treatments-xml/data/03/8A/87/038A87DEFFC37310FEC78DACFDA17F28.xml

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<mods:title id="C70B50070FC705609B4A9E49BA70398C">Ceraspis jaliscoensis, a new species from México (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="F312D4C9DA70378A667F1C8F5AE5AA7B">Delgado, Leonardo</mods:namePart>
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<mods:namePart id="5D65ADC590D917CB28953B9EB15D0A0D">Navarrete-Heredia, José Luis</mods:namePart>
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<subSubSection id="C3396543FFC37315FEC78DACFB627F8B" box="[264,1220,483,510]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8B9C36C8FFC37315FEC78DACFB627F8B" blockId="1.[264,1220,483,510]" box="[264,1220,483,510]" pageId="1" pageNumber="2">
<heading id="D0D481A4FFC37315FEC78DACFB627F8B" bold="true" box="[264,1220,483,510]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B957EADAFFC37315FEC78DACFB627F8B" bold="true" box="[264,1220,483,510]" pageId="1" pageNumber="2">
<taxonomicName id="4C234D4BFFC37315FEC78DACFC3B7F8B" authority="Delgado and Navarrete-Heredia" authorityName="Delgado and Navarrete-Heredia" box="[264,925,483,510]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jaliscoensis" status="sp. nov.">
<emphasis id="B957EADAFFC37315FEC78DACFDA77F88" bold="true" box="[264,513,484,510]" italics="true" pageId="1" pageNumber="2">Ceraspis jaliscoensis</emphasis>
Delgado and NavarreteHeredia
</taxonomicName>
,
<taxonomicNameLabel id="A26457A1FFC37315FC658DABFB9C7F8B" box="[938,1082,483,509]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="13182A4DFFC37315FB858DABFB1A7F8B" box="[1098,1212,483,509]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,383,1520,1544]" captionTargetBox="[269,1315,572,1509]" captionTargetId="figure@2.[267,1320,569,1514]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Parameres of male genitalia of Ceraspis species. 1, C. jaliscoensis holotype frontal view. 2, Same lateral view of left paramere. 3, C. oaxacaensis frontal view. 4, Same lateral view of left paramere. Scale line = 1 mm." httpUri="https://zenodo.org/record/169632/files/figure.png" pageId="1" pageNumber="2">Figs. 1–2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396543FFC37315FEC78E66FB737D17" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8B9C36C8FFC37315FEC78E66FB737D17" blockId="1.[264,1324,558,1905]" pageId="1" pageNumber="2">
<emphasis id="B957EADAFFC37315FEC78E66FE187C3E" bold="true" box="[264,446,558,584]" pageId="1" pageNumber="2">
<typeStatus id="5498886AFFC37315FEC78E66FEE57C3E" box="[264,323,558,584]" pageId="1" pageNumber="2">Type</typeStatus>
material:
</emphasis>
<typeStatus id="5498886AFFC37315FE088E67FD927C3F" box="[455,564,559,585]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
male deposited at
<collectionCode id="ED32AE0DFFC37315FCC08E67FCF97C3F" box="[783,863,559,585]" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="1" pageNumber="2">CZUG</collectionCode>
labeled: &quot;
<emphasis id="B957EADAFFC37315FC1A8E66FBFC7C3E" bold="true" box="[981,1114,558,584]" pageId="1" pageNumber="2">
<collectingCountry id="F3347658FFC37315FC1A8E66FBF37C3E" box="[981,1109,558,584]" name="Mexico" pageId="1" pageNumber="2">MÉXICO</collectingCountry>
:
</emphasis>
Jalisco, Mascota, gasolinería de la desviación a Talpa,
<date id="FF9D1008FFC37315FD178E1FFCF07C07" box="[728,854,599,625]" pageId="1" pageNumber="2" value="2004-06-05">5VI2004</date>
, Alt.
<quantity id="4CDB9B2DFFC37315FC6E8E1FFBAC7C07" box="[929,1034,599,625]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="2" unit="m" value="1400.0">1,400 m</quantity>
, vegetación secundaria, sobre pasto, G. A. QuirozRocha col.&quot; Three male
<typeStatus id="5498886AFFC37315FC498E37FC507CEF" box="[902,1014,639,665]" pageId="1" pageNumber="2" type="paratype">paratypes</typeStatus>
deposited at
<collectionCode id="ED32AE0DFFC37315FB6C8E37FB557CEF" box="[1187,1267,639,665]" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="1" pageNumber="2">CZUG</collectionCode>
and
<collectionCode id="ED32AE0DFFC37315FEC78EEFFEF47CB7" box="[264,338,679,705]" pageId="1" pageNumber="2">IEXA</collectionCode>
. Two
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labeled: &quot;
<emphasis id="B957EADAFFC37315FD4C8EEEFCAE7CB6" bold="true" box="[643,776,678,704]" pageId="1" pageNumber="2">
<collectingCountry id="F3347658FFC37315FD4C8EEEFCA57CB6" box="[643,771,678,704]" name="Mexico" pageId="1" pageNumber="2">MÉXICO</collectingCountry>
:
</emphasis>
Jalisco, Mascota, Cerro La Mona,
<date id="FF9D1008FFC37315FB678EEFFA807CB7" box="[1192,1318,679,705]" pageId="1" pageNumber="2" value="2004-06-05">5VI2004</date>
, Alt.
<quantity id="4CDB9B2DFFC37315FEF28E87FE047C9F" box="[317,418,719,745]" metricMagnitude="3" metricUnit="m" metricValue="1.433" pageId="1" pageNumber="2" unit="m" value="1433.0">1,433 m</quantity>
, bosque de encinopino,
<geoCoordinate id="EE17500FFFC37315FD048E87FC267C9F" box="[715,896,719,745]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="20.462444">20º 27' 44.8'' N</geoCoordinate>
<geoCoordinate id="EE17500FFFC37315FC468E87FBE07C9F" box="[905,1094,719,745]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="1" value="-104.75061">104º 45' 2.2'' W</geoCoordinate>
, bajo musgo, G. A. QuirozRocha col.” (
<collectionCode id="ED32AE0DFFC37315FE318EBFFDE97D67" box="[510,591,759,785]" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="1" pageNumber="2">CZUG</collectionCode>
,
<collectionCode id="ED32AE0DFFC37315FD938EBFFD017D67" box="[604,679,759,785]" pageId="1" pageNumber="2">IEXA</collectionCode>
). One
<typeStatus id="5498886AFFC37315FD3A8EBFFCFC7D67" box="[757,858,759,785]" pageId="1" pageNumber="2" type="paratype">paratype</typeStatus>
labeled: &quot;
<emphasis id="B957EADAFFC37315FC1C8EBEFBFE7D66" bold="true" box="[979,1112,758,784]" pageId="1" pageNumber="2">
<collectingCountry id="F3347658FFC37315FC1C8EBEFBF57D66" box="[979,1107,758,784]" name="Mexico" pageId="1" pageNumber="2">MÉXICO</collectingCountry>
:
</emphasis>
Jalisco, Mascota, El Atajo, carr. a San Sebastián del Oeste,
<collectionCode id="ED32AE0DFFC37315FD3C8F57FC987D4F" box="[755,830,799,825]" httpUri="http://grbio.org/cool/e0n2-eyvk" name="Buergermeister Mueller, Museum" pageId="1" pageNumber="2">BMM</collectionCode>
,
<geoCoordinate id="EE17500FFFC37315FC858F57FC7C7D4F" box="[842,986,799,825]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="20.633638">20º38'1.1''N</geoCoordinate>
,
<geoCoordinate id="EE17500FFFC37315FC298F57FB277D4F" box="[998,1153,799,825]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="15" value="-104.862495">104º51'45''W</geoCoordinate>
, Alt.
<quantity id="4CDB9B2DFFC37315FB0D8F57FA807D4F" box="[1218,1318,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="1" pageNumber="2" unit="m" value="1413.0">1,413 m</quantity>
, NTP calamar,
<date id="FF9D1008FFC37315FE7E8F0FFDD57D17" box="[433,627,839,865]" pageId="1" pageNumber="2" value="2004-06-05" valueMax="2004-07-03" valueMin="2004-06-05">5.VI3.VII.2004</date>
, J.L. NavarreteHeredia and V.H. Gómez&quot; (
<collectionCode id="ED32AE0DFFC37315FBBD8F0FFB6F7D17" box="[1138,1225,839,865]" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="1" pageNumber="2">CZUG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396543FFC37316FEF78F26FDDD7915" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8B9C36C8FFC37315FEF78F26FB187A0F" blockId="1.[264,1324,558,1905]" pageId="1" pageNumber="2">
<emphasis id="B957EADAFFC37315FEF78F26FE697DFE" bold="true" box="[312,463,878,904]" pageId="1" pageNumber="2">Description.</emphasis>
Male, total length
<quantity id="4CDB9B2DFFC37315FD748F27FC8F7DFF" box="[699,809,879,905]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="1" pageNumber="2" unit="mm" value="13.7">13.7 mm</quantity>
, maximum width (at middle of elytra)
<quantity id="4CDB9B2DFFC37315FAC98F27FE9F7DC7" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="2" unit="mm" value="5.8">5.8 mm</quantity>
. Body ovalelongate; color dull black on frons, pronotum, most of abdomen and thorax; dull reddishbrown on elytra and middle of abdomen; red on clypeus and legs; surface with small, lanceolate, whitish scales. Clypeus trapezoidal, sides straight, apex bilobed, sides and apex perpendicularly reflexed, surface flat; clypeus and frons densely and coarsely punctate with dense, recumbent scales; vertex sparsely punctate with sparse scales; antennae 9segmented, club slightly shorter than segments 2–6 combined.
</paragraph>
<paragraph id="8B9C36C8FFC37316FEF788CFFDFB7F88" blockId="1.[264,1324,558,1905]" lastBlockId="2.[264,1323,284,510]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
Pronotum swollen, semihexagonal, somewhat wider than long, anterior angles acute and posterior angles obtuse; base as wide as base of elytra, base anterior to scutellum fitting in scutellar base; pronotal surface with appressed scales in a pattern more or less longitudinal, forming glabrous areas as follows: along midline, 2 anterolaterad and 2 posterolaterad to midline and at lateral foveae. Scutellum heartshaped with moderately dense, recumbent scales. Elytra longer than wide, elytral surface distinctly shagreened with costae scarcely elevated; elytral intervals with scarce, appressed scales smaller than those of pronotum, scales of first and second intervals coalescing at apical region; humeral umbones obtuse and prominent. Venter covered with scales (at sides), scalelike setae and slender and long setae (at midline). Prosternal process slightly curved, extending almost to apex of procoxae. Abdomen in lateral view strongly concave, first visible abdominal sternite broadly concave at middle; sternites 2–5 distinctly longer at sides than at middle; last abdominal sternite with longitudinal central 1/4 produced at apex, with moderately dense, long, yellowish setae, apical margin finely denticulate with narrow membranous border. Pygidium longer than wide, convex in lateral view, surface shagreened, with appressed scales except at basal 2/3 of midline, apex with dense, yellowish setae. Femora and tibiae with sparse, short, yellowish setae; tarsi with dense, long, golden setae. Protibiae tridentate with basal tooth small, without internal spur, with ventral keel serrate; meso and metatibiae with 2 short, apical spurs. Pro and metatarsi (minus claws) short and thickened, shorter than respective tibiae; first protarsomere with stout, spinose process on ventrolateral face, second protarsomere with same process but smaller; metatarsi (minus claws) slightly longer than metatibiae, less thickened than pro and mesotarsi, metatarsomeres 3–4 as long as wide. All claws simple; unguitractor plate distinctly produced beyond base of claws. Male genitalia with parameres asymmetrical, short, broadly separated, curved at sides and produced at base (
<figureCitation id="13182A4DFFC07316FE128DACFDE97F88" box="[477,591,484,510]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,383,1520,1544]" captionTargetBox="[269,1315,572,1509]" captionTargetId="figure@2.[267,1320,569,1514]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Parameres of male genitalia of Ceraspis species. 1, C. jaliscoensis holotype frontal view. 2, Same lateral view of left paramere. 3, C. oaxacaensis frontal view. 4, Same lateral view of left paramere. Scale line = 1 mm." httpUri="https://zenodo.org/record/169632/files/figure.png" pageId="2" pageNumber="3">Figs. 1–2</figureCitation>
).
</paragraph>
<caption id="DF5C6640FFC07316FEC789B8FDCA783B" httpUri="https://zenodo.org/record/169632/files/figure.png" pageId="2" pageNumber="3" targetBox="[269,1315,572,1509]" targetPageId="2">
<paragraph id="8B9C36C8FFC07316FEC789B8FDCA783B" blockId="2.[264,1323,1520,1613]" pageId="2" pageNumber="3">
<emphasis id="B957EADAFFC07316FEC789B8FE15787E" bold="true" box="[264,435,1520,1544]" pageId="2" pageNumber="3">FIGURES 1–4.</emphasis>
Parameres of male genitalia of
<taxonomicName id="4C234D4BFFC07316FCCA89B9FCC2787E" box="[773,868,1521,1544]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B957EADAFFC07316FCCA89B9FCC2787E" box="[773,868,1521,1544]" italics="true" pageId="2" pageNumber="3">Ceraspis</emphasis>
</taxonomicName>
species. 1,
<taxonomicName id="4C234D4BFFC07316FC1189B9FBDF787E" box="[990,1145,1521,1544]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC07316FC1189B9FBDF787E" box="[990,1145,1521,1544]" italics="true" pageId="2" pageNumber="3">C. jaliscoensis</emphasis>
</taxonomicName>
holotype frontal view. 2, Same lateral view of left paramere. 3,
<taxonomicName id="4C234D4BFFC07316FD338A5BFC39785C" box="[764,927,1555,1578]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC07316FD338A5BFC39785C" box="[764,927,1555,1578]" italics="true" pageId="2" pageNumber="3">C. oaxacaensis</emphasis>
</taxonomicName>
frontal view. 4, Same lateral view of left paramere. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8B9C36C8FFC07316FEF78AE0FDBA78B5" blockId="2.[264,1325,1704,1891]" box="[312,540,1704,1731]" pageId="2" pageNumber="3">
<emphasis id="B957EADAFFC07316FEF78AE0FE3578B4" bold="true" box="[312,403,1704,1730]" pageId="2" pageNumber="3">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B9C36C8FFC07316FEF78A98FDDD7915" blockId="2.[264,1325,1704,1891]" pageId="2" pageNumber="3">
<emphasis id="B957EADAFFC07316FEF78A98FDF7789C" bold="true" box="[312,593,1744,1770]" pageId="2" pageNumber="3">
Variation in
<typeStatus id="5498886AFFC07316FE1A8A98FDF7789C" box="[469,593,1744,1770]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
</emphasis>
. Three males, total length
<quantity id="4CDB9B2DFFC07316FC448A99FB9E789D" box="[907,1080,1745,1771]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.22" metricValueMin="1.1199999999999999" pageId="2" pageNumber="3" unit="mm" value="11.7" valueMax="12.2" valueMin="11.2">11.2–12.2 mm</quantity>
, maximum width (at middle of elytra)
<quantity id="4CDB9B2DFFC07316FE158AB1FDCB7965" box="[474,621,1785,1811]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="2" pageNumber="3" unit="mm" value="5.0" valueMax="5.1" valueMin="4.9">4.9–5.1 mm</quantity>
. The color on frons and pronotum is slightly shining black and density of setae and scales slightly varies. In other respects the
<typeStatus id="5498886AFFC07316FBEF8B69FB36794D" box="[1056,1168,1825,1851]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
do not differ significantly from the
<typeStatus id="5498886AFFC07316FDC28B01FDD17915" box="[525,631,1865,1891]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396543FFC17317FEF78D53FD307D48" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8B9C36C8FFC17317FEF78D53FE3F7C98" blockId="3.[264,1323,283,830]" pageId="3" pageNumber="4">
<emphasis id="B957EADAFFC17317FEF78D53FE177F43" bold="true" box="[312,433,283,309]" pageId="3" pageNumber="4">Remarks.</emphasis>
<taxonomicName id="4C234D4BFFC17317FE748D54FD137F43" box="[443,693,284,309]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FE748D54FD137F43" box="[443,693,284,309]" italics="true" pageId="3" pageNumber="4">Ceraspis jaliscoensis</emphasis>
</taxonomicName>
is included in the Frey's group 3 based on the presence of scales on the dorsum and venter.
<taxonomicName id="4C234D4BFFC17317FD238D0CFC317F2B" box="[748,919,324,349]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FD238D0CFC317F2B" box="[748,919,324,349]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
</taxonomicName>
is most similar to
<emphasis id="B957EADAFFC17317FBBE8D0CFE127FF3" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C234D4BFFC17317FBBE8D0CFA807F2B" box="[1137,1318,324,349]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">C. oaxacaensis</taxonomicName>
,
<taxonomicName id="4C234D4BFFC17317FEC78D24FE127FF3" box="[264,436,364,389]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">C. jaliscoensis</taxonomicName>
</emphasis>
is distinguished from other species of this group (except
<taxonomicName id="4C234D4BFFC17317FBA98D24FABF7FF3" box="[1126,1305,364,389]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FBA98D24FABF7FF3" box="[1126,1305,364,389]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
</taxonomicName>
), by the following combination of characters: pronotum not rounded, scales of pronotum and elytra arranged in a longitudinal pattern, claws simple and color of pronotum and elytra black or reddish brown.
<taxonomicName id="4C234D4BFFC17317FDA08DACFCBF7F8B" box="[623,793,484,509]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FDA08DACFCBF7F8B" box="[623,793,484,509]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="4C234D4BFFC17317FC3D8DACFB027F8B" box="[1010,1188,484,509]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FC3D8DACFB027F8B" box="[1010,1188,484,509]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
</taxonomicName>
by the dorsum that is distinctly shagreened and dull or slightly shining (dorsum with scarce shagreened and shining or slightly dull in
<taxonomicName id="4C234D4BFFC17317FCCC8E7CFC117C3B" box="[771,951,564,589]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FCCC8E7CFC117C3B" box="[771,951,564,589]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
</taxonomicName>
); by the color of elytra wholly reddish brown (wholly black or only with the elytral margins reddish brown in
<taxonomicName id="4C234D4BFFC17317FB1E8E14FECB7CEB" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FB1E8E14FECB7CEB" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
</taxonomicName>
); and by the shape of male genitalia with the parameres shorter and produced at the base (
<figureCitation id="13182A4DFFC17317FEB18EE4FE577CB0" box="[382,497,684,710]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,383,1520,1544]" captionTargetBox="[269,1315,572,1509]" captionTargetId="figure@2.[267,1320,569,1514]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Parameres of male genitalia of Ceraspis species. 1, C. jaliscoensis holotype frontal view. 2, Same lateral view of left paramere. 3, C. oaxacaensis frontal view. 4, Same lateral view of left paramere. Scale line = 1 mm." httpUri="https://zenodo.org/record/169632/files/figure.png" pageId="3" pageNumber="4">Figs. 1–2</figureCitation>
) (parameres longer and not produced at the base in
<taxonomicName id="4C234D4BFFC17317FBBA8EE4FA8D7CB3" box="[1141,1323,684,709]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FBBA8EE4FA8D7CB3" box="[1141,1323,684,709]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
</taxonomicName>
[
<figureCitation id="13182A4DFFC17317FEC08E9CFE227C98" box="[271,388,724,750]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,383,1520,1544]" captionTargetBox="[269,1315,572,1509]" captionTargetId="figure@2.[267,1320,569,1514]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Parameres of male genitalia of Ceraspis species. 1, C. jaliscoensis holotype frontal view. 2, Same lateral view of left paramere. 3, C. oaxacaensis frontal view. 4, Same lateral view of left paramere. Scale line = 1 mm." httpUri="https://zenodo.org/record/169632/files/figure.png" pageId="3" pageNumber="4">Figs. 3–4</figureCitation>
]).
</paragraph>
<paragraph id="8B9C36C8FFC17317FEF78EB4FD307D48" blockId="3.[264,1323,283,830]" pageId="3" pageNumber="4">
The key of the Mexican species of
<taxonomicName id="4C234D4BFFC17317FD288EB4FBB37D60" authority="Delgado 2001" authorityName="Delgado" authorityYear="2001" box="[743,1045,764,790]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B957EADAFFC17317FD288EB4FCE97D63" box="[743,847,764,789]" italics="true" pageId="3" pageNumber="4">Ceraspis</emphasis>
(
<bibRefCitation id="EFB24B39FFC17317FCAD8EB4FBAB7D60" author="Delgado" box="[866,1037,764,790]" pageId="3" pageNumber="6" refString="Delgado, L. (2001) A new species of Ceraspis, with a key to the Mexican species of the genus (Coleoptera: Melolonthidae; Melolonthinae). Journal of the New York Entomological Society, 109 (1), 167 - 170." type="journal article" year="2001">Delgado 2001</bibRefCitation>
)
</taxonomicName>
should be modified as follows to include
<taxonomicName id="4C234D4BFFC17317FE2C8F6CFD2B7D4B" box="[483,653,804,829]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FE2C8F6CFD2B7D4B" box="[483,653,804,829]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
</taxonomicName>
:
</paragraph>
</subSubSection>
<subSubSection id="C3396543FFC17317FEC78F3CFA8C7BE0" pageId="3" pageNumber="4" type="key">
<key id="FFB261F9FFC17317FEC78F3CFA8C7BE0" pageId="3" pageNumber="4">
<keyStep id="30D72E6DFFC17317FEC78F3CFA8D7DA8" pageId="3" pageNumber="4">
<paragraph id="8B9C36C8FFC17317FEC78F3CFA8D7DC0" blockId="3.[264,1324,884,1430]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC78F3CFA8D7DC0" pageId="3" pageNumber="4">1 Dorsum with vestiture of setae or scalelike setae (acicular shape and longer than wide) ....................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B9C36C8FFC17317FEC78F8CFA8D7DA8" blockId="3.[264,1324,884,1430]" box="[264,1323,964,990]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC78F8CFA8D7DA8" box="[264,1323,964,990]" pageId="3" pageNumber="4"> Dorsum with vestiture of scales (spearshaped and broader)........................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D72E6DFFC17317FEC78FA4FA8C7A20" pageId="3" pageNumber="4">
<paragraph id="8B9C36C8FFC17317FEC78FA4FA8C7A70" blockId="3.[264,1324,884,1430]" box="[264,1322,1004,1030]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC78FA4FA8C7A70" box="[264,1322,1004,1030]" pageId="3" pageNumber="4">
2 Clypeus bilobed; pronotum almost evenly convex
<emphasis id="B957EADAFFC17317FC4B8FA4FB9A7A73" box="[900,1084,1004,1029]" italics="true" pageId="3" pageNumber="4">.........................</emphasis>
<taxonomicName id="4C234D4BFFC17317FB8C8FA4FA8C7A70" authority="(Sharp)" authorityName="Sharp" baseAuthorityName="Sharp" box="[1091,1322,1004,1030]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B957EADAFFC17317FB8C8FA4FB6A7A73" box="[1091,1228,1004,1029]" italics="true" pageId="3" pageNumber="4">C. centralis</emphasis>
(Sharp)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C36C8FFC17317FEC7885CFA8C7A20" blockId="3.[264,1324,884,1430]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC7885CFA8C7A20" pageId="3" pageNumber="4">
 Clypeus straight or broadly emarginate; pronotum distinctly elevated...........................
<emphasis id="B957EADAFFC17317FEF48874FB747A23" box="[315,1234,1084,1109]" italics="true" pageId="3" pageNumber="4">
........................................................................................................
<taxonomicName id="4C234D4BFFC17317FBF68874FB747A23" authority="Harold" authorityName="Harold" box="[1081,1234,1084,1109]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mexicana">C. mexicana</taxonomicName>
</emphasis>
Harold
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D72E6DFFC17317FEC7882CFA8D7A80" pageId="3" pageNumber="4">
<paragraph id="8B9C36C8FFC17317FEC7882CFA8C7AD0" blockId="3.[264,1324,884,1430]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC7882CFA8C7AD0" pageId="3" pageNumber="4">
3 Pronotum almost entirely covered with recumbent scales (reclining on the surface), not forming a longitudinal pattern
<emphasis id="B957EADAFFC17317FD0988C4FBED7AD3" box="[710,1099,1164,1189]" italics="true" pageId="3" pageNumber="4">.....................................................</emphasis>
<taxonomicName id="4C234D4BFFC17317FB8188C4FA8C7AD0" authority="(Bates)" authorityName="Bates" baseAuthorityName="Bates" box="[1102,1322,1164,1190]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="velutina">
<emphasis id="B957EADAFFC17317FB8188C4FB687AD3" box="[1102,1230,1164,1189]" italics="true" pageId="3" pageNumber="4">C. velutina</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C36C8FFC17317FEC788FCFA8D7A80" blockId="3.[264,1324,884,1430]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC788FCFA8D7A80" pageId="3" pageNumber="4"> Pronotum with appressed scales (closely applied to the surface), forming a longitudi nal pattern...................................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D72E6DFFC17317FEC7894CFA8A7B30" pageId="3" pageNumber="4">
<paragraph id="8B9C36C8FFC17317FEC7894CFA8C7B68" blockId="3.[264,1324,884,1430]" box="[264,1322,1284,1310]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC7894CFA8C7B68" box="[264,1322,1284,1310]" pageId="3" pageNumber="4">
4 Pronotum and elytra red .....................................................................
<taxonomicName id="4C234D4BFFC17317FB92894CFA8C7B68" authority="Harold" authorityName="Harold" box="[1117,1322,1284,1310]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pilatei">
<emphasis id="B957EADAFFC17317FB92894CFB747B6B" box="[1117,1234,1284,1309]" italics="true" pageId="3" pageNumber="4">C. pilatei</emphasis>
Harold
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C36C8FFC17317FEC78964FA8A7B30" blockId="3.[264,1324,884,1430]" box="[264,1324,1324,1350]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC78964FA8A7B30" box="[264,1324,1324,1350]" pageId="3" pageNumber="4"> Pronotum and elytra black or reddish brown ................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D72E6DFFC17317FEC7891CFA8C7BE0" pageId="3" pageNumber="4">
<paragraph id="8B9C36C8FFC17317FEC7891CFA8C7B18" blockId="3.[264,1324,884,1430]" box="[264,1322,1364,1390]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC7891CFA8C7B18" box="[264,1322,1364,1390]" pageId="3" pageNumber="4">
5 Elytra almost wholly shining black..........................................
<taxonomicName id="4C234D4BFFC17317FBC1891CFA8C7B18" authority="Delgado" authorityName="Delgado" box="[1038,1322,1364,1390]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FBC1891CFB667B1B" box="[1038,1216,1364,1389]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
Delgado
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9C36C8FFC17317FEC78934FA8C7BE0" blockId="3.[264,1324,884,1430]" box="[264,1322,1404,1430]" pageId="3" pageNumber="4">
<keyLead id="30D295FDFFC17317FEC78934FA8C7BE0" box="[264,1322,1404,1430]" pageId="3" pageNumber="4">
 Elytra wholly dull reddish brown
<emphasis id="B957EADAFFC17317FD098934FC0B7BE3" box="[710,941,1404,1429]" italics="true" pageId="3" pageNumber="4">
........
<taxonomicName id="4C234D4BFFC17317FCCB8934FC0B7BE3" authority="Delgado and Navarrete-Heredia" authorityName="Delgado and Navarrete-Heredia" box="[772,941,1404,1429]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">C. jaliscoensis</taxonomicName>
</emphasis>
Delgado and NavarreteHeredia
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C3396543FFC17317FEF78989FB1C791D" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8B9C36C8FFC17317FEF78989FDFF7885" blockId="3.[264,1324,1473,1899]" pageId="3" pageNumber="4">
<emphasis id="B957EADAFFC17317FEF78989FE7E7BAD" bold="true" box="[312,472,1473,1499]" pageId="3" pageNumber="4">Distribution.</emphasis>
The discovery of
<taxonomicName id="4C234D4BFFC17317FD768989FCCE7BAC" box="[697,872,1473,1498]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FD768989FCCE7BAC" box="[697,872,1473,1498]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
</taxonomicName>
extends the distribution of the genus
<taxonomicName id="4C234D4BFFC17317FEC789A1FED67874" box="[264,368,1513,1538]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B957EADAFFC17317FEC789A1FED67874" box="[264,368,1513,1538]" italics="true" pageId="3" pageNumber="4">Ceraspis</emphasis>
</taxonomicName>
from
<geoCoordinate id="EE17500FFFC17317FE7589A1FD8F7875" box="[442,553,1513,1539]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="18.75">18º 45' N</geoCoordinate>
to
<geoCoordinate id="EE17500FFFC17317FD9F89A1FD187875" box="[592,702,1513,1539]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="20.45">20º 27' N</geoCoordinate>
(
<figureCitation id="13182A4DFFC17317FD1F89A1FCE47875" box="[720,834,1513,1539]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="4.[264,368,1050,1074]" captionStartId-1="4.[264,368,1841,1865]" captionStartId-2="5.[264,368,923,947]" captionTargetBox-0="[408,1192,460,1050]" captionTargetBox-1="[428,1192,1222,1786]" captionTargetBox-2="[411,1174,328,892]" captionTargetId-0="figure@4.[407,1192,448,1050]" captionTargetId-1="figure@4.[414,1207,1207,1823]" captionTargetId-2="figure@5.[400,1187,315,916]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 5. Distribution of the Mexican species of Ceraspis. C. jaliscoensis (triangles), C. oaxacaensis (square) and, C. velutina (circles)." captionText-1="FIGURE 6. Distribution of the Mexican species of Ceraspis. C. mexicana (circles) and, C. centralis (squares)." captionText-2="FIGURE 7. Distribution of Ceraspis pilatei (circles; empty diamonds, country record without specific data)." httpUri-0="https://zenodo.org/record/158247/files/figure.png" httpUri-1="https://zenodo.org/record/158248/files/figure.png" httpUri-2="https://zenodo.org/record/158249/files/figure.png" pageId="3" pageNumber="4">Figs. 5–7</figureCitation>
). The new species is found at the highest altitude of any species of this genus (
<quantity id="4CDB9B2DFFC17317FD738A59FC85785D" box="[700,803,1553,1579]" metricMagnitude="3" metricUnit="m" metricValue="1.433" pageId="3" pageNumber="4" unit="m" value="1433.0">1,433 m</quantity>
). In
<collectingCountry id="F3347658FFC17317FC988A59FC12785D" box="[855,948,1553,1579]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
, others species are found below
<quantity id="4CDB9B2DFFC17317FEC78A71FECE7825" box="[264,360,1593,1619]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="1000.0">1,000 m</quantity>
(
<bibRefCitation id="EFB24B39FFC17317FEB78A71FDC37825" author="Delgado" box="[376,613,1593,1619]" pageId="3" pageNumber="6" refString="Delgado, L., Deloya, C. &amp; Moron, M. A. (1987) Descripcion del macho de Ceraspis velutina (Bates) y nuevos registros de C. centralis (Sharp) para Mexico (Coleoptera: Melolonthidae; Macrodactylini). Folia Entomologica Mexicana, 71, 47 - 54." type="journal article" year="1987">Delgado et al. 1987</bibRefCitation>
). Additionally, the distribution of
<taxonomicName id="4C234D4BFFC17317FC3C8A71FB387824" box="[1011,1182,1593,1618]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FC3C8A71FB387824" box="[1011,1182,1593,1618]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
</taxonomicName>
extends the range more than
<quantity id="4CDB9B2DFFC17317FE008A29FD8F780D" box="[463,553,1633,1659]" metricMagnitude="5" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="4" unit="km" value="670.0">670 km</quantity>
west in
<collectingCountry id="F3347658FFC17317FD438A29FD43780D" box="[652,741,1633,1659]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
(
<geoCoordinate id="EE17500FFFC17317FD398A29FCD9780D" box="[758,895,1633,1659]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="-104.75">104º 45' W</geoCoordinate>
). Three species of
<taxonomicName id="4C234D4BFFC17317FB938A29FB62780C" box="[1116,1220,1633,1658]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B957EADAFFC17317FB938A29FB62780C" box="[1116,1220,1633,1658]" italics="true" pageId="3" pageNumber="4">Ceraspis</emphasis>
</taxonomicName>
are now endemic to
<collectingCountry id="F3347658FFC17317FE5C8AC1FE5778D5" box="[403,497,1673,1699]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
:
<taxonomicName id="4C234D4BFFC17317FE328AC1FC8178D5" authority="(Oaxaca)" baseAuthorityName="Oaxaca" box="[509,807,1673,1699]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oaxacaensis">
<emphasis id="B957EADAFFC17317FE328AC1FD1778D4" box="[509,689,1673,1698]" italics="true" pageId="3" pageNumber="4">C. oaxacaensis</emphasis>
(Oaxaca)
</taxonomicName>
,
<taxonomicName id="4C234D4BFFC17317FCFB8AC1FB9A78D5" authority="(Guerrero)" baseAuthorityName="Guerrero" box="[820,1084,1673,1699]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="velutina">
<emphasis id="B957EADAFFC17317FCFB8AC1FC1178D4" box="[820,951,1673,1698]" italics="true" pageId="3" pageNumber="4">C. velutina</emphasis>
(Guerrero)
</taxonomicName>
, and
<taxonomicName id="4C234D4BFFC17317FBB18AC1FECD78BD" authority="(Jalisco)" baseAuthorityName="Jalisco" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC17317FBB18AC1FA8D78D4" box="[1150,1323,1673,1698]" italics="true" pageId="3" pageNumber="4">C. jaliscoensis</emphasis>
(Jalisco)
</taxonomicName>
(
<figureCitation id="13182A4DFFC17317FEB18AF9FE6E78BD" box="[382,456,1713,1739]" captionStart="FIGURE 5" captionStartId="4.[264,368,1050,1074]" captionTargetBox="[408,1192,460,1050]" captionTargetId="figure@4.[407,1192,448,1050]" captionTargetPageId="4" captionText="FIGURE 5. Distribution of the Mexican species of Ceraspis. C. jaliscoensis (triangles), C. oaxacaensis (square) and, C. velutina (circles)." httpUri="https://zenodo.org/record/158247/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
). The remaining nonendemic species occur in southern Mexican states south to
<collectingCountry id="F3347658FFC17317FEA28A91FE6C7885" box="[365,458,1753,1779]" name="Panama" pageId="3" pageNumber="4">Panamá</collectingCountry>
(
<figureCitation id="13182A4DFFC17317FE158A91FDEA7885" box="[474,588,1753,1779]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="4.[264,368,1841,1865]" captionStartId-1="5.[264,368,923,947]" captionTargetBox-0="[428,1192,1222,1786]" captionTargetBox-1="[411,1174,328,892]" captionTargetId-0="figure@4.[414,1207,1207,1823]" captionTargetId-1="figure@5.[400,1187,315,916]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 6. Distribution of the Mexican species of Ceraspis. C. mexicana (circles) and, C. centralis (squares)." captionText-1="FIGURE 7. Distribution of Ceraspis pilatei (circles; empty diamonds, country record without specific data)." httpUri-0="https://zenodo.org/record/158248/files/figure.png" httpUri-1="https://zenodo.org/record/158249/files/figure.png" pageId="3" pageNumber="4">Figs. 6–7</figureCitation>
).
</paragraph>
<paragraph id="8B9C36C8FFC17317FEF78B49FB1C791D" blockId="3.[264,1324,1473,1899]" pageId="3" pageNumber="4">
<emphasis id="B957EADAFFC17317FEF78B49FE67796D" bold="true" box="[312,449,1793,1819]" pageId="3" pageNumber="4">Bionomics.</emphasis>
The adults were collected on grass (
<specimenCount id="9D25FD41FFC17317FCBE8B49FC5E796D" box="[881,1016,1793,1819]" pageId="3" pageNumber="4" type="generic">1 specimen</specimenCount>
), under moss (2) and, in a carrion trap baited with squid (1). The carrionbaited trap was probably an accidental capture, since the adults of this genus feed on flowers (
<bibRefCitation id="EFB24B39FFC17317FCAC8B19FB0A791D" author="Capistran" box="[867,1196,1873,1899]" pageId="3" pageNumber="6" refString="Capistran, F. &amp; Aquino, O. (1992) Observaciones sobre la biologia de Ceraspis pilatei Harold (Coleoptera: Melolonthidae), en el area de Catemaco, Veracruz, Mexico. Folia Entomologica Mexicana, 85, 119 - 122." type="journal article" year="1992">Capistrán and Aquino 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396543FFC67310FEF78D53FDA17F28" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8B9C36C8FFC67310FEF78D53FDA17F28" blockId="4.[264,1322,283,350]" pageId="4" pageNumber="5">
<emphasis id="B957EADAFFC67310FEF78D53FE627F43" bold="true" box="[312,452,283,309]" pageId="4" pageNumber="5">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C234D4BFFC67310FD008D54FCFE7F43" box="[719,856,284,309]" class="Insecta" family="Melolonthidae" genus="Ceraspis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jaliscoensis">
<emphasis id="B957EADAFFC67310FD008D54FCFE7F43" box="[719,856,284,309]" italics="true" pageId="4" pageNumber="5">jaliscoensis</emphasis>
</taxonomicName>
, refers to Jalisco, the state where was collected this species.
</paragraph>
</subSubSection>
</treatment>
</document>