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<document id="26CCB097EEF01A1E17A3DCFDC873DFDF" ID="10.11646/zootaxa.4139.3.9" ID-DOI="10.11646/zootaxa.4139.3.9" ID-GBIF-Dataset="e4bda244-3f74-4c60-897e-54d0a74fc3bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="264129" ID-ZooBank="ED8CB393-11E7-43E2-A6B0-D5A11BDEBD71" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469092900070" checkinUser="plazi" docAuthor="Páll-Gergely, Barna" docDate="2016" docId="038A87EDFFE7F953FF10FF64C483E35C" docLanguage="en" docName="zootaxa.4139.3.9.pdf" docOrigin="Zootaxa 4139 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vargapupa humilis Páll-Gergely, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="433" masterDocId="FFB3FF95FFE6F951FF87FFF3C778E563" masterDocTitle="A new species of Vargapupa Páll-Gergely, 2015 and a new synonym of Pseudopomatias Möllendorff, 1885 (Gastropoda: Caenogastropoda: Pupinidae)" masterLastPageNumber="434" masterPageNumber="431" pageNumber="432" updateTime="1698365521852" updateUser="plazi">
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<mods:title id="C617D34BF1FC6037A924912623E0C73E">A new species of Vargapupa Páll-Gergely, 2015 and a new synonym of Pseudopomatias Möllendorff, 1885 (Gastropoda: Caenogastropoda: Pupinidae)</mods:title>
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<mods:namePart id="CFB1FEB51278D24811AD6B04FC04A787">Páll-Gergely, Barna</mods:namePart>
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<mods:title id="CF3E80E753B9A148C2F11C94BA5CBFDC">Zootaxa</mods:title>
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<mods:date id="A0116A70CD4DCFA519F6A1DD66DF8BDF">2016</mods:date>
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<mods:identifier id="24E15C472BB50D98CD785E2D12AB0653" type="DOI">10.11646/zootaxa.4139.3.9</mods:identifier>
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<treatment id="038A87EDFFE7F953FF10FF64C483E35C" ID-DOI="http://doi.org/10.5281/zenodo.6085585" ID-GBIF-Taxon="123198632" ID-Zenodo-Dep="6085585" LSID="urn:lsid:plazi:treatment:038A87EDFFE7F953FF10FF64C483E35C" httpUri="http://treatment.plazi.org/id/038A87EDFFE7F953FF10FF64C483E35C" lastPageId="2" lastPageNumber="433" pageId="1" pageNumber="432">
<subSubSection id="C3396570FFE7F950FF10FF64C653E5B2" pageId="1" pageNumber="432" type="nomenclature">
<paragraph id="8B9C36FBFFE7F950FF10FF64C6BFE5D1" blockId="1.[151,1436,151,452]" box="[151,455,151,178]" pageId="1" pageNumber="432">
<heading id="D0D48197FFE7F950FF10FF64C6BFE5D1" bold="true" box="[151,455,151,178]" fontSize="11" level="1" pageId="1" pageNumber="432" reason="1">
<emphasis id="B957EAE9FFE7F950FF10FF64C6BFE5D1" bold="true" box="[151,455,151,178]" pageId="1" pageNumber="432">
<taxonomicName id="4C234D78FFE7F950FF10FF64C605E5D2" box="[151,381,151,177]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="humilis" status="sp. nov.">
<emphasis id="B957EAE9FFE7F950FF10FF64C605E5D2" bold="true" box="[151,381,151,177]" italics="true" pageId="1" pageNumber="432">Vargapupa humilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2645792FFE7F950FE03FF6BC6BFE5D1" box="[388,455,152,178]" pageId="1" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36FBFFE7F950FF10FF49C653E5B2" blockId="1.[151,1436,151,452]" box="[151,299,186,209]" pageId="1" pageNumber="432">
<figureCitation id="13182A7EFFE7F950FF10FF49C781E5B2" box="[151,249,186,209]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="1" pageNumber="432">Figures 3</figureCitation>
AI.
