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<document id="447483E9B79067B1A90B9FA40137D814" ID-CLB-Dataset="22375" ID-DOI="10.5852/ejt.2015.150" ID-GBIF-Dataset="f492ad9b-a83d-4d8e-bdf9-20ef759b75c1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3787588" ID-ZooBank="D546A676-35F8-4025-8660-F123A0BA67F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588704123207" checkinUser="carolina" docAuthor="Enghoff, Henrik &amp; Frederiksen, Sara B." docDate="2015" docId="03A487F2FFFFFFB775646A139727B3FD" docLanguage="en" docName="ejt-2015-150.pdf.imf" docOrigin="European Journal of Taxonomy 150" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor submajor Enghoff 2015, sp. nov." docType="treatment" docUuid="690C121A-D087-400D-95BC-42BCA7B224BE" docUuidSource="ZooBank" docVersion="7" lastPageNumber="10" masterDocId="FF9DFF8AFFF9FFBE775F69219518B21B" masterDocTitle="A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1698838626763" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F6F9E5BF6EB78F1205C469375C0BD45D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9A39237BE5A61D3697326D60A049A3C">
<mods:title id="8D8638BF30FBA816914A68748216D57D">A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
</mods:titleInfo>
<mods:name id="F2D392E13E6914A2443E1ED534705F18" type="personal">
<mods:role id="ECE5CED7046F74BAB08B0BF9BDA42659">
<mods:roleTerm id="AAB6E8D7DBD2EFB658F0BC99D7734587">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="499CCCE43405E1645B0F6FBB6E3D1D4C">Enghoff, Henrik</mods:namePart>
<mods:affiliation id="095AC340434C5CB6CDAF3D6F5968B859">Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark &amp; Corresponding author: henghoff @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635</mods:affiliation>
<mods:nameIdentifier id="D1F119C8642AEE3652DD024EF441A718" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:name id="ACC234D16C46C84BBF2011493C28F95E" type="personal">
<mods:role id="ABD3324FE43A1D53E44371D52804FA66">
<mods:roleTerm id="7B63126BCAEB0DFB235E7C95A45CD99E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8317E8C47A5D3C8A6732FCFCFE9078F3">Frederiksen, Sara B.</mods:namePart>
<mods:nameIdentifier id="0D66F1FB5723C7DF8CF99A9690FC2FBA" type="LSID">6DE62DD5-0055-424A-A136-0EF290AF5AFD</mods:nameIdentifier>
<mods:affiliation id="6511895A2EE32B640F2F30EE31226F5D">Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark &amp; Department of Ecology, Animal Ecology, University of Marburg, Karl-von-Frisch-Strasse 8, D- 35032 Marburg, Germany &amp; Email: sara. frederiksen @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: 6 DE 62 DD 5 - 0055 - 424 A-A 136 - 0 EF 290 AF 5 AFD</mods:affiliation>
<mods:nameIdentifier id="39CCE2A08D864344FB15E93F3806DD65" type="email">sara.frederiksen@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E6731C50DC0F7E91D6D55D5EFC60A493">text</mods:typeOfResource>
<mods:relatedItem id="6457B70FCF3C1B302397C79B3D67750F" type="host">
<mods:titleInfo id="D08FBEDC875421BF93EE47814C36E9F1">
<mods:title id="6955FB45C5DBB9C5E4648FABB91ED0A7">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D2DF4BD9C90958D6811CE51CE9721CF3">
<mods:date id="4435EE76B0334190C8BB34D541892A8A">2015</mods:date>
<mods:detail id="F5EFCD3CC793C190CA1F288E7E104B97" type="pubDate">
<mods:number id="CDF61A3FBD0F9FB8CF9A8FCF6EED8996">2015-11-06</mods:number>
</mods:detail>
<mods:detail id="CC8E6F458AFA367DE7A326DB853BEC61" type="issue">
<mods:number id="30666021D8E8C80BE041C1118D329158">150</mods:number>
</mods:detail>
<mods:extent id="A05BEE81AE02B469D7A820C97BFF5AFB" unit="page">
<mods:start id="EB39963E93935BFE99BE52CC70D00767">1</mods:start>
<mods:end id="16A5B982B1A4A3217F8337EDE7B20A84">25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="87F10C5702DB3D826A990361A3F19EE1">journal article</mods:classification>
<mods:identifier id="0A9A3F67780C22A6C467D8D007151E7D" type="CLB-Dataset">22375</mods:identifier>
<mods:identifier id="59999214C715CEC51D89615F60C7DB09" type="DOI">10.5852/ejt.2015.150</mods:identifier>
<mods:identifier id="99D4241EAC033DB827902307431C5B5A" type="GBIF-Dataset">f492ad9b-a83d-4d8e-bdf9-20ef759b75c1</mods:identifier>
<mods:identifier id="E558D78896A0DFD34845A2080E181A0F" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="A02D537019CD22CFF2F4D85AB4789FB9" type="Zenodo-Dep">3787588</mods:identifier>
<mods:identifier id="D0D64103F5658B2249645B5232FC18F2" type="ZooBank">D546A676-35F8-4025-8660-F123A0BA67F7</mods:identifier>
</mods:mods>
<treatment id="03A487F2FFFFFFB775646A139727B3FD" ID-DOI="http://doi.org/10.5281/zenodo.3793789" ID-GBIF-Taxon="163963422" ID-Zenodo-Dep="3793789" LSID="urn:lsid:zoobank.org:act:690C121A-D087-400D-95BC-42BCA7B224BE" httpUri="http://treatment.plazi.org/id/03A487F2FFFFFFB775646A139727B3FD" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
<subSubSection id="C317656FFFFFFFB875646A1396E1B156" box="[571,1017,818,845]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BB236E4FFFFFFB875646A1396E1B156" blockId="6.[361,1227,818,915]" box="[571,1017,818,845]" pageId="6" pageNumber="7">
<heading id="D0FA8188FFFFFFB875646A1396E1B156" box="[571,1017,818,845]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="4C0D4D67FFFFFFB875646A13968FB157" ID-CoL="FZL5" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[571,919,818,845]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="submajor" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB875646A139628B156" bold="true" box="[571,816,818,845]" italics="true" pageId="6" pageNumber="7">Aquattuor submajor</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB874C16A1296E1B156" box="[926,1017,819,845]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFFFFB876366A749655B188" pageId="6" pageNumber="7" type="description">
<paragraph id="8BB236E4FFFFFFB876366A7491D3B16B" blockId="6.[361,1227,818,915]" box="[361,1227,853,880]" pageId="6" pageNumber="7">
<uri id="FF9C3AE6FFFFFFB876366A7491D3B16B" box="[361,1227,853,880]" pageId="6" pageNumber="7">
urn:lsid:zoobank.org:act:
<uuid id="FFAB0C31FFFFFFB875D96A7491D3B16B" box="[646,1227,853,880]" pageId="6" pageNumber="7">690C121A-D087-400D-95BC-42BCA7B224BE</uuid>
</uri>
</paragraph>
<paragraph id="8BB236E4FFFFFFB875B86A589655B188" blockId="6.[361,1227,818,915]" box="[743,845,888,915]" pageId="6" pageNumber="7">
<figureCitation id="13362A61FFFFFFB875B86A589655B188" box="[743,845,888,915]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1860,1886]" captionStartId-1="8.[189,232,1890,1916]" captionTargetBox-0="[228,1353,267,1835]" captionTargetBox-1="[198,1390,269,1859]" captionTargetId-0="figure@7.[227,1360,265,1835]" captionTargetId-1="figure@8.[194,1393,266,1862]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 4. Aquattuor submajor Enghoff sp. nov., paratype. A. Gonopods in situ, right gonopod coloured. B. Left gonopod coxa and basal part of telopodite. C. Left gonopod telopodite and part of coxa, dorsal (basal) view.Articifical colours: orange = coxa; blue = basomere; yellow = solenomere; pink = telomere; red = gonopod sternum; light green = ventral part of eighth pleurotergite (right side); dark green = sternum of reduced ninth leg-pair. mp = metaplica; pa = palette; pp = proplica. Scale bars = 0.1 mm." captionText-1="Fig. 5. Aquattuor submajor Enghoff sp. nov., paratype, left gonopod. A. Anterior view. B. Close-up of mesal incision. C. Tip of telomere. D. Posterior view. btl = basal telomeral lamella; li = lateral incision; mi = mesal incision; mpl = mesal-posterior lamella of telomere; pa = palette; pn = posttorsal narrowing. Scale bars: A, D = 0.1 mm; B–C = 0.01 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787596" figureDoi-1="http://doi.org/10.5281/zenodo.3787598" httpUri-0="https://zenodo.org/record/3787596/files/figure.png" httpUri-1="https://zenodo.org/record/3787598/files/figure.png" pageId="6" pageNumber="7">Figs 4–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFFFFB877E26A9B91CEB677" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BB236E4FFFFFFB877E26A9B942EB1CF" blockId="6.[189,310,954,980]" box="[189,310,954,980]" pageId="6" pageNumber="7">
<heading id="D0FA8188FFFFFFB877E26A9B942EB1CF" bold="true" box="[189,310,954,980]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B979EAF6FFFFFFB877E26A9B942EB1CF" bold="true" box="[189,310,954,980]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26AC891CEB677" blockId="6.[189,1399,1000,1132]" pageId="6" pageNumber="7">
Diameter
<quantity id="4CF59B01FFFFFFB8766C6AC994FAB618" box="[307,482,1000,1027]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" metricValueMax="1.87" metricValueMin="1.55" pageId="6" pageNumber="7" unit="mm" value="1.71" valueMax="1.87" valueMin="1.55">1.55–1.87 mm</quantity>
, 45–48 podous rings. Gonopods: lateral incision of coxa (
<emphasis id="B979EAF6FFFFFFB873CB6AC891BCB618" box="[1172,1188,1001,1027]" italics="true" pageId="6" pageNumber="7">li</emphasis>
) absent to poorly developed; apical palette (
<emphasis id="B979EAF6FFFFFFB876A76D2C970DB63D" box="[504,533,1037,1062]" italics="true" pageId="6" pageNumber="7">pa</emphasis>
) rounded, not longer than broad; telomere describing a
<emphasis id="B979EAF6FFFFFFB873E96D2C91D2B63D" box="[1206,1226,1037,1062]" italics="true" pageId="6" pageNumber="7">c.</emphasis>
270° curve in almost one plane; mesal-posterior lamella of telomere (
<emphasis id="B979EAF6FFFFFFB874196D0E966AB652" box="[838,882,1071,1097]" italics="true" pageId="6" pageNumber="7">mpl</emphasis>
) low; anterior margin of telomere without a triangular tooth; telomere tip with finely dentate margins, internal surface microspiculate.
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFFFFB877E26DB5973EB738" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BB236E4FFFFFFB877E26DB5945CB6B5" blockId="6.[189,324,1172,1198]" box="[189,324,1172,1198]" pageId="6" pageNumber="7">
<heading id="D0FA8188FFFFFFB877E26DB5945CB6B5" bold="true" box="[189,324,1172,1198]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B979EAF6FFFFFFB877E26DB5945CB6B5" bold="true" box="[189,324,1172,1198]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26DE3973EB738" blockId="6.[189,1398,1218,1315]" pageId="6" pageNumber="7">
The name is an artificial Latin adjective meaning “below
<taxonomicName id="4C0D4D67FFFFFFB874036DE296BAB6C7" authorityName="Enghoff" authorityYear="2015" box="[860,930,1219,1244]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B979EAF6FFFFFFB874036DE296BAB6C7" box="[860,930,1219,1244]" italics="true" pageId="6" pageNumber="7">major</emphasis>
</taxonomicName>
” and refers to 1) the smaller size of this species compared to
<taxonomicName id="4C0D4D67FFFFFFB876ED6DC79798B6E4" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[434,640,1253,1280]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB876ED6DC79700B6E4" box="[434,536,1254,1280]" italics="true" pageId="6" pageNumber="7">A. major</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB875D86DC797F9B71B" box="[647,737,1254,1280]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
and 2) the fact that this species occurs at lower altitudes than
<taxonomicName id="4C0D4D67FFFFFFB877A86C2894DDB738" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[247,453,1289,1315]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B979EAF6FFFFFFB877A86C289445B739" box="[247,349,1289,1315]" italics="true" pageId="6" pageNumber="7">A. major</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFFFFB876936C28973EB738" box="[460,550,1289,1315]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFFFFB877E26C6B91E8B4EF" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BB236E4FFFFFFB877E26C6B9724B77E" blockId="6.[189,572,1354,1381]" box="[189,572,1354,1381]" pageId="6" pageNumber="7">
<emphasis id="B979EAF6FFFFFFB877E26C6B9494B77F" bold="true" box="[189,396,1354,1380]" pageId="6" pageNumber="7">Material studied</emphasis>
(total:
<specimenCount id="9D0BFD6DFFFFFFB876BE6C6B972BB77E" box="[481,563,1354,1381]" pageId="6" pageNumber="7" type="male">17 ♂♂</specimenCount>
)
</paragraph>
<materialsCitation id="3B653CB9FFFFFFB877B56C589604B7C2" ID-GBIF-Occurrence="2610408307" collectingDate="1982-08-18" collectionCode="ZMUC" collectorName="Mwanihana FR &amp; M. Stoltze &amp; N. Scharff" country="Tanzania" elevation="1000" location="Udzungwa Mts" municipality="Udzungwa Mts" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
<paragraph id="8BB236E4FFFFFFB877B56C589444B788" blockId="6.[189,1399,1401,1498]" box="[234,348,1401,1427]" pageId="6" pageNumber="7">
<emphasis id="B979EAF6FFFFFFB877B56C589444B788" bold="true" box="[234,348,1401,1427]" pageId="6" pageNumber="7">
<typeStatus id="54B68846FFFFFFB877B56C589444B788" box="[234,348,1401,1427]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26CBD9604B7C2" blockId="6.[189,1399,1401,1498]" pageId="6" pageNumber="7">
<collectingCountry id="F31A7674FFFFFFB877E26CBD944FB7AD" box="[189,343,1436,1462]" name="Tanzania" pageId="6" pageNumber="7">TANZANIA</collectingCountry>
:
<specimenCount id="9D0BFD6DFFFFFFB8763E6CBD9461B7AD" box="[353,377,1436,1462]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
,
<collectingRegion id="49C9F806FFFFFFB876D96CBD9744B7AD" box="[390,604,1436,1462]" country="Tanzania" name="Morogoro" pageId="6" pageNumber="7">Morogoro Region</collectingRegion>
,
<collectingMunicipality id="6BD6AC9EFFFFFFB875386CBD9606B7AD" box="[615,798,1436,1462]" pageId="6" pageNumber="7">Udzungwa Mts</collectingMunicipality>
,
<collectorName id="26F85332FFFFFFB874756CBD96FAB7AD" box="[810,994,1436,1462]" pageId="6" pageNumber="7">Mwanihana FR</collectorName>
,
<locationDeviation id="9A002FC6FFFFFFB874B16CBD9167B7AD" box="[1006,1151,1436,1462]" pageId="6" pageNumber="7">above Sanje</locationDeviation>
,
<elevation id="0020D1D7FFFFFFB873D46CBD9016B7AD" box="[1163,1294,1436,1462]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="1000.0">
<quantity id="4CF59B01FFFFFFB873D46CBD91FCB7AD" box="[1163,1252,1436,1462]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="1000.0">1000 m</quantity>
asl
</elevation>
,
<date id="FFB31024FFFFFFB872456CBD95E2B7C2" pageId="6" pageNumber="7" value="1982-08-18">
<collectingDate id="EFF7E9CCFFFFFFB872456CBD95E2B7C2" pageId="6" pageNumber="7" value="1982-08-18">18 Aug. 1982</collectingDate>
</date>
, in log,
<collectorName id="26F85332FFFFFFB876066CE194CAB7C2" box="[345,466,1471,1498]" pageId="6" pageNumber="7">M. Stoltze</collectorName>
&amp;
<collectorName id="26F85332FFFFFFB876A76CE1976AB7C2" box="[504,626,1471,1498]" pageId="6" pageNumber="7">N. Scharff</collectorName>
leg. (
<collectionCode id="ED1CAE21FFFFFFB875EB6C9E9608B7C2" box="[692,784,1471,1497]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="7">ZMUC</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BB236E4FFFFFFB877B56F20947FB400" blockId="6.[189,1399,1537,1669]" box="[234,359,1537,1563]" pageId="6" pageNumber="7">
<emphasis id="B979EAF6FFFFFFB877B56F20947FB400" bold="true" box="[234,359,1537,1563]" pageId="6" pageNumber="7">
<typeStatus id="54B68846FFFFFFB877B56F20947FB400" box="[234,359,1537,1563]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26F059650B49F" blockId="6.[189,1399,1537,1669]" pageId="6" pageNumber="7">
<collectingCountry id="F31A7674FFFFFFB877E26F05944FB425" box="[189,343,1572,1598]" name="Tanzania" pageId="6" pageNumber="7">TANZANIA</collectingCountry>
:
<specimenCount id="9D0BFD6DFFFFFFB8763E6F0594BCB425" box="[353,420,1572,1598]" pageId="6" pageNumber="7" type="male">7 ♂♂</specimenCount>
, same data as
<typeStatus id="54B68846FFFFFFB875166F0597ABB425" box="[585,691,1572,1598]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
, but in litter, log and pitfall trap, M. Stoltze &amp; N. Scharff leg. (ZMUC);
<specimenCount id="9D0BFD6DFFFFFFB8766A6F66947BB47A" box="[309,355,1607,1633]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, same data but
<quantity id="4CF59B01FFFFFFB875476F669768B47A" box="[536,624,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="7" unit="m" value="1100.0">1100 m</quantity>
asl, pitfall trap (ZMUC);
<specimenCount id="9D0BFD6DFFFFFFB874FF6F6696FCB47A" box="[928,996,1607,1633]" pageId="6" pageNumber="7" type="male">8 ♂♂</specimenCount>
, same data but
<quantity id="4CF59B01FFFFFFB873C66F6691EAB47A" box="[1177,1266,1607,1634]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="7" unit="m" value="1250.0">1250 m</quantity>
asl,
<date id="FFB31024FFFFFFB872776F6695E2B49F" pageId="6" pageNumber="7" value="1982-08-01">1 Aug. 1982</date>
, under log, M. Stoltze &amp; N. Scharff leg. (ZMUC).
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26F8D9445B4DE" blockId="6.[189,349,1707,1734]" box="[189,349,1707,1734]" pageId="6" pageNumber="7">
<heading id="D0FA8188FFFFFFB877E26F8D9445B4DE" bold="true" box="[189,349,1707,1734]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B979EAF6FFFFFFB877E26F8D9445B4DE" bold="true" box="[189,349,1707,1734]" pageId="6" pageNumber="7">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26FFB91E8B4EF" blockId="6.[189,1264,1754,1781]" box="[189,1264,1754,1781]" pageId="6" pageNumber="7">
<collectingCountry id="F31A7674FFFFFFB877E26FFB944FB4EF" box="[189,343,1754,1780]" name="Tanzania" pageId="6" pageNumber="7">TANZANIA</collectingCountry>
:
<collectingRegion id="49C9F806FFFFFFB8763D6FFA9720B4EF" box="[354,568,1754,1781]" country="Tanzania" name="Morogoro" pageId="6" pageNumber="7">Morogoro Region</collectingRegion>
, Udzungwa Mts, Mwanihana FR, above Sanje,
<quantity id="4CF59B01FFFFFFB873376FFB91D9B4EE" box="[1128,1217,1754,1781]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="1000.0">1000 m</quantity>
asl.
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFFFFFB877E26E3A944DB59C" pageId="6" pageNumber="7" type="description">
<paragraph id="8BB236E4FFFFFFB877E26E3A944DB52E" blockId="6.[189,341,1819,1845]" box="[189,341,1819,1845]" pageId="6" pageNumber="7">
<heading id="D0FA8188FFFFFFB877E26E3A944DB52E" bold="true" box="[189,341,1819,1845]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B979EAF6FFFFFFB877E26E3A944DB52E" bold="true" box="[189,341,1819,1845]" pageId="6" pageNumber="7">Colouration</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFFFFFB877E26E6B944DB59C" blockId="6.[189,1398,1866,1927]" pageId="6" pageNumber="7">After 33 years in alcohol faded, only some darker markings on head and collum left. No traces of a middorsal stripe.</paragraph>
</subSubSection>
<caption id="DF72666CFFFEFFB977E26E659044B5F7" ID-DOI="http://doi.org/10.5281/zenodo.3787596" ID-Zenodo-Dep="3787596" httpUri="https://zenodo.org/record/3787596/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1860,1886]" targetBox="[228,1353,267,1835]" targetPageId="7">
<paragraph id="8BB236E4FFFEFFB977E26E659044B5F7" blockId="7.[189,1399,1860,2028]" pageId="7" pageNumber="8">
<emphasis id="B979EAF6FFFEFFB977E26E659414B545" bold="true" box="[189,268,1860,1886]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="4C0D4D67FFFEFFB9764A6E649775B545" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[277,621,1860,1887]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="submajor" status="sp. nov.">
<emphasis id="B979EAF6FFFEFFB9764A6E64971CB545" box="[277,516,1860,1887]" italics="true" pageId="7" pageNumber="8">Aquattuor submajor</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFFEFFB9752A6E6497CCB544" box="[629,724,1861,1887]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, paratype.
<emphasis id="B979EAF6FFFEFFB974086E659674B545" bold="true" box="[855,876,1860,1886]" pageId="7" pageNumber="8">A</emphasis>
. Gonopods
<emphasis id="B979EAF6FFFEFFB974A16E649150B545" box="[1022,1096,1861,1886]" italics="true" pageId="7" pageNumber="8">in situ</emphasis>
, right gonopod coloured.
<emphasis id="B979EAF6FFFEFFB977E26E4995C9B599" bold="true" box="[189,209,1896,1922]" pageId="7" pageNumber="8">B</emphasis>
. Left gonopod coxa and basal part of telopodite.
<emphasis id="B979EAF6FFFEFFB974486E469634B59A" bold="true" box="[791,812,1895,1921]" pageId="7" pageNumber="8">C</emphasis>
. Left gonopod telopodite and part of coxa, dorsal (basal) view.Articifical colours: orange = coxa; blue = basomere; yellow = solenomere; pink = telomere; red = gonopod sternum; light green = ventral part of eighth pleurotergite (right side); dark green = sternum of reduced ninth leg-pair.
<emphasis id="B979EAF6FFFEFFB9750C6EF3976FB5F0" box="[595,631,2002,2027]" italics="true" pageId="7" pageNumber="8">mp</emphasis>
= metaplica;
<emphasis id="B979EAF6FFFEFFB974486EF3962CB5F0" box="[791,820,2002,2027]" italics="true" pageId="7" pageNumber="8">pa</emphasis>
= palette;
<emphasis id="B979EAF6FFFEFFB974EE6EF396D6B5F0" box="[945,974,2002,2027]" italics="true" pageId="7" pageNumber="8">pp</emphasis>
= proplica. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="DF72666CFFF1FFB677E26E4397D1B5FD" ID-DOI="http://doi.org/10.5281/zenodo.3787598" ID-Zenodo-Dep="3787598" httpUri="https://zenodo.org/record/3787598/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1890,1916]" targetBox="[198,1390,269,1859]" targetPageId="8">
<paragraph id="8BB236E4FFF1FFB677E26E4397D1B5FD" blockId="8.[189,1399,1890,2022]" pageId="8" pageNumber="9">
<emphasis id="B979EAF6FFF1FFB677E26E439413B567" bold="true" box="[189,267,1890,1916]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
<taxonomicName id="4C0D4D67FFF1FFB6764C6E439771B567" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[275,617,1890,1916]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="submajor" status="sp. nov.">
<emphasis id="B979EAF6FFF1FFB6764C6E439719B567" box="[275,513,1890,1916]" italics="true" pageId="8" pageNumber="9">Aquattuor submajor</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFF1FFB6752E6E4297D6B566" box="[625,718,1891,1917]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, left gonopod.
<emphasis id="B979EAF6FFF1FFB674AB6E439111B567" bold="true" box="[1012,1033,1890,1916]" pageId="8" pageNumber="9">A</emphasis>
. Anterior view.
<emphasis id="B979EAF6FFF1FFB673966E4391C5B567" bold="true" box="[1225,1245,1890,1916]" pageId="8" pageNumber="9">B</emphasis>
. Close-up of mesal incision.
<emphasis id="B979EAF6FFF1FFB6762B6EA49491B584" bold="true" box="[372,393,1925,1951]" pageId="8" pageNumber="9">C</emphasis>
. Tip of telomere.
<emphasis id="B979EAF6FFF1FFB675046EA49768B584" bold="true" box="[603,624,1925,1951]" pageId="8" pageNumber="9">D</emphasis>
. Posterior view.
<emphasis id="B979EAF6FFF1FFB6746A6EA4964CB584" box="[821,852,1925,1951]" italics="true" pageId="8" pageNumber="9">btl</emphasis>
= basal telomeral lamella;
<emphasis id="B979EAF6FFF1FFB673CD6EA491BAB584" box="[1170,1186,1925,1951]" italics="true" pageId="8" pageNumber="9">li</emphasis>
= lateral incision;
<emphasis id="B979EAF6FFF1FFB677E26E8895C2B5D9" box="[189,218,1961,1986]" italics="true" pageId="8" pageNumber="9">mi</emphasis>
= mesal incision;
<emphasis id="B979EAF6FFF1FFB676EF6E8994C4B5D9" box="[432,476,1960,1986]" italics="true" pageId="8" pageNumber="9">mpl</emphasis>
= mesal-posterior lamella of telomere;
<emphasis id="B979EAF6FFF1FFB674F66E8896DEB5D9" box="[937,966,1961,1986]" italics="true" pageId="8" pageNumber="9">pa</emphasis>
= palette;
<emphasis id="B979EAF6FFF1FFB6731D6E889147B5D9" box="[1090,1119,1961,1986]" italics="true" pageId="8" pageNumber="9">pn</emphasis>
= posttorsal narrowing. Scale bars: A, D = 0.1 mm; B–C = 0.01 mm.
</paragraph>
</caption>
<subSubSection id="C317656FFFF0FFB777E2682A911AB348" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BB236E4FFF0FFB777E2682A94F6B33E" blockId="9.[189,494,267,293]" box="[189,494,267,293]" pageId="9" pageNumber="10">
<heading id="D0FA8188FFF0FFB777E2682A94F6B33E" bold="true" box="[189,494,267,293]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B979EAF6FFF0FFB777E2682A94F6B33E" bold="true" box="[189,494,267,293]" pageId="9" pageNumber="10">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFF0FFB777E2681B911AB348" blockId="9.[189,1026,313,340]" box="[189,1026,313,340]" pageId="9" pageNumber="10">
Known only from Mwanihana FR. Altitudinal range:
<quantity id="4CF59B01FFF0FFB7746F681896CBB34F" box="[816,979,313,340]" metricMagnitude="3" metricUnit="m" metricValue="1.125" metricValueMax="1.25" metricValueMin="1.0" pageId="9" pageNumber="10" unit="m" value="1125.0" valueMax="1250.0" valueMin="1000.0">1000–1250 m</quantity>
asl.
</paragraph>
</subSubSection>
<subSubSection id="C317656FFFF0FFB777E2685A9727B3FD" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BB236E4FFF0FFB777E2685A9486B38E" blockId="9.[189,414,379,405]" box="[189,414,379,405]" pageId="9" pageNumber="10">
<heading id="D0FA8188FFF0FFB777E2685A9486B38E" bold="true" box="[189,414,379,405]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B979EAF6FFF0FFB777E2685A9486B38E" bold="true" box="[189,414,379,405]" pageId="9" pageNumber="10">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="8BB236E4FFF0FFB777E2688B9727B3FD" blockId="9.[189,1398,425,487]" pageId="9" pageNumber="10">
<taxonomicName id="4C0D4D67FFF0FFB777E2688B94F4B3D8" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[189,492,425,452]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="B979EAF6FFF0FFB777E2688B949BB3D8" box="[189,387,426,452]" italics="true" pageId="9" pageNumber="10">Aquattuor major</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="A24A578DFFF0FFB776AA688B974AB3DF" box="[501,594,426,452]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0D4D67FFF0FFB775D168889182B3D8" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[654,1178,425,451]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mwanihanensis">
<emphasis id="B979EAF6FFF0FFB775D1688896FEB3D8" box="[654,998,425,451]" italics="true" pageId="9" pageNumber="10">Chaleponcus mwanihanensis</emphasis>
<bibRefCitation id="EF9C4B15FFF0FFB774B068889182B3D8" author="Enghoff H." box="[1007,1178,425,451]" pageId="9" pageNumber="10" pagination="1 - 75" refId="ref8067" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
occur in the same area, but only at higher altitudes.
</paragraph>
</subSubSection>
</treatment>
</document>
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