228 lines
30 KiB
XML
228 lines
30 KiB
XML
<document id="B7C129F1C8CF93776E48D3B0D56C6DED" ID-DOI="10.11646/zootaxa.4718.3.12" ID-GBIF-Dataset="577b9b02-aced-4d2a-86f5-c0daaebf564d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3751922" ID-ZooBank="90EF5A00-4B9E-493A-B42E-D713BE4499E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586891079281" checkinUser="carolina" docAuthor="Cumming, Jeffrey M. & Brooks, Scott E." docDate="2020" docId="038A87F9FF8DFFDCFF5049EBFE35D4FF" docLanguage="en" docName="zootaxa.4718.3.12.pdf.imf" docOrigin="Zootaxa 4718 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philetus kitsi Cumming & Brooks, 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="447" masterDocId="FFB3FF81FF8DFFDDFFC74A2FFFB9D405" masterDocTitle="A new species of Philetus Melander (Diptera: Empididae) from the Yukon Territory, Canada" masterLastPageNumber="450" masterPageNumber="447" pageNumber="447" updateTime="1698737838582" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CADE76D94330FC9985CD4AEA393E25AB">A new species of Philetus Melander (Diptera: Empididae) from the Yukon Territory, Canada</mods:title>
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<mods:name id="74F14A033B6C20AC30B0D033FD01AEC1" type="personal">
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<mods:namePart id="85477272E74EBC7BBEFE8A2A95E63F88">Cumming, Jeffrey M.</mods:namePart>
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<mods:affiliation id="67CC227FA7B9A66DAEB4F67161403FB6">Canadian National Collection of Insects, Agriculture and Agri-Food Canada</mods:affiliation>
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<mods:nameIdentifier id="5A005D07EC6F2CB6578202CDAFC8CAED" type="email">jeff.cumming@canada.ca</mods:nameIdentifier>
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<mods:namePart id="9967F7A087ECC1FC0F9838D42C5B378A">Brooks, Scott E.</mods:namePart>
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<mods:affiliation id="CBC58E0550DBBFBD63B8437B4265CE96">Canadian National Collection of Insects, Agriculture and Agri-Food Canada</mods:affiliation>
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<mods:nameIdentifier id="D9F7402249BA93350D0B2A206C0D7AD1" type="email">scott.brooks@canada.ca</mods:nameIdentifier>
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<mods:title id="9920D98226870C0AF477CE82EAAA3337">Zootaxa</mods:title>
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<mods:part id="5449FA77698F5059CA5CB5E9D244A4D9">
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<mods:date id="575A3524908000AE6C79BE192CAB94DE">2020</mods:date>
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<mods:number id="88415C7FE000C34FB698C86A1D787C0A">2020-01-07</mods:number>
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<mods:identifier id="B9EF403CD666310517C4C6184FE87C96" type="DOI">10.11646/zootaxa.4718.3.12</mods:identifier>
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<treatment id="038A87F9FF8DFFDCFF5049EBFE35D4FF" ID-DOI="http://doi.org/10.5281/zenodo.3808691" ID-GBIF-Taxon="163631365" ID-Zenodo-Dep="3808691" LSID="urn:lsid:plazi:treatment:038A87F9FF8DFFDCFF5049EBFE35D4FF" httpUri="http://treatment.plazi.org/id/038A87F9FF8DFFDCFF5049EBFE35D4FF" lastPageId="1" lastPageNumber="447" pageId="0" pageNumber="447">
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<subSubSection id="C3396564FF8DFFDDFF5049EBFDA7D7DB" box="[151,542,964,991]" pageId="0" pageNumber="447" type="nomenclature">
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<paragraph id="8B9C36EFFF8DFFDDFF5049EBFDA7D7DB" blockId="0.[151,1437,964,2034]" box="[151,542,964,991]" pageId="0" pageNumber="447">
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<heading id="D0D48183FF8DFFDDFF5049EBFDA7D7DB" bold="true" box="[151,542,964,991]" fontSize="11" level="1" pageId="0" pageNumber="447" reason="1">
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<emphasis id="B957EAFDFF8DFFDDFF5049EBFDA7D7DB" bold="true" box="[151,542,964,991]" pageId="0" pageNumber="447">
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<taxonomicName id="4C234D6CFF8DFFDDFF5049EBFE8CD7DA" ID-CoL="9HWR7" box="[151,309,964,991]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="kitsi" status="sp. nov.">
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<emphasis id="B957EAFDFF8DFFDDFF5049EBFE8CD7DA" bold="true" box="[151,309,964,991]" italics="true" pageId="0" pageNumber="447">Philetus kitsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2645786FF8DFFDDFEFB49EAFE22D7DA" box="[316,411,965,991]" pageId="0" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="13182A6AFF8DFFDDFE6D49EBFDACD7DB" box="[426,533,964,990]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3−5" captionStartId-0="1.[151,258,1933,1955]" captionStartId-1="2.[151,258,1768,1790]" captionTargetBox-0="[160,1428,563,1913]" captionTargetBox-1="[160,1428,342,1737]" captionTargetId-0="figure@1.[160,1428,539,1913]" captionTargetId-1="figure@2.[151,1436,338,1748]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–2. Habitus photograph and collection locality. 1. Philetus kitsi sp. nov. habitus photograph, male holotype. 2. Shrub tundra locality in the Richardson Mountains, Yukon Territory, Canada, where the holotype was collected using a Malaise trap (photograph: J.H. Kits)." captionText-1="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi-0="http://doi.org/10.5281/zenodo.3751924" figureDoi-1="http://doi.org/10.5281/zenodo.3751926" httpUri-0="https://zenodo.org/record/3751924/files/figure.png" httpUri-1="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Figs 1–5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396564FF8DFFDDFF5049C5FC27D007" box="[151,926,1002,1026]" pageId="0" pageNumber="447" type="description">
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<paragraph id="8B9C36EFFF8DFFDDFF5049C5FC27D007" blockId="0.[151,1437,964,2034]" box="[151,926,1002,1026]" pageId="0" pageNumber="447">
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urn:lsid:zoobank.org:act:
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<uuid id="FF850C3AFF8DFFDDFE6749C5FC27D007" box="[416,926,1002,1026]" pageId="0" pageNumber="447">2E951D56-6D9E-4FF2-89B5-E3B88BC411EE</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396564FF8DFFDDFF504E21FE21D097" pageId="0" pageNumber="447" type="materials_examined">
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<paragraph id="8B9C36EFFF8DFFDDFF504E21FE21D097" blockId="0.[151,1437,964,2034]" pageId="0" pageNumber="447">
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<emphasis id="B957EAFDFF8DFFDDFF504E21FE85D023" bold="true" box="[151,316,1038,1062]" pageId="0" pageNumber="447">Type material.</emphasis>
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<materialsCitation id="3B4B3CB2FF8DFFDDFE864E21FE21D097" ID-GBIF-Occurrence="2597661837" collectingDate="2019-07-29" collectingMethod="Malaise trap" collectionCode="CNC" collectorName="J. H. Kits" country="Canada" elevation="850" latitude="67.77868" location="CANADA" longLatPrecision="1" longitude="-136.69885" pageId="0" pageNumber="447" specimenCode="CNC1614247" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B957EAFDFF8DFFDDFE864E21FE61D023" bold="true" box="[321,472,1038,1062]" pageId="0" pageNumber="447">
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<typeStatus id="5498884DFF8DFFDDFE864E21FE61D023" box="[321,472,1038,1062]" pageId="0" pageNumber="447" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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,
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<specimenCount id="9D25FD66FF8DFFDDFE224E21FE40D020" box="[485,505,1038,1061]" pageId="0" pageNumber="447" type="male">♂</specimenCount>
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labelled: “
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<collectingCountry id="F334767FFF8DFFDDFDAD4E21FD5BD023" box="[618,738,1038,1062]" name="Canada" pageId="0" pageNumber="447">CANADA</collectingCountry>
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: YT: Richardson/ Mtns.,
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<geoCoordinate id="EE175028FF8DFFDDFC2D4E23FB3ED023" box="[1002,1159,1036,1062]" degrees="67" direction="north" minutes="46" orientation="latitude" pageId="0" pageNumber="447" precision="1" seconds="43.22" value="67.77868">67°46ʹ43.22ʺN</geoCoordinate>
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/
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<geoCoordinate id="EE175028FF8DFFDDFB524E23FAF1D023" box="[1173,1352,1036,1062]" degrees="136" direction="west" minutes="41" orientation="longitude" pageId="0" pageNumber="447" precision="1" seconds="55.86" value="-136.69885">136°41ʹ55.86ʺW</geoCoordinate>
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,
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<quantity id="4CDB9B0AFF8DFFDDFA964E21FA29D023" box="[1361,1424,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="0" pageNumber="447" unit="m" value="850.0">
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<elevation id="000ED1DCFF8DFFDDFA964E21FA29D023" box="[1361,1424,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="0" pageNumber="447" unit="m" value="850.0">850m</elevation>
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</quantity>
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,/
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<date id="FF9D102FFF8DFFDDFF504E1DFEAAD04F" box="[151,275,1074,1098]" pageId="0" pageNumber="447" value="2019-07-29">
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<collectingDate id="EFD9E9C7FF8DFFDDFF504E1DFEAAD04F" box="[151,275,1074,1098]" pageId="0" pageNumber="447" value="2019-07-29">29.vii.2019</collectingDate>
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</date>
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,
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<collectingMethod id="52624EF8FF8DFFDDFED84E1DFE1ED04F" box="[287,423,1074,1098]" pageId="0" pageNumber="447">Malaise trap</collectingMethod>
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,/ shrub tundra, JK19-075,/
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<collectorName id="26D65339FF8DFFDDFD164E1DFC8AD04F" box="[721,819,1074,1098]" pageId="0" pageNumber="447">J.H. Kits</collectorName>
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,
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<specimenCode id="DB859E94FF8DFFDDFCF84E1DFC61D04F" box="[831,984,1074,1098]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="0" pageNumber="447">CNC1614247</specimenCode>
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”; “HOLOTYPE/
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<taxonomicName id="4C234D6CFF8DFFDDFB514E1DFA9BD04F" box="[1174,1314,1074,1098]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="kitsi">
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<emphasis id="B957EAFDFF8DFFDDFB514E1DFA9BD04F" box="[1174,1314,1074,1098]" italics="true" pageId="0" pageNumber="447">Philetus kitsi</emphasis>
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</taxonomicName>
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/ Cumming & Brooks [red label]” [dissected terminalia in microvial]. Type deposited in the Canadian National Collection of Insects, Ottawa, Canada (
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<collectionCode id="ED32AE2AFF8DFFDDFE974E55FE35D097" box="[336,396,1146,1170]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="0" pageNumber="447">CNC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396564FF8DFFDDFF004EB1FBB5D140" pageId="0" pageNumber="447" type="diagnosis">
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<paragraph id="8B9C36EFFF8DFFDDFF004EB1FBB5D140" blockId="0.[151,1437,964,2034]" pageId="0" pageNumber="447">
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<emphasis id="B957EAFDFF8DFFDDFF004EB1FE85D0B3" bold="true" box="[199,316,1182,1206]" pageId="0" pageNumber="447">Diagnosis.</emphasis>
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<taxonomicName id="4C234D6CFF8DFFDDFE814EB1FE6DD0B3" box="[326,468,1182,1206]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="kitsi" status="sp. nov.">
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<emphasis id="B957EAFDFF8DFFDDFE814EB1FE6DD0B3" box="[326,468,1182,1206]" italics="true" pageId="0" pageNumber="447">Philetus kitsi</emphasis>
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</taxonomicName>
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<emphasis id="B957EAFDFF8DFFDDFE184EB1FD81D0B3" bold="true" box="[479,568,1182,1206]" pageId="0" pageNumber="447">
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<taxonomicNameLabel id="A2645786FF8DFFDDFE184EB1FD81D0B3" box="[479,568,1182,1206]" pageId="0" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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is characterized by male terminalia with elongate simple cercus connected to lengthened ventral subepandrial lobe with digitiform apex (
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<figureCitation id="13182A6AFF8DFFDDFCD94EEDFCE6D0DF" box="[798,863,1218,1242]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 3</figureCitation>
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) and thumb-like preapical medially-directed process (
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<figureCitation id="13182A6AFF8DFFDDFF584EC9FF66D0FB" box="[159,223,1254,1278]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 5</figureCitation>
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), large epandrial lobe with posteriorly hooked dorsal process, and hypandrium with dorsally prolonged narrow median apical process (
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<figureCitation id="13182A6AFF8DFFDDFEB64F25FE6BD127" box="[369,466,1290,1314]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Figs 3, 4</figureCitation>
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). In addition, the longer more narrowly produced apex of the antennal postpedicel (similar to
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<taxonomicName id="4C234D6CFF8DFFDDFF744F00FEE3D143" authorityName="Melander" authorityYear="1927" box="[179,346,1326,1350]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="memorandus">
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<emphasis id="B957EAFDFF8DFFDDFF744F00FF7AD143" box="[179,195,1327,1350]" italics="true" pageId="0" pageNumber="447">P</emphasis>
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||
.
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<emphasis id="B957EAFDFF8DFFDDFF174F01FEE3D143" box="[208,346,1326,1350]" italics="true" pageId="0" pageNumber="447">memorandus</emphasis>
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</taxonomicName>
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) separates this species from most specimens of
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<taxonomicName id="4C234D6CFF8DFFDDFC9D4F00FBBCD143" authorityName="Melander. The" authorityYear="1927" box="[858,1029,1326,1350]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="schizophorus">
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<emphasis id="B957EAFDFF8DFFDDFC9D4F00FCD3D143" box="[858,874,1327,1350]" italics="true" pageId="0" pageNumber="447">P</emphasis>
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||
.
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<emphasis id="B957EAFDFF8DFFDDFCBF4F01FBBCD143" box="[888,1029,1326,1350]" italics="true" pageId="0" pageNumber="447">schizophorus</emphasis>
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</taxonomicName>
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.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396564FF8DFFDDFF004F7DFE21D2AB" pageId="0" pageNumber="447" type="description">
|
||
<paragraph id="8B9C36EFFF8DFFDDFF004F7DFAF5D28F" blockId="0.[151,1437,964,2034]" pageId="0" pageNumber="447">
|
||
<emphasis id="B957EAFDFF8DFFDDFF004F7DFE20D16F" bold="true" box="[199,409,1362,1386]" pageId="0" pageNumber="447">Description. Male.</emphasis>
|
||
Body length
|
||
<quantity id="4CDB9B0AFF8DFFDDFDEF4F7DFDC2D16C" box="[552,635,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="447" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, dark brownish grey, covered with silvery grey tomentum (
|
||
<figureCitation id="13182A6AFF8DFFDDFB284F7DFA94D16F" box="[1263,1325,1362,1386]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,1933,1955]" captionTargetBox="[160,1428,563,1913]" captionTargetId="figure@1.[160,1428,539,1913]" captionTargetPageId="1" captionText="FIGURES 1–2. Habitus photograph and collection locality. 1. Philetus kitsi sp. nov. habitus photograph, male holotype. 2. Shrub tundra locality in the Richardson Mountains, Yukon Territory, Canada, where the holotype was collected using a Malaise trap (photograph: J.H. Kits)." figureDoi="http://doi.org/10.5281/zenodo.3751924" httpUri="https://zenodo.org/record/3751924/files/figure.png" pageId="0" pageNumber="447">Fig. 1</figureCitation>
|
||
). Antenna dark brown to black; postpedicel pyriform with apex narrowly produced towards stylus. Legs brown to greyish brown. Wing hyaline with brownish veins. Halter brownish white. Terminalia: hypandrium with long hook-shaped apical rods arising above base of long, dorsally directed, narrow median apical process (
|
||
<figureCitation id="13182A6AFF8DFFDDFC784F91FC47D1D3" box="[959,1022,1470,1494]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 3</figureCitation>
|
||
); postgonite projected dorsally as wide rounded lobe; phallus tubular, apex partially desclerotized and slightly expanded in dorsal view (
|
||
<figureCitation id="13182A6AFF8DFFDDFB794FCDFABBD1FF" box="[1214,1282,1506,1530]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 4</figureCitation>
|
||
); anterodorsal margin of hypandrium partially fused with anteroventral margin of epandrium; epandrial lobe large with series of short setae on outer and inner margins and posteriorly hooked dorsal process; ventral subepandrial lobe lengthened, shiny, with digitiform apex (
|
||
<figureCitation id="13182A6AFF8DFFDDFE8D4C61FE31D263" box="[330,392,1614,1638]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 3</figureCitation>
|
||
) and thumb-like preapical medially-directed process (
|
||
<figureCitation id="13182A6AFF8DFFDDFC0F4C61FBB1D263" box="[968,1032,1614,1638]" captionStart="FIGURES 3−5" captionStartId="2.[151,258,1768,1790]" captionTargetBox="[160,1428,342,1737]" captionTargetId="figure@2.[151,1436,338,1748]" captionTargetPageId="2" captionText="FIGURES 3−5. Philetus kitsi sp. nov. male terminalia. 3. Lateral view. 4. Hypandrium and phallus, dorsal view. 5. Epandrium and proctiger, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; epand lb—epandrial lobe; goncx apod—gonocoxal apodeme; hypd—hypandrium; hypd rd—hypandrial rod; m ap hypd proc—median apical hypandrial process; pgt—postgonite; ph—phallus; v sbepand lb—ventral subepandrial lobe." figureDoi="http://doi.org/10.5281/zenodo.3751926" httpUri="https://zenodo.org/record/3751926/files/figure.png" pageId="0" pageNumber="447">Fig. 5</figureCitation>
|
||
); ventral subepandrial lobe connected basally to cercus; medial subepandrial lobe absent; cercus elongate, simple, without bifid apex, rounded apically.
|
||
</paragraph>
|
||
<paragraph id="8B9C36EFFF8DFFDDFF004CB9FE21D2AB" blockId="0.[151,1437,964,2034]" box="[199,408,1686,1710]" pageId="0" pageNumber="447">
|
||
<emphasis id="B957EAFDFF8DFFDDFF004CB9FE98D2AB" bold="true" box="[199,289,1686,1710]" pageId="0" pageNumber="447">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396564FF8DFFDDFF004C95FC6FD2F3" pageId="0" pageNumber="447" type="distribution">
|
||
<paragraph id="8B9C36EFFF8DFFDDFF004C95FC6FD2F3" blockId="0.[151,1437,964,2034]" pageId="0" pageNumber="447">
|
||
<emphasis id="B957EAFDFF8DFFDDFF004C95FDCDD2D7" bold="true" box="[199,628,1722,1746]" pageId="0" pageNumber="447">Distribution and seasonal occurrence.</emphasis>
|
||
Known from shrub tundra in the Richardson Mountains of the
|
||
<collectingRegion id="49E7F80DFF8DFFDDFAD34C95FF7AD2F3" country="Canada" name="Yukon" pageId="0" pageNumber="447">Yukon Territory</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334767FFF8DFFDDFF094CF1FEA7D2F3" box="[206,286,1758,1782]" name="Canada" pageId="0" pageNumber="447">Canada</collectingCountry>
|
||
(
|
||
<figureCitation id="13182A6AFF8DFFDDFEEB4CF1FED7D2F3" box="[300,366,1758,1782]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,1933,1955]" captionTargetBox="[160,1428,563,1913]" captionTargetId="figure@1.[160,1428,539,1913]" captionTargetPageId="1" captionText="FIGURES 1–2. Habitus photograph and collection locality. 1. Philetus kitsi sp. nov. habitus photograph, male holotype. 2. Shrub tundra locality in the Richardson Mountains, Yukon Territory, Canada, where the holotype was collected using a Malaise trap (photograph: J.H. Kits)." figureDoi="http://doi.org/10.5281/zenodo.3751924" httpUri="https://zenodo.org/record/3751924/files/figure.png" pageId="0" pageNumber="447">Fig. 2</figureCitation>
|
||
). The single known male was collected at the end of July.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396564FF8DFFDDFF004D2DFE6DD33B" pageId="0" pageNumber="447" type="etymology">
|
||
<paragraph id="8B9C36EFFF8DFFDDFF004D2DFE6DD33B" blockId="0.[151,1437,964,2034]" pageId="0" pageNumber="447">
|
||
<emphasis id="B957EAFDFF8DFFDDFF004D2DFEF1D31F" bold="true" box="[199,328,1794,1818]" pageId="0" pageNumber="447">Etymology.</emphasis>
|
||
This species is named in honour of our colleague, Dr. Joel H. Kits (CNC), who collected the only known specimen of this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396564FF8DFFDCFF004D65FE35D4FF" lastPageId="1" lastPageNumber="448" pageId="0" pageNumber="447" type="discussion">
|
||
<paragraph id="8B9C36EFFF8DFFDCFF004D65FE35D4FF" blockId="0.[151,1437,964,2034]" lastBlockId="1.[151,1437,154,250]" lastPageId="1" lastPageNumber="448" pageId="0" pageNumber="447">
|
||
<emphasis id="B957EAFDFF8DFFDDFF004D65FE8FD367" bold="true" box="[199,310,1866,1890]" pageId="0" pageNumber="447">Remarks.</emphasis>
|
||
Based on the form of the male terminalia and the shape of the postpedicel (see key couplet 1 below),
|
||
<taxonomicName id="4C234D6CFF8DFFDDFA424D64FF7AD383" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="kitsi" status="sp. nov.">
|
||
<emphasis id="B957EAFDFF8DFFDDFA424D64FA2CD367" box="[1413,1429,1867,1890]" italics="true" pageId="0" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B957EAFDFF8DFFDDFF504D41FF7AD383" box="[151,195,1902,1926]" italics="true" pageId="0" pageNumber="447">kitsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EAFDFF8DFFDDFF0D4D41FEA6D383" bold="true" box="[202,287,1902,1926]" pageId="0" pageNumber="447">
|
||
<taxonomicNameLabel id="A2645786FF8DFFDDFF0D4D41FEA6D383" box="[202,287,1902,1926]" pageId="0" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears more similar to
|
||
<taxonomicName id="4C234D6CFF8DFFDDFDE34D40FD72D383" authorityName="Melander" authorityYear="1927" box="[548,715,1902,1926]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="memorandus">
|
||
<emphasis id="B957EAFDFF8DFFDDFDE34D40FD8DD383" box="[548,564,1903,1926]" italics="true" pageId="0" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B957EAFDFF8DFFDDFD864D41FD72D383" box="[577,715,1902,1926]" italics="true" pageId="0" pageNumber="447">memorandus</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C234D6CFF8DFFDDFCD84D40FC70D383" authorityName="Melander. The" authorityYear="1927" box="[799,969,1902,1926]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="schizophorus">
|
||
<emphasis id="B957EAFDFF8DFFDDFCD84D40FC96D383" box="[799,815,1903,1926]" italics="true" pageId="0" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B957EAFDFF8DFFDDFCFB4D41FC70D383" box="[828,969,1902,1926]" italics="true" pageId="0" pageNumber="447">schizophorus</emphasis>
|
||
</taxonomicName>
|
||
. However, it differs from both these species in the habitat occupied,
|
||
<emphasis id="B957EAFDFF8DFFDDFE534DBCFE0DD3AF" box="[404,436,1939,1962]" italics="true" pageId="0" pageNumber="447">i.e.</emphasis>
|
||
, shrub tundra rather than montane and coastal forests.
|
||
<bibRefCitation id="EFB24B1EFF8DFFDDFC314DBDFB5BD3AF" author="Cumming, J. M. & Brooks, S. E. & Sinclair, B. J." box="[1014,1250,1938,1962]" pageId="0" pageNumber="447" pagination="261 - 274" refId="ref1519" refString="Cumming, J. M., Brooks, S. E. & Sinclair, B. J. (2016) Review of the little-known western Nearctic fly genus Philetus Melander (Diptera: Empididae), with a discussion of its phylogenetic assignment. Zootaxa, 4093 (2), 261 - 274. https: // doi. org / 10.11646 / zootaxa. 4093.2.7" type="journal article" year="2016">
|
||
Cumming
|
||
<emphasis id="B957EAFDFF8DFFDDFBA04DBCFB2CD3AF" box="[1127,1173,1938,1962]" italics="true" pageId="0" pageNumber="447">et al</emphasis>
|
||
. (2016
|
||
</bibRefCitation>
|
||
, Appendix) listed several undetermined female specimens of
|
||
<taxonomicName id="4C234D6CFF8DFFDDFDB64D99FD71D3CB" box="[625,712,1974,1998]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAFDFF8DFFDDFDB64D99FD71D3CB" box="[625,712,1974,1998]" italics="true" pageId="0" pageNumber="447">Philetus</emphasis>
|
||
</taxonomicName>
|
||
, including three in particular from the
|
||
<collectingRegion id="49E7F80DFF8DFFDDFBBF4D99FA9CD3CB" box="[1144,1317,1974,1998]" country="Canada" name="Yukon" pageId="0" pageNumber="447">Yukon Territory</collectingRegion>
|
||
(two from the Dempster Highway taken on tundra and one from Whitehorse). Some of these
|
||
<collectingRegion id="49E7F80DFF8DFFDDFBD24DF5FBE2D3F7" box="[1045,1115,2010,2034]" country="Canada" name="Yukon" pageId="0" pageNumber="447">Yukon</collectingRegion>
|
||
specimens possibly represent females of the new species described here. Unfortunately because of their older age and method of capture (from pan and flight intercept traps) they are unlikely to yield mitochondrial DNA barcode (COI) sequences, which could help confirm their species identities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |