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<document id="15316D2B2CA3566ED1F615C28ABD6F4D" ID-CLB-Dataset="50742" ID-DOI="10.5281/zenodo.172563" ID-GBIF-Dataset="d325feef-2116-4c36-9571-ea93d68c89a4" ID-ISSN="1175­5326" ID-Zenodo-Dep="172563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459758407235" checkinUser="plazi" docAuthor="Cuezzo, Maria Gabriela" docDate="2006" docId="03A4A163FFCEE36DBA4041659DCA4A31" docLanguage="en" docName="zt01221p023.pdf" docOrigin="Zootaxa 1221" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Labyrinthus subplanatus Petit 1843" docType="treatment" docVersion="10" lastPageNumber="12" masterDocId="FF9DD91BFFC6E366BB48435199524E1C" masterDocTitle="On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="9" updateTime="1698218777145" updateUser="ExternalLinkService">
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<mods:title id="21300C5287ED95B48B845C6D97F7F7CE">On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)</mods:title>
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<mods:date id="1D249AD5180AEF427C96E60E31B45E52">2006</mods:date>
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<treatment id="03A4A163FFCEE36DBA4041659DCA4A31" ID-DOI="http://doi.org/10.5281/zenodo.6261407" ID-GBIF-Taxon="119344237" ID-Zenodo-Dep="6261407" LSID="urn:lsid:plazi:treatment:03A4A163FFCEE36DBA4041659DCA4A31" httpUri="http://treatment.plazi.org/id/03A4A163FFCEE36DBA4041659DCA4A31" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="C31743FEFFCEE36EBA40416598E24C6D" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BB21075FFCEE36EBA4041659B874C51" blockId="8.[264,725,563,625]" box="[264,725,563,590]" pageId="8" pageNumber="9">
<heading id="D0FAA719FFCEE36EBA4041659B874C51" bold="true" box="[264,725,563,590]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="4C0D6BF6FFCEE36EBA4041659B874C51" ID-CoL="7RSLJ" authority="Petit, 1843" authorityName="Petit" authorityYear="1843" box="[264,725,563,590]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCEE36EBA4041659B874C51" bold="true" box="[264,725,563,590]" pageId="8" pageNumber="9">
<emphasis id="B979CC67FFCEE36EBA4041659B644C52" bold="true" box="[264,566,564,590]" italics="true" pageId="8" pageNumber="9">Labyrinthus subplanatus</emphasis>
(
<bibRefCitation id="EF9C6D84FFCEE36EB90D41629B9F4C51" author="Petit" box="[581,717,563,589]" pageId="8" pageNumber="23" refString="Petit, S. (1843) Description de quatre coquilles nouvelles provenant du voyage de M. Goudot a la Nouvelle-Grenade. Revue Zoologique, Societe Cuvier, 6, 238 - 239." type="journal article" year="1843">Petit, 1843</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB21075FFCEE36EBA40410698E24C6D" blockId="8.[264,725,563,625]" box="[264,432,599,625]" pageId="8" pageNumber="9">
<figureCitation id="13360CF0FFCEE36EBA40410698E24C6D" box="[264,432,599,625]" captionStart-0="FIGURES 13 16" captionStart-1="FIGURES 17 22" captionStartId-0="9.[264,382,1622,1646]" captionStartId-1="10.[264,382,1541,1565]" captionTargetBox-0="[286,1317,290,1592]" captionTargetBox-1="[271,1315,284,1516]" captionTargetId-0="figure@9.[276,1330,281,1614]" captionTargetId-1="figure@10.[271,1315,284,1518]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." captionText-1="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri-0="https://zenodo.org/record/172567/files/figure.png" httpUri-1="https://zenodo.org/record/172568/files/figure.png" pageId="8" pageNumber="9">Figures 1322</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31743FEFFCEE36EBA4041F69BE14CA1" box="[264,691,678,702]" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BB21075FFCEE36EBA4041F69BE14CA1" blockId="8.[264,691,678,702]" box="[264,691,678,702]" pageId="8" pageNumber="9">
<taxonomicName id="4C0D6BF6FFCEE36EBA4041F69B2B4CA1" authority="Petit, 1843" authorityName="Petit" authorityYear="1843" box="[264,633,678,702]" class="Gastropoda" family="Geomitridae" genus="Caracolla" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCEE36EBA4041F698AE4CA2" box="[264,508,679,702]" italics="true" pageId="8" pageNumber="9">Caracolla subplanatus</emphasis>
<bibRefCitation id="EF9C6D84FFCEE36EB94B41F79B2B4CA1" author="Petit" box="[515,633,678,701]" pageId="8" pageNumber="23" refString="Petit, S. (1843) Description de quatre coquilles nouvelles provenant du voyage de M. Goudot a la Nouvelle-Grenade. Revue Zoologique, Societe Cuvier, 6, 238 - 239." type="journal article" year="1843">Petit, 1843</bibRefCitation>
</taxonomicName>
, 238.
</paragraph>
</subSubSection>
<subSubSection id="C31743FEFFCEE36DBA4041BF9BC84FB1" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
<paragraph id="8BB21075FFCEE36EBA4041BF9DFA4D15" blockId="8.[264,1323,750,1817]" box="[264,1192,750,777]" pageId="8" pageNumber="9">
<emphasis id="B979CC67FFCEE36EBA4041BF98F04D14" bold="true" box="[264,418,750,776]" pageId="8" pageNumber="9">Description:</emphasis>
<emphasis id="B979CC67FFCEE36EBAF941BE98BE4D14" box="[433,492,751,776]" italics="true" pageId="8" pageNumber="9">Shell</emphasis>
: Morphology as described by
<bibRefCitation id="EF9C6D84FFCEE36EB81941BE9D7D4D15" author="Solem" box="[849,1071,751,777]" pageId="8" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem (1966: 116)</bibRefCitation>
, (
<figureCitation id="13360CF0FFCEE36EBF0B41BE9DC84D15" box="[1091,1178,751,777]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BB21075FFCEE36EBA704046987F4DE5" blockId="8.[264,1323,750,1817]" pageId="8" pageNumber="9">
<emphasis id="B979CC67FFCEE36EBA70404698BB4D2C" box="[312,489,791,816]" italics="true" pageId="8" pageNumber="9">Pallial Organs</emphasis>
: Kidney long, thin, occupying about 60% of pulmonary roof length. Surface of pulmonary roof densely speckled with dark spots. Thin transversal veins abundant between kidney and rectum. Main pulmonary vein branched before reaching mantle collar. Pulmonary roof not expanded beyond top of kidney. Secondary ureter closed up to mantle collar. Ureteric interramus triangular, deeply excavated. Diaphragm thick, muscular.
</paragraph>
<paragraph id="8BB21075FFCEE36EBA70475698084905" blockId="8.[264,1323,750,1817]" pageId="8" pageNumber="9">
<emphasis id="B979CC67FFCEE36EBA7047569B7F4A3C" box="[312,557,1031,1056]" italics="true" pageId="8" pageNumber="9">Reproductive system</emphasis>
(
<figureCitation id="13360CF0FFCEE36EB90A47569B9F4A3D" box="[578,717,1031,1057]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Figs 1416</figureCitation>
): Ovotestis formed by elongated digitiform nonramified acini. Albumen gland yellowish, bean­shaped. FPSC undifferentiated. Hermaphroditic duct reflexed over itself at base of albumen gland. Albumen gland continuous with spermoviduct. Folds of uterine portion transversal with respect to longitudinal axis of spermoviduct (
<figureCitation id="13360CF0FFCEE36EBAF647F69B4F4ADD" box="[446,541,1191,1217]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
). Free oviduct short, cylindrical, naturally forming angle with vagina. Prostatic duct of spermoviduct extending into proximal portion of free oviduct. Vagina as long as penis. Proximal portion of vagina with inner thick longitudinal ridges in zigzag pattern. Distally, ridges become thinner, straight. Hooked denticles absent in entire length of vagina. Duct of bursa copulatrix long, thick, parallel to spermoviduct up to junction with albumen gland. Bursa copulatrix sac globose, reflexed over duct (
<figureCitation id="13360CF0FFCEE36EBF3E463E9D9F4B95" box="[1142,1229,1391,1417]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
). Penial complex formed by flagellum, epiphallus, penis (
<figureCitation id="13360CF0FFCEE36EB80546C69AF74BAD" box="[845,933,1431,1457]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
). Diameter of flagellum progressively decreasing towards tip. Thick internal pilaster extending between flagellum and epiphallus. Pilaster forming papilla­like structure transversally divided at junction between flagellum and epiphallus, point of insertion of vas deferens (
<figureCitation id="13360CF0FFCEE36EB894455E9D674835" box="[988,1077,1551,1577]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig. 16</figureCitation>
). Penis sheath thick, muscular overlapping distal zone of penis. Epiphallus reflexed over penis, wrapped with muscular fibres and connective tissue. Distal zone of penis thicker than proximal. Penis retractor muscle thick, inserting at proximal portion of penis close to junction with epiphallus. Inner penis wall with thick folds, deeper in proximal zone. Dorsal surface of folds with white, oval, hooked pustules, regularly distributed as shown in
<figureCitation id="13360CF0FFCEE36EBFC245869D8B48ED" box="[1162,1241,1751,1777]" captionStart="FIGURES 13 16" captionStartId="9.[264,382,1622,1646]" captionTargetBox="[286,1317,290,1592]" captionTargetId="figure@9.[276,1330,281,1614]" captionTargetPageId="9" captionText="FIGURES 13 16: Labyrinthus subplanatus. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar = 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar = 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar = 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath." httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="8" pageNumber="9">Fig.16</figureCitation>
. Verge absent.
</paragraph>
<caption id="DF7240FDFFCFE36FBA4045079A7C497C" httpUri="https://zenodo.org/record/172567/files/figure.png" pageId="9" pageNumber="10" targetBox="[286,1317,290,1592]" targetPageId="9">
<paragraph id="8BB21075FFCFE36FBA4045079A7C497C" blockId="9.[264,1325,1622,1888]" pageId="9" pageNumber="10">
<emphasis id="B979CC67FFCFE36FBA40450798984872" bold="true" box="[264,458,1622,1646]" pageId="9" pageNumber="10">FIGURES 1316</emphasis>
:
<taxonomicName id="4C0D6BF6FFCFE36FBA9245079BB74871" box="[474,741,1622,1645]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCFE36FBA9245079BB74871" box="[474,741,1622,1645]" italics="true" pageId="9" pageNumber="10">Labyrinthus subplanatus</emphasis>
</taxonomicName>
. 13: Morphology of shell aperture. 14: General morphology of reproductive system in ventral view, ovotestis not figured, scale bar= 5 mm. Abbreviations: ag: albumen gland; bc: bursa copulatrix; f: flagellum; fo: free oviduct; go: genital opening; hd: hermaphroditic duct; p: penis; s: spermoviduct; v: vagina; ps: penis sheath. 15: Detail of penial complex, dorsal view, scale bar= 5 mm. Abbreviations: f: flagellum; p: penis; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath. 16: Inner structure of penial complex, scale bar= 2 mm. Abbreviations: a: atrium; d: denticles; e: epiphallus; f: flagellum; p: penis; pi: pilaster; pr: penis retractor muscle; vd: vas deferens; ps: penis sheath.
</paragraph>
</caption>
<caption id="DF7240FDFFCCE36CBA404554989B48BB" httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="10" pageNumber="11" targetBox="[271,1315,284,1516]" targetPageId="10">
<paragraph id="8BB21075FFCCE36CBA404554989B48BB" blockId="10.[264,1323,1541,1703]" pageId="10" pageNumber="11">
<emphasis id="B979CC67FFCCE36CBA40455498954801" bold="true" box="[264,455,1541,1565]" pageId="10" pageNumber="11">FIGURES 1722</emphasis>
:
<taxonomicName id="4C0D6BF6FFCCE36CBA9D45579BB64801" box="[469,740,1542,1565]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCCE36CBA9D45579BB64801" box="[469,740,1542,1565]" italics="true" pageId="10" pageNumber="11">Labyrinthus subplanatus</emphasis>
</taxonomicName>
. 17: Dorsal view of jaw, scale bar= 100 m. 18: Detail of the jaw in dorsal view, scale bar= 100 m. 19: Central and first lateral tooth, scale bar= 10 m. 20: Lateral teeth No. 79 to the left of central tooth, scale bar= 10 m. 21: First marginal teeth in the same row of previous view, scale bar= 10 m. 22: Last marginal teeth close to radular margins, scale bar= 10 m.
</paragraph>
</caption>
<paragraph id="8BB21075FFCCE36CBA7044529C7B4971" blockId="10.[264,1323,1795,1901]" pageId="10" pageNumber="11">
<emphasis id="B979CC67FFCCE36CBA7044529B524900" box="[312,512,1795,1820]" italics="true" pageId="10" pageNumber="11">Digestive system</emphasis>
: Jaw (
<figureCitation id="13360CF0FFCCE36CB91D44529B8D4901" box="[597,735,1795,1821]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="10" pageNumber="11">Figs 1718</figureCitation>
) arched, without vertical ribs (
<figureCitation id="13360CF0FFCCE36CBF1444529DE54901" box="[1116,1207,1795,1821]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="10" pageNumber="11">Fig. 17</figureCitation>
). Surface with thin transverse striation crossed by thin, shallow perpendicular lines giving appearance of reticulated surface (
<figureCitation id="13360CF0FFCCE36CB91B44029BFE4971" box="[595,684,1875,1901]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="10" pageNumber="11">Fig. 18</figureCitation>
). Lower edge without medial protruded cutting edge.
</paragraph>
<paragraph id="8BB21075FFCDE36DBA40424D9BC84FB1" blockId="11.[264,1323,284,1070]" pageId="11" pageNumber="12">
Central tooth of radula unicuspid, triangular (
<figureCitation id="13360CF0FFCDE36DB87E424D9AC34F2A" box="[822,913,284,310]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="11" pageNumber="12">Fig. 19</figureCitation>
). Fourteen lateral teeth unicuspid (
<figureCitation id="13360CF0FFCDE36DBA58421598C54F42" box="[272,407,324,350]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="11" pageNumber="12">Figs 19, 20</figureCitation>
). Marginal teeth tricuspid (
<figureCitation id="13360CF0FFCDE36DB99A42159A7B4F42" box="[722,809,324,350]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="11" pageNumber="12">Fig. 21</figureCitation>
). Outermost marginal teeth becoming wider but always tricuspid with more pronounced mesocones (
<figureCitation id="13360CF0FFCDE36DB8EE423D9AAD4F9A" box="[934,1023,364,390]" captionStart="FIGURES 17 22" captionStartId="10.[264,382,1541,1565]" captionTargetBox="[271,1315,284,1516]" captionTargetId="figure@10.[271,1315,284,1518]" captionTargetPageId="10" captionText="FIGURES 17 22: Labyrinthus subplanatus. 17: Dorsal view of jaw, scale bar = 100 m. 18: Detail of the jaw in dorsal view, scale bar = 100 m. 19: Central and first lateral tooth, scale bar = 10 m. 20: Lateral teeth No. 7 9 to the left of central tooth, scale bar = 10 m. 21: First marginal teeth in the same row of previous view, scale bar = 10 m. 22: Last marginal teeth close to radular margins, scale bar = 10 m." httpUri="https://zenodo.org/record/172568/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
). Buccal mass oval. Rest of digestive tract as in
<emphasis id="B979CC67FFCDE36DB95242C59BC84FB1" box="[538,666,404,429]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C0D6BF6FFCDE36DB95242C59BC44FB1" box="[538,662,404,429]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="dunkeri">L. dunkeri</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31743FEFFCDE36DBA7042EA9DCF4C6A" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BB21075FFCDE36DBA7042EA9DCF4C6A" blockId="11.[264,1323,284,1070]" pageId="11" pageNumber="12">
<emphasis id="B979CC67FFCDE36DBA7042EA9BBE4FC9" bold="true" box="[312,748,443,469]" pageId="11" pageNumber="12">
Material examined:
<collectingCountry id="F31A50E5FFCDE36DB90E42EA9BBE4FC9" box="[582,748,443,469]" name="Colombia" pageId="11" pageNumber="12">COLOMBIA</collectingCountry>
</emphasis>
:
<collectionCode id="ED1C88B0FFCDE36DB84C42ED9A0B4FCA" LSID="urn:lsid:biocol.org:col:34795" box="[772,857,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
163706,
<emphasis id="B979CC67FFCDE36DB89F42ED9D924FC9" box="[983,1216,444,469]" italics="true" pageId="11" pageNumber="12">Choco Department</emphasis>
, Choco, Caño Taparral, 20 Km to the north of
<collectingCountry id="F31A50E5FFCDE36DB9F742B59A7E4FE2" box="[703,812,484,510]" name="Palestine" pageId="11" pageNumber="12">Palestina</collectingCountry>
, on San Juan River.
<date id="FFB336B5FFCDE36DBF5042B59C564FE2" box="[1048,1284,484,510]" pageId="11" pageNumber="12" value="1968-11-25">November 25, 1968</date>
. B. Malkin Coll.
<collectionCode id="ED1C88B0FFCDE36DBAE6415D9B514C3A" LSID="urn:lsid:biocol.org:col:34795" box="[430,515,524,550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
173866,
<emphasis id="B979CC67FFCDE36DB932415D9A0D4C39" box="[634,863,524,549]" italics="true" pageId="11" pageNumber="12">Choco Department</emphasis>
, San Juan River, Quebrada Docordo.
<date id="FFB336B5FFCDE36DBA40416598ED4C52" box="[264,447,564,590]" pageId="11" pageNumber="12" value="1969-06-05" valueMax="1969-06-08" valueMin="1969-06-05">June 58, 1969</date>
. B. Malkin Coll.
<collectionCode id="ED1C88B0FFCDE36DB9C341659BB24C52" LSID="urn:lsid:biocol.org:col:34795" box="[651,736,564,590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
223561,
<emphasis id="B979CC67FFCDE36DB80741659D624C51" box="[847,1072,564,589]" italics="true" pageId="11" pageNumber="12">Choco Department</emphasis>
, San Juan River, 110 Km to the north of
<collectingCountry id="F31A50E5FFCDE36DBAA1410D9B044C6A" box="[489,598,604,630]" name="Palestine" pageId="11" pageNumber="12">Palestine</collectingCountry>
.
<date id="FFB336B5FFCDE36DB92A410D9A744C6A" box="[610,806,604,630]" pageId="11" pageNumber="12" value="1971-01-20">January 20,1971</date>
. B. Malkin &amp; P. Bouchard Coll.
</paragraph>
</subSubSection>
<subSubSection id="C31743FEFFCDE36DBA7041D59DCA4A31" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BB21075FFCDE36DBA7041D59DCA4A31" blockId="11.[264,1323,284,1070]" pageId="11" pageNumber="12">
<emphasis id="B979CC67FFCDE36DBA7041D598E14C82" bold="true" box="[312,435,644,670]" pageId="11" pageNumber="12">Remarks:</emphasis>
<taxonomicName id="4C0D6BF6FFCDE36DBAF541D59BB04C81" box="[445,738,644,669]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DBAF541D59BB04C81" box="[445,738,644,669]" italics="true" pageId="11" pageNumber="12">Labyrinthus subplanatus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C0D6BF6FFCDE36DB8EA41D59ABC4C81" box="[930,1006,644,669]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DB8EA41D59ABC4C81" box="[930,1006,644,669]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
species complex that was supported as a monophyletic clade within the genus (
<bibRefCitation id="EF9C6D84FFCDE36DB83A41FD9D414CDA" author="Cuezzo" box="[882,1043,684,710]" pageId="11" pageNumber="23" refString="Cuezzo, M. G. (2003) Phylogenetic analysis of the Camaenidae with special emphasis on the American taxa (Mollusca: Stylommatophora). Zoological Journal of the Linnean Society, 138, 449 - 476." type="journal article" year="2003">Cuezzo 2003</bibRefCitation>
). The group of
<taxonomicName id="4C0D6BF6FFCDE36DBF8F41FD9C424CD9" box="[1223,1296,684,709]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DBF8F41FD9C424CD9" box="[1223,1296,684,709]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
is composed by
<taxonomicName id="4C0D6BF6FFCDE36DBAE341859B994CF2" authority="Lightfoot, 1786" authorityName="Lightfoot" authorityYear="1786" box="[427,715,724,750]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DBAE3418598A64CF1" box="[427,500,724,749]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
(Lightfoot, 1786)
</taxonomicName>
,
<taxonomicName id="4C0D6BF6FFCDE36DB99F41859AD64CF1" box="[727,900,724,749]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DB99F41859AD64CF1" box="[727,900,724,749]" italics="true" pageId="11" pageNumber="12">L. subplanatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0D6BF6FFCDE36DB8D241859DA64CF2" authority="Pilsbry, 1910" authorityName="Pilsbry" authorityYear="1910" box="[922,1268,724,750]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="marmatensis">
<emphasis id="B979CC67FFCDE36DB8D241859D1D4CF1" box="[922,1103,724,749]" italics="true" pageId="11" pageNumber="12">L. marmatensis</emphasis>
Pilsbry, 1910
</taxonomicName>
, and
<taxonomicName id="4C0D6BF6FFCDE36DBA4041AD98DA4D09" box="[264,392,764,789]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="plicatus">
<emphasis id="B979CC67FFCDE36DBA4041AD98DA4D09" box="[264,392,764,789]" italics="true" pageId="11" pageNumber="12">L. plicatus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0D6BF6FFCDE36DBAD341AD9B934D09" box="[411,705,764,789]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DBAD341AD9B934D09" box="[411,705,764,789]" italics="true" pageId="11" pageNumber="12">Labyrinthus subplanatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C0D6BF6FFCDE36DB82441AD9AE84D09" box="[876,954,764,789]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DB82441AD9AE84D09" box="[876,954,764,789]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
and both have been confused mostly because their distributional ranges are partly overlapping. However,
<taxonomicName id="4C0D6BF6FFCDE36DBFC440759D844D21" box="[1164,1238,804,829]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DBFC440759D844D21" box="[1164,1238,804,829]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0D6BF6FFCDE36DBA02401D98AA4D79" box="[330,504,844,869]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DBA02401D98AA4D79" box="[330,504,844,869]" italics="true" pageId="11" pageNumber="12">L. subplanatus</emphasis>
</taxonomicName>
mainly in having a deep supraperipheral groove and in being much larger (
<taxonomicName id="4C0D6BF6FFCDE36DBA17402598F84D91" box="[351,426,884,909]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DBA17402598F84D91" box="[351,426,884,909]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
shell diameter up to
<quantity id="4CF5BD90FFCDE36DB9E140259A534D92" box="[681,769,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="12" unit="mm" value="58.0">58 mm</quantity>
, while
<taxonomicName id="4C0D6BF6FFCDE36DB81040259D544D91" box="[856,1030,884,909]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DB81040259D544D91" box="[856,1030,884,909]" italics="true" pageId="11" pageNumber="12">L. subplanatus</emphasis>
</taxonomicName>
diameter up to
<quantity id="4CF5BD90FFCDE36DBF8D40259C4D4D92" box="[1221,1311,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="mm" value="45.0">45 mm</quantity>
).
<taxonomicName id="4C0D6BF6FFCDE36DBA4040CD9B664DA9" box="[264,564,924,949]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subplanatus">
<emphasis id="B979CC67FFCDE36DBA4040CD9B664DA9" box="[264,564,924,949]" italics="true" pageId="11" pageNumber="12">Labyrinthus subplanatus</emphasis>
</taxonomicName>
also present a periphery acutely angulated with protruding keel, and the peristome forming a peripheral notch sometimes slightly twisted (see
<bibRefCitation id="EF9C6D84FFCDE36DBFDC40959C704DC2" author="Solem" box="[1172,1314,964,990]" pageId="11" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">Solem 1966</bibRefCitation>
) which is absent in
<taxonomicName id="4C0D6BF6FFCDE36DBAAC40BD9B7C4A19" box="[484,558,1004,1029]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">
<emphasis id="B979CC67FFCDE36DBAAC40BD9B7C4A19" box="[484,558,1004,1029]" italics="true" pageId="11" pageNumber="12">L. otis</emphasis>
</taxonomicName>
. Differences in the genitalia between these two species are difficult to evaluate because of the poor descriptions presently available of
<emphasis id="B979CC67FFCDE36DBF0F47459DCA4A31" box="[1095,1176,1044,1069]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C0D6BF6FFCDE36DBF0F47459DC64A31" box="[1095,1172,1044,1069]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="otis">L. otis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>