</paragraph>
</subSubSection>
<subSubSection id="C3396570FFE7F950FF10FF2EC202E43F" pageId="1" pageNumber="432" type="materials_examined">
<paragraph id="8B9C36FBFFE7F950FF10FF2EC202E43F" blockId="1.[151,1436,151,452]" pageId="1" pageNumber="432">
<emphasis id="B957EAE9FFE7F950FF10FF2EC60FE597" bold="true" box="[151,375,221,244]" pageId="1" pageNumber="432">Material examined.</emphasis>
<typeStatus id="54988859FFE7F950FEF9FF2EC699E597" box="[382,481,221,244]" pageId="1" pageNumber="432" type="holotype">Holotype</typeStatus>
(one empty shell,
<collectionCode id="ED32AE3EFFE7F950FD2AFF2EC587E597" box="[685,767,221,244]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="432">MNHN</collectionCode>
<collectionCode id="ED32AE3EFFE7F950FC80FF2EC449E597" box="[775,817,221,244]" pageId="1" pageNumber="432">IM-</collectionCode>
2012-27159,
<figureCitation id="13182A7EFFE7F950FC47FF2EC37AE597" box="[960,1026,221,244]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="1" pageNumber="432">Figs 3</figureCitation>
AE),
<typeStatus id="54988859FFE7F950FBCBFF2EC3D0E597" box="[1100,1192,221,244]" pageId="1" pageNumber="432" type="paratype">paratype</typeStatus>
(one empty shell with broken apex,
<collectionCode id="ED32AE3EFFE7F950FEABFEF3C606E474" box="[300,382,256,279]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="432">MNHN</collectionCode>
2012-27160,
<figureCitation id="13182A7EFFE7F950FD9FFEF3C523E474" box="[536,603,256,279]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="1" pageNumber="432">Figs 3</figureCitation>
FI), and one non-type (one juvenile empty shell,
<collectionCode id="ED32AE3EFFE7F950FB00FEF3C3A1E474" box="[1159,1241,256,279]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="432">MNHN</collectionCode>
<collectionCode id="ED32AE3EFFE7F950FB65FEF3C275E474" box="[1250,1293,256,279]" pageId="1" pageNumber="432">IM-</collectionCode>
2012-27161). Central
<collectingCountry id="F334766BFFE7F950FF69FED1C65CE45A" box="[238,292,290,313]" name="Laos" pageId="1" pageNumber="432">Laos</collectingCountry>
,
<collectingRegion id="49E7F819FFE7F950FEB6FED1C544E45A" box="[305,572,290,313]" country="Laos" name="Louangphabang" pageId="1" pageNumber="432">Luang Prabang province</collectingRegion>
, just NE of Phou Khoun, under rocks in old secondary forest above large cave,
<quantity id="4CDB9B1EFFE7F950FF10FEB6C79EE43F" box="[151,230,325,348]" metricMagnitude="3" metricUnit="m" metricValue="1.177" pageId="1" pageNumber="432" unit="m" value="1177.0">1177 m</quantity>
a.s.l.,
<geoCoordinate id="EE17503CFFE7F950FEABFEB6C6CDE43F" box="[300,437,325,348]" direction="north" orientation="latitude" pageId="1" pageNumber="432" precision="1" value="19.4464">19°26.784'N</geoCoordinate>
,
<geoCoordinate id="EE17503CFFE7F950FE38FEB6C52AE43F" box="[447,594,325,348]" direction="east" orientation="longitude" pageId="1" pageNumber="432" precision="1" value="102.43816">102°26.290'E</geoCoordinate>
, leg. Ahmed Abdou &amp; Igor V. Muratov,
<date id="FF9D103BFFE7F950FB86FEB6C303E43F" box="[1025,1147,325,348]" pageId="1" pageNumber="432" value="2006-11-15">15.11.2006</date>
. (locality code: 32L06).
</paragraph>
</subSubSection>
<subSubSection id="C3396570FFE7F950FF40FE9BC681E4C2" pageId="1" pageNumber="432" type="etymology">
<paragraph id="8B9C36FBFFE7F950FF40FE9BC681E4C2" blockId="1.[151,1436,151,452]" pageId="1" pageNumber="432">
<emphasis id="B957EAE9FFE7F950FF40FE9BC63FE41C" bold="true" box="[199,327,360,383]" pageId="1" pageNumber="432">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C234D78FFE7F950FDA1FE94C50CE41C" box="[550,628,359,383]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="humilis">
<emphasis id="B957EAE9FFE7F950FDA1FE94C50CE41C" box="[550,628,359,383]" italics="true" pageId="1" pageNumber="432">humilis</emphasis>
</taxonomicName>
(Latin: low, small, slight) refers to the basal keel, which is weaker than in the other two
<taxonomicName id="4C234D78FFE7F950FEADFE79C6E6E4C2" box="[298,414,394,417]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B957EAE9FFE7F950FEADFE79C6E6E4C2" box="[298,414,394,417]" italics="true" pageId="1" pageNumber="432">Vargapupa</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C3396570FFE7F950FF40FE5EC225E4A7" box="[199,1373,429,452]" pageId="1" pageNumber="432" type="diagnosis">
<paragraph id="8B9C36FBFFE7F950FF40FE5EC225E4A7" blockId="1.[151,1436,151,452]" box="[199,1373,429,452]" pageId="1" pageNumber="432">
<emphasis id="B957EAE9FFE7F950FF40FE5EC644E4A7" bold="true" box="[199,316,429,452]" pageId="1" pageNumber="432">Diagnosis.</emphasis>
A
<taxonomicName id="4C234D78FFE7F950FEDBFE5EC6A8E4A7" box="[348,464,429,452]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B957EAE9FFE7F950FEDBFE5EC6A8E4A7" box="[348,464,429,452]" italics="true" pageId="1" pageNumber="432">Vargapupa</emphasis>
</taxonomicName>
species with rather spindle-shaped shell, nearly rounded aperture and low basal keel.
</paragraph>
</subSubSection>
<caption id="DF5C6673FFE7F950FF10FB23C6EEE060" httpUri="https://zenodo.org/record/264130/files/figure.png" pageId="1" pageNumber="432" targetBox="[150,1435,503,1210]" targetPageId="1">
<paragraph id="8B9C36FBFFE7F950FF10FB23C6EEE060" blockId="1.[151,1436,1232,1283]" pageId="1" pageNumber="432">
<emphasis id="B957EAE9FFE7F950FF10FB23C676E185" bold="true" box="[151,270,1232,1254]" pageId="1" pageNumber="432">FIGURE 1.</emphasis>
Type sample and labels of
<emphasis id="B957EAE9FFE7F950FD9BFB23C513E186" box="[540,619,1232,1253]" italics="true" pageId="1" pageNumber="432">Mellaca</emphasis>
<taxonomicName id="4C234D78FFE7F950FDF4FB23C42DE186" authority="Mabille, 1889" authorityName="Mabille" authorityYear="1889" box="[627,853,1232,1253]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="balansai">
<emphasis id="B957EAE9FFE7F950FDF4FB23C5BEE186" box="[627,710,1232,1253]" italics="true" pageId="1" pageNumber="432">balansai</emphasis>
Mabille, 1889
</taxonomicName>
(synonym of
<taxonomicName id="4C234D78FFE7F950FC58FB23C2EDE186" authority="Mollendorff, 1885" authorityName="Mollendorff" authorityYear="1885" box="[991,1429,1232,1253]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="amoenus">
<emphasis id="B957EAE9FFE7F950FC58FB23C3A2E186" box="[991,1242,1232,1253]" italics="true" pageId="1" pageNumber="432">Pseudopomatias amoenus</emphasis>
Möllendorff, 1885
</taxonomicName>
) (MNHN IM-2000-31807).
</paragraph>
</caption>
<caption id="DF5C6673FFE7F950FF10F85AC304E2BF" httpUri="https://zenodo.org/record/264131/files/figure.png" pageId="1" pageNumber="432" targetBox="[151,1435,1360,1940]" targetPageId="1">
<paragraph id="8B9C36FBFFE7F950FF10F85AC304E2BF" blockId="1.[151,1435,1961,2012]" pageId="1" pageNumber="432">
<emphasis id="B957EAE9FFE7F950FF10F85AC677E2DC" bold="true" box="[151,271,1961,1983]" pageId="1" pageNumber="432">FIGURE 2.</emphasis>
Lectotype (MNHN IM-2000-32496, Fig. B) and paralectotypes (MNHN IM-2000-31807, Figs A and C) of
<emphasis id="B957EAE9FFE7F950FACAF859C2E3E2DC" box="[1357,1435,1962,1983]" italics="true" pageId="1" pageNumber="432">Mellaca</emphasis>
<taxonomicName id="4C234D78FFE7F950FF10F834C60EE2BF" authority="Mabille, 1889" authorityName="Mabille" authorityYear="1889" box="[151,374,1991,2012]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="balansai">
<emphasis id="B957EAE9FFE7F950FF10F834C792E2BF" box="[151,234,1991,2012]" italics="true" pageId="1" pageNumber="432">balansai</emphasis>
Mabille, 1889
</taxonomicName>
(synonym of
<taxonomicName id="4C234D78FFE7F950FE7BF834C4D7E2BF" authority="Mollendorff, 1885" authorityName="Mollendorff" authorityYear="1885" box="[508,943,1991,2012]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="432" phylum="Mollusca" rank="species" species="amoenus">
<emphasis id="B957EAE9FFE7F950FE7BF834C58DE2BF" box="[508,757,1991,2012]" italics="true" pageId="1" pageNumber="432">Pseudopomatias amoenus</emphasis>
Möllendorff, 1885
</taxonomicName>
). Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="DF5C6673FFE4F953FF10FC8EC352E6D3" httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="2" pageNumber="433" targetBox="[151,1436,191,872]" targetPageId="2">
<paragraph id="8B9C36FBFFE4F953FF10FC8EC352E6D3" blockId="2.[151,1436,893,944]" pageId="2" pageNumber="433">
<emphasis id="B957EAE9FFE4F953FF10FC8EC674E6F0" bold="true" box="[151,268,893,915]" pageId="2" pageNumber="433">FIGURE 3.</emphasis>
Holotype (MNHN IM-2012-27159; Figs AE) and paratype (MNHN IM-2012-27160; Figs FI) of
<taxonomicName id="4C234D78FFE4F953FB49FC8EC2FBE6F1" box="[1230,1411,893,914]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="humilis" status="sp. nov.">
<emphasis id="B957EAE9FFE4F953FB49FC8EC2FBE6F1" box="[1230,1411,893,914]" italics="true" pageId="2" pageNumber="433">Vargapupa humilis</emphasis>
</taxonomicName>
<emphasis id="B957EAE9FFE4F953FA0EFC8EC7CCE6D3" bold="true" pageId="2" pageNumber="433">
<taxonomicNameLabel id="A2645792FFE4F953FA0EFC8EC7CCE6D3" pageId="2" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs BI.
</paragraph>
</caption>
<subSubSection id="C3396570FFE4F953FF40FC28C27BE02F" pageId="2" pageNumber="433" type="description">
<paragraph id="8B9C36FBFFE4F953FF40FC28C220E049" blockId="2.[151,1436,987,1600]" pageId="2" pageNumber="433">
<emphasis id="B957EAE9FFE4F953FF40FC28C62AE691" bold="true" box="[199,338,987,1010]" pageId="2" pageNumber="433">Description.</emphasis>
Shell rather spindle shaped, light brownish-greyish, with 9 regularly increasing, moderately bulging whorls; protoconch of slightly less than one whorl, very finely granulated, not glossy, without radial sculpture; teleoconch with regular, strong but blunt ribs without spiral striation; ribs lacking on neck region above keel (this area is irregularly, finely wrinkled), but 4 to 5 rather regular ribs present behind peristome; ribs stronger between keel and umbilicus than elsewhere; basal keel blunt, low, starts one half whorl before the aperture, above umbilicus, and ends just a bit before the peristome; groove corresponding to keel inside aperture absent; ribs present on keel, especially in the
<typeStatus id="54988859FFE4F953FF10FB58C78BE1A1" box="[151,243,1195,1218]" pageId="2" pageNumber="433" type="paratype">paratype</typeStatus>
(
<figureCitation id="13182A7EFFE4F953FE85FB58C63AE1A1" box="[258,322,1195,1218]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="2" pageNumber="433">Fig. 3</figureCitation>
H); the white line visible above keel of the
<typeStatus id="54988859FFE4F953FC9DFB58C40FE1A1" box="[794,887,1195,1218]" pageId="2" pageNumber="433" type="holotype">holotype</typeStatus>
(
<figureCitation id="13182A7EFFE4F953FC01FB58C4C7E1A1" box="[902,959,1195,1218]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="2" pageNumber="433">Fig 3</figureCitation>
E) is not a thickening of inner shell surface; aperture nearly rounded, palatal-parietal transition angled, columellar-parietal transition not angled; peristome slightly oblique from lateral view (
<figureCitation id="13182A7EFFE4F953FE47FB03C57EE064" box="[448,518,1264,1287]" captionStart="FIGURE 3" captionStartId="2.[151,244,893,915]" captionTargetBox="[151,1436,191,872]" captionTargetId="figure@2.[151,1436,191,872]" captionTargetPageId="2" captionText="FIGURE 3. Holotype (MNHN IM- 2012 - 27159; Figs A E) and paratype (MNHN IM- 2012 - 27160; Figs F I) of Vargapupa humilis n. sp. Leftmost scale = 2 mm and refers to Fig. A, rightmost scale = 1 mm, and refers to Figs B I." httpUri="https://zenodo.org/record/264132/files/figure.png" pageId="2" pageNumber="433">Figs 3</figureCitation>
C, I); parietal callus well developed; apertural rim lighter colour than rest of shell, thickened and slightly reflexed, especially in the direction of the umbilicus; umbilicus very slightly open, slit-like.
</paragraph>
<paragraph id="8B9C36FBFFE4F953FF40FAC6C27BE02F" blockId="2.[151,1436,987,1600]" box="[199,1283,1333,1356]" pageId="2" pageNumber="433">
<emphasis id="B957EAE9FFE4F953FF40FAC6C55CE02F" bold="true" box="[199,548,1333,1356]" pageId="2" pageNumber="433">
Measurements of the
<typeStatus id="54988859FFE4F953FE3CFAC6C567E02F" box="[443,543,1333,1356]" pageId="2" pageNumber="433" type="holotype">holotype</typeStatus>
.
</emphasis>
H:
<quantity id="4CDB9B1EFFE4F953FDCBFAC6C5D5E02F" box="[588,685,1333,1356]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" pageId="2" pageNumber="433" unit="mm" value="8.55">8.55 mm</quantity>
, W: 3.0 mm. The other two examined shells are broken.
</paragraph>
</subSubSection>
<subSubSection id="C3396570FFE4F953FF40FAABC424E37F" pageId="2" pageNumber="433" type="discussion">
<paragraph id="8B9C36FBFFE4F953FF40FAABC424E37F" blockId="2.[151,1436,987,1600]" pageId="2" pageNumber="433">
<emphasis id="B957EAE9FFE4F953FF40FAABC64DE00C" bold="true" box="[199,309,1368,1391]" pageId="2" pageNumber="433">Remarks.</emphasis>
<taxonomicName id="4C234D78FFE4F953FEBAFAAAC578E00C" box="[317,512,1367,1392]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="oharai">
<emphasis id="B957EAE9FFE4F953FEBAFAAAC578E00C" box="[317,512,1367,1392]" italics="true" pageId="2" pageNumber="433">Vargapupa oharai</emphasis>
</taxonomicName>
is much larger, possesses a very strong main basal keel and a weaker additional one, has a channel in the basal part of the aperture corresponding with the main keel, a nearly rectangular aperture, and a less oblique peristome (from lateral view).
<taxonomicName id="4C234D78FFE4F953FDB9FA6DC4B3E0D7" authority="Pall-Gergely, 2015" authorityName="Pall-Gergely" authorityYear="2015" box="[574,971,1437,1461]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="biheli">
<emphasis id="B957EAE9FFE4F953FDB9FA6DC580E0D6" box="[574,760,1437,1461]" italics="true" pageId="2" pageNumber="433">Vargapupa biheli</emphasis>
<bibRefCitation id="EFB24B0AFFE4F953FC86FA6EC4B3E0D7" author="Pall-Gergely" box="[769,971,1437,1460]" pageId="2" pageNumber="433" refString="Pall-Gergely, B., Feher, Z., Hunyadi, A. &amp; Asami, T. (2015) Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa, 3937 (1), 1 - 49. http: // dx. doi. org / 10.11646 / zootaxa. 3937.1" type="journal article" year="2015">Páll-Gergely, 2015</bibRefCitation>
</taxonomicName>
possesses a single basal keel that is much stronger than that of the new species, and has a slender triangular shell, which is rather spindle-shaped in
<taxonomicName id="4C234D78FFE4F953FB72FA32C224E0B4" box="[1269,1372,1471,1496]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="humilis" status="sp. nov.">
<emphasis id="B957EAE9FFE4F953FB72FA32C224E0B4" box="[1269,1372,1471,1496]" italics="true" pageId="2" pageNumber="433">V. humilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2645792FFE4F953FAE4FA33C2E4E0B4" box="[1379,1436,1472,1495]" pageId="2" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
Moreover, similarly to
<taxonomicName id="4C234D78FFE4F953FE21FA10C568E099" box="[422,528,1506,1530]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="oharai">
<emphasis id="B957EAE9FFE4F953FE21FA10C568E099" box="[422,528,1506,1530]" italics="true" pageId="2" pageNumber="433">V. oharai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C234D78FFE4F953FDA2FA10C5FDE099" box="[549,645,1506,1530]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="biheli">
<emphasis id="B957EAE9FFE4F953FDA2FA10C5FDE099" box="[549,645,1506,1530]" italics="true" pageId="2" pageNumber="433">V. biheli</emphasis>
</taxonomicName>
also has a less oblique peristome, a slight apertural basal channel corresponding with the keel, and a somewhat rectangular aperture.
</paragraph>
</subSubSection>
<subSubSection id="C3396570FFE4F953FF40F9DBC483E35C" box="[199,1019,1575,1600]" pageId="2" pageNumber="433" type="distribution">
<paragraph id="8B9C36FBFFE4F953FF40F9DBC483E35C" blockId="2.[151,1436,987,1600]" box="[199,1019,1575,1600]" pageId="2" pageNumber="433">
<emphasis id="B957EAE9FFE4F953FF40F9DBC621E35C" bold="true" box="[199,345,1576,1599]" pageId="2" pageNumber="433">Distribution.</emphasis>
<taxonomicName id="4C234D78FFE4F953FED8F9DAC550E35C" box="[351,552,1575,1600]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="433" phylum="Mollusca" rank="species" species="humilis" status="sp. nov.">
<emphasis id="B957EAE9FFE4F953FED8F9DAC550E35C" box="[351,552,1575,1600]" italics="true" pageId="2" pageNumber="433">Vargapupa humilis</emphasis>
</taxonomicName>
<emphasis id="B957EAE9FFE4F953FDA8F9DBC513E35C" bold="true" box="[559,619,1576,1599]" pageId="2" pageNumber="433">
<taxonomicNameLabel id="A2645792FFE4F953FDA8F9DBC513E35C" box="[559,619,1576,1599]" pageId="2" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is known from the
<typeStatus id="54988859FFE4F953FCBCF9DBC411E35C" box="[827,873,1576,1599]" pageId="2" pageNumber="433">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
</treatment>
</document>