="Three new species of Helicopsyche von Siebold (Trichoptera: Helicopsychidae) from Brazil" masterLastPageNumber="353" masterPageNumber="344" pageNumber="346" updateTime="1698294712413" updateUser="ExternalLinkService">
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<mods:mods id="379A2340364423E455E86874D56C8EC3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C06B083132E59AABBEE04E9AE80E2A62">
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<mods:title id="0D8CB93F5D657AB94827FE90BFA581D0">Three new species of Helicopsyche von Siebold (Trichoptera: Helicopsychidae) from Brazil</mods:title>
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<mods:name id="50FC14CC90C111AC4FB1356AAE0EF10C" type="personal">
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<mods:namePart id="075364F4AAD70DA52ECE86999BCA7EDB">Holzenthal, Ralph W.</mods:namePart>
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<mods:namePart id="4604EE804688EC96AFE35C717C5A17AF">Blahnik, Roger J.</mods:namePart>
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<mods:name id="BE6BECABB5C2CCDD9D6D7FED91C0E86E" type="personal">
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<mods:roleTerm id="66AA44CBDB996A3EC1268E6F924CB06A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1D19C8D88D4EC275E42B4B5DED850F2E">Calor, Adolfo R.</mods:namePart>
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<mods:typeOfResource id="2570E1FFD3DCE480A20076F96BC96825">text</mods:typeOfResource>
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<mods:title id="66474533A4309BDA6C9E5D3AF90A56F2">Zootaxa</mods:title>
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<mods:part id="977F5166DA9FEB63B2352CCBFE4031F9">
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<mods:date id="F975B729C7E8A36F8A43A74AC780CB51">2016</mods:date>
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<mods:detail id="8E9426C83D166FDB97E53A21744AE509" type="volume">
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<mods:number id="979458218D0F6F6790269EB39B827BD0">4078</mods:number>
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<mods:identifier id="2FE1FA1673911317F728114290B68266" type="DOI">10.11646/zootaxa.4078.1.29</mods:identifier>
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<treatment id="03A6460CFFB1FFF1FF7FBC89FC22A6DF" ID-DOI="http://doi.org/10.5281/zenodo.6059363" ID-GBIF-Taxon="119410073" ID-Zenodo-Dep="6059363" LSID="urn:lsid:plazi:treatment:03A6460CFFB1FFF1FF7FBC89FC22A6DF" httpUri="http://treatment.plazi.org/id/03A6460CFFB1FFF1FF7FBC89FC22A6DF" lastPageNumber="346" pageId="2" pageNumber="346">
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<subSubSection id="C315A491FFB1FFF1FF7FBC89FEA6A097" pageId="2" pageNumber="346" type="nomenclature">
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<paragraph id="8BB0F71AFFB1FFF1FF7FBC89FD46A0B6" blockId="2.[151,677,438,498]" box="[151,677,438,464]" pageId="2" pageNumber="346">
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<heading id="D0F84076FFB1FFF1FF7FBC89FD46A0B6" bold="true" box="[151,677,438,464]" fontSize="11" level="1" pageId="2" pageNumber="346" reason="1">
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<emphasis id="B97B2B08FFB1FFF1FF7FBC89FD46A0B6" bold="true" box="[151,677,438,464]" pageId="2" pageNumber="346">
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<taxonomicName id="4C0F8C99FFB1FFF1FF7FBC89FDA3A0B6" box="[151,576,438,464]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov." subGenus="Feropsyche">
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<emphasis id="B97B2B08FFB1FFF1FF7FBC89FEDBA0B6" bold="true" box="[151,312,438,464]" italics="true" pageId="2" pageNumber="346">Helicopsyche</emphasis>
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||
(
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||
<emphasis id="B97B2B08FFB1FFF1FEA1BC89FE36A0B6" bold="true" box="[329,469,438,464]" italics="true" pageId="2" pageNumber="346">Feropsyche</emphasis>
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)
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<emphasis id="B97B2B08FFB1FFF1FE0EBC89FDA3A0B6" bold="true" box="[486,576,438,464]" italics="true" pageId="2" pageNumber="346">angeloi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2489673FFB1FFF1FDAFBC89FD46A0B6" box="[583,677,438,464]" pageId="2" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BB0F71AFFB1FFF1FF7FBCE6FEA6A097" blockId="2.[151,677,438,498]" box="[151,325,473,498]" pageId="2" pageNumber="346">
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(
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<figureCitation id="1334EB9FFFB1FFF1FF76BCE6FEE7A097" box="[158,260,473,498]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Figures 1</figureCitation>
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A–E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C315A491FFB1FFF1FF7FBF1EFC0BA218" pageId="2" pageNumber="346" type="materials_examined">
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||
<paragraph id="8BB0F71AFFB1FFF1FF7FBF1EFC0BA218" blockId="2.[151,1437,545,1977]" pageId="2" pageNumber="346">
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<emphasis id="B97B2B08FFB1FFF1FF7FBF1EFC92A35C" bold="true" box="[151,881,545,570]" pageId="2" pageNumber="346">
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Material examined.
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<typeStatus id="54B449B8FFB1FFF1FE63BF1EFE1BA35C" box="[395,504,545,570]" pageId="2" pageNumber="346" type="holotype">Holotype</typeStatus>
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male.
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<collectingCountry id="F318B78AFFB1FFF1FDA2BF1EFD54A35C" box="[586,695,545,570]" name="Brazil" pageId="2" pageNumber="346">BRAZIL</collectingCountry>
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. Minas Gerais:
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||
</emphasis>
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Córrego das Águas Pretas & tribs., ca.
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<quantity id="4CF75AFFFFB1FFF1FAD3BF1EFA67A35C" box="[1339,1412,545,570]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="346" unit="km" value="15.0">15 km</quantity>
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S Aiuruoca,
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<geoCoordinate id="EE3B91DDFFB1FFF1FEFCBF79FE40A338" box="[276,419,582,606]" direction="south" orientation="latitude" pageId="2" pageNumber="346" precision="1" value="-22.061733">22°03.704’S</geoCoordinate>
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,
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||
<geoCoordinate id="EE3B91DDFFB1FFF1FE5DBF79FDADA338" box="[437,590,582,606]" direction="west" orientation="longitude" pageId="2" pageNumber="346" precision="1" value="-44.63735">44°38.241’W</geoCoordinate>
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||
, el.
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<quantity id="4CF75AFFFFB1FFF1FD60BF7AFD05A33B" box="[648,742,581,606]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="2" pageNumber="346" unit="m" value="1386.0">1386 m</quantity>
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||
,
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||
<date id="FFB1D1DAFFB1FFF1FD10BF7AFC91A338" box="[760,882,581,606]" pageId="2" pageNumber="346" value="2001-11-21">21.xi.2001</date>
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||
, Holzenthal, Paprocki, Blahnik and Neto leg.,
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||
<collectionCode id="ED1E6FDFFFB1FFF1FF7FBF56FF00A3E4" box="[151,227,617,642]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="2" pageNumber="346">UMSP</collectionCode>
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||
000082582 (pinned) (
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||
<collectionCode id="ED1E6FDFFFB1FFF1FE0BBF55FDD1A3E4" box="[483,562,618,642]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="346">MZSP</collectionCode>
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||
).
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||
<emphasis id="B97B2B08FFB1FFF1FDA4BF56FC68A3E4" bold="true" box="[588,907,617,642]" pageId="2" pageNumber="346">
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||
<typeStatus id="54B449B8FFB1FFF1FDA4BF56FD2BA3E4" box="[588,712,617,642]" pageId="2" pageNumber="346" type="paratype">Paratypes</typeStatus>
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||
: Minas Gerais:
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||
</emphasis>
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||
same data as
|
||
<typeStatus id="54B449B8FFB1FFF1FBA9BF56FB45A3E4" box="[1089,1190,617,642]" pageId="2" pageNumber="346" type="holotype">holotype</typeStatus>
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||
,
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||
<specimenCount id="9D093C93FFB1FFF1FB5FBF55FAF4A3E4" box="[1207,1303,617,642]" pageId="2" pageNumber="346" type="male">2 males</specimenCount>
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,
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||
<specimenCount id="9D093C93FFB1FFF1FAC1BF55FA7FA3E4" box="[1321,1436,617,642]" pageId="2" pageNumber="346" type="female">2 females</specimenCount>
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||
(pinned) (
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||
<collectionCode id="ED1E6FDFFFB1FFF1FEEFBFB1FEB4A3C0" box="[263,343,654,678]" pageId="2" pageNumber="346">UFBA</collectionCode>
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),
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||
<specimenCount id="9D093C93FFB1FFF1FE84BFB1FE24A3C0" box="[364,455,653,678]" pageId="2" pageNumber="346" type="male">7 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D093C93FFB1FFF1FE3DBFB1FDA0A3C0" box="[469,579,653,678]" pageId="2" pageNumber="346" type="female">7 females</specimenCount>
|
||
(pinned) (
|
||
<collectionCode id="ED1E6FDFFFB1FFF1FD54BFB1FCEDA3C0" box="[700,782,654,678]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="2" pageNumber="346">UMSP</collectionCode>
|
||
); Rio Tanque, ca.
|
||
<quantity id="4CF75AFFFFB1FFF1FC08BFB1FBCAA3C0" box="[992,1065,653,678]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="346" unit="km" value="12.0">12 km</quantity>
|
||
(rd) from Ipoema,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FAE0BFB1FA7BA3C0" box="[1288,1432,654,678]" direction="south" orientation="latitude" pageId="2" pageNumber="346" precision="1" value="-19.5368">19°32.208’S</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE3B91DDFFB1FFF1FF7FBF8EFECCA3AC" box="[151,303,689,714]" direction="west" orientation="longitude" pageId="2" pageNumber="346" precision="1" value="-43.447968">43°26.878’W</geoCoordinate>
|
||
, el.
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||
<quantity id="4CF75AFFFFB1FFF1FEB7BF8EFE4AA3AF" box="[351,425,689,714]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="346" unit="m" value="750.0">750 m</quantity>
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||
,
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||
<date id="FFB1D1DAFFB1FFF1FE5FBF8EFDCFA3AC" box="[439,556,689,714]" pageId="2" pageNumber="346" value="1998-05-16">16.V.1998</date>
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||
, Holzenthal and Paprocki leg.,
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||
<specimenCount id="9D093C93FFB1FFF1FC79BF8DFC3DA3AC" box="[913,990,689,714]" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
|
||
(alcohol) (
|
||
<collectionCode id="ED1E6FDFFFB1FFF1FBB4BF8DFB4DA3AC" box="[1116,1198,690,714]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="2" pageNumber="346">UMSP</collectionCode>
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||
); Parque Estadual de São Gonçalo do Rio Preto, Córrego das Eguas,
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||
<geoCoordinate id="EE3B91DDFFB1FFF1FD32BFEAFC89A388" box="[730,874,725,750]" direction="south" orientation="latitude" pageId="2" pageNumber="346" precision="1" value="-18.145267">18°08.716’S</geoCoordinate>
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||
,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FC93BFEAFBF7A388" box="[891,1044,725,750]" direction="west" orientation="longitude" pageId="2" pageNumber="346" precision="1" value="-43.36928">43°22.157’W</geoCoordinate>
|
||
, el.
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||
<quantity id="4CF75AFFFFB1FFF1FBA5BFE9FB7FA38B" box="[1101,1180,726,750]" metricMagnitude="2" metricUnit="m" metricValue="8.91" pageId="2" pageNumber="346" unit="m" value="891.0">891 m</quantity>
|
||
,
|
||
<date id="FFB1D1DAFFB1FFF1FB46BFE9FAC3A388" box="[1198,1312,726,750]" pageId="2" pageNumber="346" value="2000-10-14">14.x.2000</date>
|
||
, Paprocki, Amarante and Isaac leg.,
|
||
<specimenCount id="9D093C93FFB1FFF1FE21BFC5FDFAA274" box="[457,537,761,786]" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
|
||
(alcohol) (
|
||
<collectionCode id="ED1E6FDFFFB1FFF1FD77BFC5FD0DA274" box="[671,750,762,786]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="346">MZSP</collectionCode>
|
||
);
|
||
<emphasis id="B97B2B08FFB1FFF1FCEFBFC6FC6EA274" bold="true" box="[775,909,761,786]" pageId="2" pageNumber="346">São Paulo:</emphasis>
|
||
Estação Biológica de Boracéia, Rio Coruja,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FF7FBE21FEFAA250" box="[151,281,798,822]" direction="south" orientation="latitude" pageId="2" pageNumber="346" precision="9" value="-23.668333">23°40.10’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FECEBE22FE52A250" box="[294,433,797,822]" direction="west" orientation="longitude" pageId="2" pageNumber="346" precision="9" value="-45.899334">45°53.96’W</geoCoordinate>
|
||
, el.
|
||
<quantity id="4CF75AFFFFB1FFF1FE0BBE22FDCCA253" box="[483,559,797,822]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="346" unit="m" value="850.0">850 m</quantity>
|
||
,
|
||
<date id="FFB1D1DAFFB1FFF1FDD6BE22FD55A250" box="[574,694,797,822]" pageId="2" pageNumber="346" value="1998-04-18">18.iv.1998</date>
|
||
, Holzenthal, Melo and Froehlich leg.,
|
||
<specimenCount id="9D093C93FFB1FFF1FB9DBE21FB20A250" box="[1141,1219,797,822]" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
|
||
(pinned) (
|
||
<collectionCode id="ED1E6FDFFFB1FFF1FAD5BE21FA6CA250" box="[1341,1423,798,822]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="2" pageNumber="346">UMSP</collectionCode>
|
||
);
|
||
<emphasis id="B97B2B08FFB1FFF1FF7FBE7EFEB0A23C" bold="true" box="[151,339,833,858]" pageId="2" pageNumber="346">Rio de Janeiro:</emphasis>
|
||
Rio Macacu (2nd order) on RJ 116, km 62,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FCB2BE7DFC09A23C" box="[858,1002,834,858]" direction="south" orientation="latitude" pageId="2" pageNumber="346" precision="1" value="-22.386683">22°23.201’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B91DDFFB1FFF1FC1FBE7EFB6CA23C" box="[1015,1167,833,858]" direction="west" orientation="longitude" pageId="2" pageNumber="346" precision="1" value="-42.56575">42°33.945’W</geoCoordinate>
|
||
, el.
|
||
<quantity id="4CF75AFFFFB1FFF1FB2ABE7DFAEDA23F" box="[1218,1294,834,858]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="346" unit="m" value="840.0">840 m</quantity>
|
||
,
|
||
<date id="FFB1D1DAFFB1FFF1FAF4BE7EFA74A23C" box="[1308,1431,833,858]" pageId="2" pageNumber="346" value="1996-03-17">17.iii.1996</date>
|
||
, Holzenthal, Rochetti and Oliveira leg.,
|
||
<specimenCount id="9D093C93FFB1FFF1FDA7BE59FD7EA218" box="[591,669,869,894]" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D093C93FFB1FFF1FD40BE5AFCF6A218" box="[680,789,869,894]" pageId="2" pageNumber="346" type="female">6 females</specimenCount>
|
||
(pinned) (
|
||
<collectionCode id="ED1E6FDFFFB1FFF1FC61BE59FC38A218" box="[905,987,870,894]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="2" pageNumber="346">UMSP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C315A491FFB1FFF1FF2FBEB6FDBFA5A4" pageId="2" pageNumber="346" type="diagnosis">
|
||
<paragraph id="8BB0F71AFFB1FFF1FF2FBEB6FDBFA5A4" blockId="2.[151,1437,545,1977]" pageId="2" pageNumber="346">
|
||
<emphasis id="B97B2B08FFB1FFF1FF2FBEB6FEA2A2C4" bold="true" box="[199,321,905,930]" pageId="2" pageNumber="346">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FEA6BEB6FDA5A2C7" box="[334,582,905,929]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FEA6BEB6FDA5A2C7" box="[334,582,905,929]" italics="true" pageId="2" pageNumber="346">Helicopsyche angeloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FDBABEB6FD51A2C4" box="[594,690,905,930]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FDBABEB6FD51A2C4" bold="true" box="[594,690,905,930]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FCB4BEB5FA93A2C4" authority="Silva, Santos & Nessimian, 2014" authorityName="Silva, Santos & Nessimian" authorityYear="2014" box="[860,1392,905,930]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="timbira">
|
||
<emphasis id="B97B2B08FFB1FFF1FCB4BEB5FC93A2C7" box="[860,880,906,929]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FC6CBEB6FC34A2C7" box="[900,983,905,929]" italics="true" pageId="2" pageNumber="346">timbira</emphasis>
|
||
<bibRefCitation id="EF9E8AEBFFB1FFF1FC0BBEB6FA93A2C4" author="Silva" box="[995,1392,905,930]" pageId="2" pageNumber="346" refString="Silva, A. L. R. da, Santos, A. P. M. & Nessimian, J. L. (2014) Helicopsyche (Feropsyche) timbira sp. nov. (Trichoptera: Helicopsychidae), a new species from southeastern Brazil. Zootaxa, 3847 (3), 446 - 448. http: // dx. doi. org / 10.11646 / zootaxa. 3847.3.9" type="journal article" year="2014">Silva, Santos & Nessimian, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FA69BEB5FDCDA2A0" authority="Johanson & Malm, 2006" authorityName="Johanson & Malm" authorityYear="2006" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="cipoensis">
|
||
<emphasis id="B97B2B08FFB1FFF1FA69BEB5FA76A2C7" box="[1409,1429,906,929]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FF7FBE91FF1CA2A3" box="[151,255,942,965]" italics="true" pageId="2" pageNumber="346">cipoensis</emphasis>
|
||
<bibRefCitation id="EF9E8AEBFFB1FFF1FEE2BE92FDCDA2A0" author="Johanson" box="[266,558,941,966]" pageId="2" pageNumber="346" refString="Johanson, K. A. & Malm, T. (2006) Seven new Helicopsyche (Feropsyche) Johanson, 2002 from the Neotropical region and Nearctic Mexico (Insecta: Trichoptera: Helicopsychidae). Zootaxa, 1208, 1 - 24." type="journal article" year="2006">Johanson & Malm, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FD99BE91FD39A2A3" box="[625,730,942,965]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="guara" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FD99BE91FD66A2A3" box="[625,645,942,965]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FD7FBE91FD39A2A3" box="[663,730,942,965]" italics="true" pageId="2" pageNumber="346">guara</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FD04BE92FCA9A2A0" box="[748,842,941,966]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FD04BE92FCA9A2A0" bold="true" box="[748,842,941,966]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, all from
|
||
<collectingCountry id="F318B78AFFB1FFF1FC2CBE92FBEFA2A0" box="[964,1036,941,966]" name="Brazil" pageId="2" pageNumber="346">Brazil</collectingCountry>
|
||
. All of these species have clavate inferior appendages, each without an acute apical projection on the posterior margin and with prominent basomesal lobes of the inferior appendages, evident in lateral view. All, except
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FC78BEC9FC16A56B" box="[912,1013,1014,1037]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="guara" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FC78BEC9FC47A56B" box="[912,932,1014,1037]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FC5ABEC9FC16A56B" box="[946,1013,1014,1037]" italics="true" pageId="2" pageNumber="346">guara</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FC14BECAFBB6A568" box="[1020,1109,1013,1038]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FC14BECAFBB6A568" bold="true" box="[1020,1109,1013,1038]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, also have nearly linear rows of setae on tergum X. The overall shapes of the inferior appendages are most similar between
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FB2AB925FADAA557" box="[1218,1337,1049,1073]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FB2AB925FB35A557" box="[1218,1238,1050,1073]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FB0DB926FADAA557" box="[1253,1337,1049,1073]" italics="true" pageId="2" pageNumber="346">angeloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FAA9B926FA7FA554" box="[1345,1436,1049,1074]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FAA9B926FA7FA554" bold="true" box="[1345,1436,1049,1074]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FF2FB901FED8A533" box="[199,315,1085,1109]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="timbira">
|
||
<emphasis id="B97B2B08FFB1FFF1FF2FB901FF38A533" box="[199,219,1086,1109]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FF01B902FED8A533" box="[233,315,1085,1109]" italics="true" pageId="2" pageNumber="346">timbira</emphasis>
|
||
</taxonomicName>
|
||
in being narrow in lateral view. However, in
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FCD4B901FC50A533" box="[828,947,1085,1109]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FCD4B901FCB3A533" box="[828,848,1086,1109]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FCB7B902FC50A533" box="[863,947,1085,1109]" italics="true" pageId="2" pageNumber="346">angeloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FC53B902FBF6A530" box="[955,1045,1085,1110]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FC53B902FBF6A530" bold="true" box="[955,1045,1085,1110]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the inferior appendages bear very prominent spine-like setae on their apicomesal face and mesally at midlength. These prominent setae are not present on the inferior appendages in any of the other species. Also, the apex of segment X is slightly cleft in
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FA69B9B9FF09A5A7" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="timbira">
|
||
<emphasis id="B97B2B08FFB1FFF1FA69B9B9FA76A5FB" box="[1409,1429,1158,1181]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FF7FB996FF09A5A7" box="[151,234,1193,1217]" italics="true" pageId="2" pageNumber="346">timbira</emphasis>
|
||
</taxonomicName>
|
||
, but entire in
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FE6DB995FE1FA5A7" box="[389,508,1193,1217]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov.">
|
||
<emphasis id="B97B2B08FFB1FFF1FE6DB995FE7AA5A7" box="[389,409,1194,1217]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FE40B996FE1FA5A7" box="[424,508,1193,1217]" italics="true" pageId="2" pageNumber="346">angeloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2489673FFB1FFF1FDEBB996FDBFA5A4" box="[515,604,1193,1218]" pageId="2" pageNumber="346" rank="species">
|
||
<emphasis id="B97B2B08FFB1FFF1FDEBB996FDBFA5A4" bold="true" box="[515,604,1193,1218]" pageId="2" pageNumber="346">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C315A491FFB1FFF1FF2FB9F2FA60A62C" pageId="2" pageNumber="346" type="description">
|
||
<paragraph id="8BB0F71AFFB1FFF1FF2FB9F2FA60A62C" blockId="2.[151,1437,545,1977]" pageId="2" pageNumber="346">
|
||
<emphasis id="B97B2B08FFB1FFF1FF2FB9F2FEBAA580" bold="true" box="[199,345,1229,1254]" pageId="2" pageNumber="346">Description.</emphasis>
|
||
<emphasis id="B97B2B08FFB1FFF1FE8BB9F2FE7EA583" box="[355,413,1229,1253]" italics="true" pageId="2" pageNumber="346">Male</emphasis>
|
||
. Overall color (pinned) nearly uniformly medium brown; dorsum of head with whitish setae, especially between antennae; antennae light brown; palps dark brown. Tibial spur formula 2, 4, 4. Forewing: length 4.5–5.0 mm (n = 10); covered with fine, short, brown setae, anal margin with whitish setae. Abdominal sternum VI process long, almost as long as segment VI, tubular along its length, oriented posteroventrally, nearly straight in lateral view, with microtrichiae along length, apex tapering, with lamellae apicoventrally (
|
||
<figureCitation id="1334EB9FFFB1FFF1FADDB862FA9BA410" box="[1333,1400,1373,1398]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
E).
|
||
<emphasis id="B97B2B08FFB1FFF1FF7FB8BEFEE2A4FF" box="[151,257,1409,1433]" italics="true" pageId="2" pageNumber="346">Genitalia</emphasis>
|
||
. Segment IX long ventrally; anteromesal margin well-developed; lateral apodeme well developed, located midlaterally on segment (
|
||
<figureCitation id="1334EB9FFFB1FFF1FDC8B89AFD87A4D8" box="[544,612,1445,1470]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
A). Segment X conical along basal 2/3, tapering to apex, not divided apically; mesodorsal borders inverted Y-shaped, bearing 2 nearly linear rows of 10–12 setae along length (
|
||
<figureCitation id="1334EB9FFFB1FFF1FADFB8F6FA96A484" box="[1335,1397,1481,1506]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
B); in lateral view, ventral margin almost straight, dorsal margin parallel to ventral margin along basal half, apical half angled posteroventrally (
|
||
<figureCitation id="1334EB9FFFB1FFF1FE59BB2EFE11A74C" box="[433,498,1553,1578]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
A). Superior appendage originates dorsolaterally, short, setose, rounded (
|
||
<figureCitation id="1334EB9FFFB1FFF1FADBBB2EFA97A74C" box="[1331,1396,1553,1578]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
A). Inferior appendage clavate, with anterior margin slightly concave and posterior margin convex; apicodorsal margin undulate, with long setae along apical margin (
|
||
<figureCitation id="1334EB9FFFB1FFF1FD4EBB66FD09A714" box="[678,746,1625,1650]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Figs 1</figureCitation>
|
||
A, C) and spine-like setae apicomesally; 4 spine-like setae at midlength, each on distinct protuberance (
|
||
<figureCitation id="1334EB9FFFB1FFF1FD6DBB42FD2FA7F0" box="[645,716,1661,1686]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Figs 1</figureCitation>
|
||
A, C); basomesal lobe of inferior appendage well-developed, ovate, apically bearing 8–10 spine-like setae (
|
||
<figureCitation id="1334EB9FFFB1FFF1FD42BB9EFD13A7DC" box="[682,752,1697,1722]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Figs 1</figureCitation>
|
||
A, C); basal plate of inferior appendages nearly straight in lateral view (
|
||
<figureCitation id="1334EB9FFFB1FFF1FEC5BBFAFE8EA7B8" box="[301,365,1733,1758]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
A), narrow along its length, tapering apically (
|
||
<figureCitation id="1334EB9FFFB1FFF1FC6BBBFAFC27A7B8" box="[899,964,1733,1758]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
B). Phallus tubular, slightly down-curved along its length; phallobase narrow, slightly inflated basodorsally, narrow in the middle, apicoventral half strongly sclerotized, acuminate apically; endotheca a large, long, membranous lobe extending dorsally over endophallus; endophallic membranes extended laterally as rounded lobes; phallotremal sclerite small, quadrangular (
|
||
<figureCitation id="1334EB9FFFB1FFF1FAF4BA0EFABFA62C" box="[1308,1372,1841,1866]" captionStart="FIGURE 1" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[158,1434,205,1894]" captionTargetId="figure@3.[157,1438,195,1903]" captionTargetPageId="3" captionText="FIGURE 1. Helicopsyche (Feropsyche) angeloi sp. nov., male. A. genitalia, lateral view; B. segments IX and X, dorsal view; C. inferior appendage, ventral view; D. phallus, lateral view; E. sternum VI, lateral view. Abbreviations: IX = segment IX; X = tergum X." httpUri="https://zenodo.org/record/262815/files/figure.png" pageId="2" pageNumber="346">Fig. 1</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C315A491FFB1FFF1FF2FBA6AFB83A6F3" pageId="2" pageNumber="346" type="etymology">
|
||
<paragraph id="8BB0F71AFFB1FFF1FF2FBA6AFB83A6F3" blockId="2.[151,1437,545,1977]" pageId="2" pageNumber="346">
|
||
<emphasis id="B97B2B08FFB1FFF1FF2FBA6AFEADA608" bold="true" box="[199,334,1877,1902]" pageId="2" pageNumber="346">Etymology.</emphasis>
|
||
Named in honor of Dr. Angelo B. M. Machado, the first Brazilian researcher to describe a species of
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FF5DBA43FEAFA6F2" box="[181,332,1916,1940]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97B2B08FFB1FFF1FF5DBA43FEAFA6F2" box="[181,332,1916,1940]" italics="true" pageId="2" pageNumber="346">Helicopsyche</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F8C99FFB1FFF1FEB2BA42FD2BA6F3" authority="Machado, 1957" authorityName="Machado" authorityYear="1957" box="[346,712,1916,1941]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="planorboides">
|
||
<emphasis id="B97B2B08FFB1FFF1FEB2BA42FE8DA6F2" box="[346,366,1917,1940]" italics="true" pageId="2" pageNumber="346">H</emphasis>
|
||
.
|
||
<emphasis id="B97B2B08FFB1FFF1FE94BA43FDF3A6F2" box="[380,528,1916,1940]" italics="true" pageId="2" pageNumber="346">planorboides</emphasis>
|
||
<bibRefCitation id="EF9E8AEBFFB1FFF1FDFFBA43FD2BA6F3" author="Machado" box="[535,712,1916,1941]" pageId="2" pageNumber="346" refString="Machado, A. B. M. (1957) Helicopsyche planorboides n. sp. (Trichoptera Helicopsychidae) and its mimetic relations to planorbids (Mollusca, Pulmonata). Anais da Academia Brasileira de Ciencias, 29, 193 - 200." type="journal article" year="1957">Machado, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
, on the occasion of his 80th birthday.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C315A491FFB1FFF1FF2FBA9FFC22A6DF" box="[199,961,1952,1977]" pageId="2" pageNumber="346" type="distribution">
|
||
<paragraph id="8BB0F71AFFB1FFF1FF2FBA9FFC22A6DF" blockId="2.[151,1437,545,1977]" box="[199,961,1952,1977]" pageId="2" pageNumber="346">
|
||
<emphasis id="B97B2B08FFB1FFF1FF2FBA9FFE83A6DF" bold="true" box="[199,352,1952,1977]" pageId="2" pageNumber="346">Distribution.</emphasis>
|
||
<collectingCountry id="F318B78AFFB1FFF1FE8FBA9FFE48A6DF" box="[359,427,1952,1977]" name="Brazil" pageId="2" pageNumber="346">Brazil</collectingCountry>
|
||
(Minas Gerais, Rio de Janeiro, and São Paulo).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |