156 lines
13 KiB
XML
156 lines
13 KiB
XML
<document ID-DOI="10.5281/zenodo.172563" ID-GBIF-Dataset="d325feef-2116-4c36-9571-ea93d68c89a4" ID-ISSN="11755326" ID-Zenodo-Dep="172563" checkinTime="1459758407235" checkinUser="plazi" docAuthor="Cuezzo, Maria Gabriela" docDate="2006" docId="03A4A163FFC9E376BA4046699AF54D7A" docLanguage="en" docName="zt01221p023.pdf" docOrigin="Zootaxa 1221" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@599a9cb2" docStyleName="zootaxa.0000.journal_article" docTitle="Isomeria Albers 1850" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="FF9DD91BFFC6E366BB48435199524E1C" masterDocTitle="On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="16" updateTime="1645725616964" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On a new species of Isomeria Beck and redescription of species of Labyrinthus Beck from South America (Gastropoda: Stylommatophora: Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuezzo, Maria Gabriela</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2006</mods:date>
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<mods:detail type="volume">
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<mods:number>1221</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.172563</mods:identifier>
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<mods:identifier type="GBIF-Dataset">d325feef-2116-4c36-9571-ea93d68c89a4</mods:identifier>
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<mods:identifier type="ISSN">11755326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">172563</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6261413" ID-GBIF-Taxon="119344230" ID-Zenodo-Dep="6261413" LSID="urn:lsid:plazi:treatment:03A4A163FFC9E376BA4046699AF54D7A" httpUri="http://treatment.plazi.org/id/03A4A163FFC9E376BA4046699AF54D7A" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection box="[264,535,1336,1362]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph blockId="15.[264,535,1336,1362]" box="[264,535,1336,1362]" pageId="15" pageNumber="16">
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<heading bold="true" box="[264,535,1336,1362]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<taxonomicName authority="Albers, 1850" authorityName="Albers" authorityYear="1850" box="[264,535,1336,1362]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis bold="true" box="[264,535,1336,1362]" pageId="15" pageNumber="16">
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<emphasis bold="true" box="[264,371,1336,1362]" italics="true" pageId="15" pageNumber="16">Isomeria</emphasis>
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Albers,
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<number box="[476,535,1336,1362]" pageId="15" pageNumber="16" value="1850.0">1850</number>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[264,891,1411,1437]" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph blockId="15.[264,1323,1411,1877]" box="[264,891,1411,1437]" pageId="15" pageNumber="16">
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<emphasis bold="true" box="[264,428,1411,1437]" pageId="15" pageNumber="16">
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<typeStatus box="[264,325,1411,1437]" pageId="15" pageNumber="16">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName authority="Koch, 1844" authorityName="Koch" authorityYear="1844" box="[436,717,1411,1437]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="Oreas">
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<emphasis box="[436,571,1412,1437]" italics="true" pageId="15" pageNumber="16">Helix oreas</emphasis>
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Koch,
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<number box="[656,717,1411,1437]" pageId="15" pageNumber="16" value="1844.0">1844</number>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="16" type="diagnosis">
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<paragraph blockId="15.[264,1323,1411,1877]" pageId="15" pageNumber="16">
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<emphasis bold="true" box="[312,442,1451,1477]" pageId="15" pageNumber="16">Diagnosis:</emphasis>
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Shell small to large (
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<quantity box="[692,821,1451,1477]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="7.5" metricValueMin="1.8" pageId="15" pageNumber="16" unit="mm" value="46.5" valueMax="75.0" valueMin="18.0">18–75 mm</quantity>
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) generally globose; shell surface sculpture with wrinkles, granulations and/or malleations; periphery convex to obtusely angulated; umbilicus usually closed or very narrow; lip thick, reflected with aperture subquadrate with few teeth and lamellae, toothless in some species; parietal callus usually thin, sometimes raised edge; parietal wall generally toothless or with small triangular teeth or short lamellae; basal lip with
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<number box="[549,593,1651,1677]" pageId="15" pageNumber="16" value="1.0" valueMax="2.0" valueMin="0.0">0–2</number>
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denticles, palatal lip having small triangular denticle in most species, rarely with large tooth marked by posterior indentation; presence of special structures such as penial appendages, epiphallic flagellum and/or stimulatorlike appendage in vaginaatrium; both vagina and penis with internal hooked denticles. Jaw smooth or with weakly developed plaques.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="distribution">
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<paragraph blockId="15.[264,1323,1411,1877]" lastBlockId="16.[264,1323,284,870]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<emphasis bold="true" box="[312,474,1851,1877]" pageId="15" pageNumber="16">Distribution:</emphasis>
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distributed from Sierra Nevada de Santa
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<collectingCountry box="[978,1048,1851,1877]" name="Malta" pageId="15" pageNumber="16">Marta</collectingCountry>
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in northern
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<collectingCountry box="[1200,1319,1851,1877]" name="Colombia" pageId="15" pageNumber="16">Colombia</collectingCountry>
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, Valle del Cauca, Cordillera Central and Occidental in
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<collectingCountry box="[952,1067,284,310]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
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south to Cajamarca department in
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<collectingCountry box="[436,490,324,350]" name="Peru" pageId="16" pageNumber="17">Peru</collectingCountry>
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(
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<bibRefCitation author="Solem" box="[506,653,324,350]" pageId="16" pageNumber="23" refString="Solem, A. (1966) The neotropical land snail genera Labyrinthus and Isomeria (Pulmonata, Camaenidae). Fieldiana, Zoology, 50, 1 - 226." type="journal article" year="1966">
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Solem
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<number box="[591,653,324,350]" pageId="16" pageNumber="17" value="1966.0">1966</number>
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</bibRefCitation>
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;
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<bibRefCitation author="Breure" box="[666,817,324,350]" pageId="16" pageNumber="22" refString="Breure, A. S. H. (1984) Survey of the land Mollusca of the Sierra Nevada de Santa Marta, Colombia. In: van der Hammen, T. & Ruiz, P. M. (Eds.), Studies on tropical Andean ecosystems. Vol 2. La Sierra Nevada de Santa Marta (Colombia). J. Cramer, Berlin / Stuttgart, pp. 487 - 500." type="journal article" year="1984">
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Breure
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<number box="[754,817,324,350]" pageId="16" pageNumber="17" value="1984.0">1984</number>
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</bibRefCitation>
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).
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<taxonomicName box="[839,943,324,349]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[839,943,324,349]" italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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is more abundant in
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<collectingCountry box="[1195,1293,324,350]" name="Ecuador" pageId="16" pageNumber="17">Ecuador</collectingCountry>
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at Esmeraldas and in amazonian localities close to Napo River. Its distributional range seems to be more restricted than that of
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<emphasis box="[651,800,404,429]" italics="true" pageId="16" pageNumber="17">
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<taxonomicName box="[651,796,404,429]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Labyrinthus</taxonomicName>
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,
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</emphasis>
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further collections will allow to define more precisely the distribution of
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<taxonomicName box="[595,699,444,469]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[595,699,444,469]" italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="discussion">
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<paragraph blockId="16.[264,1323,284,870]" pageId="16" pageNumber="17">
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<emphasis bold="true" box="[312,426,483,509]" pageId="16" pageNumber="17">Remarks</emphasis>
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:In the phylogenetic hypothesis proposed for
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<taxonomicName box="[1026,1171,484,510]" class="Gastropoda" family="Camaenidae" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
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by
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<bibRefCitation author="Cuezzo" pageId="16" pageNumber="23" refString="Cuezzo, M. G. (2003) Phylogenetic analysis of the Camaenidae with special emphasis on the American taxa (Mollusca: Stylommatophora). Zoological Journal of the Linnean Society, 138, 449 - 476." type="journal article" year="2003">
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Cuezzo (
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<number box="[273,337,524,550]" pageId="16" pageNumber="17" value="2003.0">2003</number>
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)
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</bibRefCitation>
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, the synapomorphies supporting the monophyly of
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<taxonomicName box="[958,1063,524,549]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[958,1063,524,549]" italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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were only shell characters. This is probable a consequence of the scarce anatomical information available for
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<taxonomicName box="[264,368,604,629]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[264,368,604,629]" italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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and the low number of species used in the cladistic analysis. The identified synapomorphies were:
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<number box="[489,502,644,670]" pageId="16" pageNumber="17" value="1.0">1</number>
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) aperture shape quadrangular;
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<number box="[867,882,644,670]" pageId="16" pageNumber="17" value="2.0">2</number>
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) presence of palatal teeth in the aperture and
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<number box="[386,401,684,710]" pageId="16" pageNumber="17" value="3.0">3</number>
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) umbilicus partially or entirely covered by lip. Characters shared with
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<taxonomicName box="[264,406,724,749]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[264,406,724,749]" italics="true" pageId="16" pageNumber="17">Labyrinthus</emphasis>
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</taxonomicName>
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are the presence of hooked denticles in the penis, which are smaller in
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<taxonomicName class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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; the penisepiphallus reflection usually wrapped in part by penis sheath. The free oviduct formed an angle with vagina. The jaw in
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<taxonomicName box="[804,908,804,829]" class="Gastropoda" family="Camaenidae" genus="Isomeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[804,908,804,829]" italics="true" pageId="16" pageNumber="17">Isomeria</emphasis>
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</taxonomicName>
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(as it is the case in some species of
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<taxonomicName box="[264,406,844,869]" class="Gastropoda" family="Labyrinthidae" genus="Labyrinthus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
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<emphasis box="[264,406,844,869]" italics="true" pageId="16" pageNumber="17">Labyrinthus</emphasis>
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</taxonomicName>
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) is smooth with transverse striae, not ribbed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |