339 lines
57 KiB
XML
339 lines
57 KiB
XML
<document id="25E1D04A1370E24F6755437DF0DD0C30" ID-CLB-Dataset="25440" ID-DOI="10.11646/zootaxa.4648.2.10" ID-GBIF-Dataset="195fa4b8-0cfa-4f61-b6b1-3a949817b2aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3354776" ID-ZooBank="649E4C40-EC59-4A68-8BAF-1E9BB50A9FAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564469048204" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos" docDate="2019" docId="038A87D5FFA9FFB480E6FCADFF2DF907" docLanguage="en" docName="zootaxa4648.2.10.pdf" docOrigin="Zootaxa 4648 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Peripus Chatzimanolis 2019, new genus" docType="treatment" docUuid="722D927C-3FB4-4A2A-AFE1-B189B8BD623A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="375" masterDocId="FFB3FFADFFA8FFB08071FFE8FFF8FFE4" masterDocTitle="Peripus, a new genus of Xanthopygina (Coleoptera: Staphylinidae) from South America" masterLastPageNumber="383" masterPageNumber="371" pageNumber="372" updateTime="1698747768664" updateUser="ExternalLinkService">
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<mods:title id="0C68EED01690911E5D69972C6A8EFA20">Peripus, a new genus of Xanthopygina (Coleoptera: Staphylinidae) from South America</mods:title>
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<mods:name id="02A521D76026455AACEE6DFD8DA06824" type="personal">
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<mods:namePart id="F13B583051061FD3C011705A14D266ED">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="859C3CF444F5A90D4052F14BEC4745E9">Zootaxa</mods:title>
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<mods:part id="AD218AD24BCA58C726E63D84FA0F1A4E">
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<mods:date id="53C8FBF40539571E73AB5EA292C11210">2019</mods:date>
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<mods:number id="95BDFA7417D0F15D6442F7958D46256E">2019-07-30</mods:number>
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<mods:number id="F381AE27A7AE1AF539D481FD076AEE9F">4648</mods:number>
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<treatment id="038A87D5FFA9FFB480E6FCADFF2DF907" ID-GBIF-Taxon="159628147" LSID="urn:lsid:zoobank.org:act:722D927C-3FB4-4A2A-AFE1-B189B8BD623A" httpUri="http://treatment.plazi.org/id/038A87D5FFA9FFB480E6FCADFF2DF907" lastPageId="4" lastPageNumber="375" pageId="1" pageNumber="372">
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<subSubSection id="C3396548FFA9FFB180E6FCADFCF1FCBB" box="[151,777,837,863]" pageId="1" pageNumber="372" type="nomenclature">
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<paragraph id="8B9C36C3FFA9FFB180E6FCADFCF1FCBB" blockId="1.[151,983,837,935]" box="[151,777,837,863]" pageId="1" pageNumber="372">
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<heading id="D0D481AFFFA9FFB180E6FCADFCF1FCBB" bold="true" box="[151,777,837,863]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
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<emphasis id="B957EAD1FFA9FFB180E6FCADFCF1FCBB" bold="true" box="[151,777,837,863]" pageId="1" pageNumber="372">
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<taxonomicName id="4C234D40FFA9FFB180E6FCADFD83FCBB" ID-CoL="8XNSG" authority="Chatzimanolis and Hightower" authorityName="Chatzimanolis" authorityYear="2019" box="[151,635,837,863]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B957EAD1FFA9FFB180E6FCADFF0CFCBB" bold="true" box="[151,244,837,863]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
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, Chatzimanolis and Hightower
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26457AAFFA9FFB182F6FCADFCF1FCBB" box="[647,777,837,863]" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3396548FFA9FFB180E6FC81FC2FFC43" pageId="1" pageNumber="372" type="description">
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<paragraph id="8B9C36C3FFA9FFB180E6FC81FED8FC67" blockId="1.[151,983,837,935]" box="[151,288,873,899]" pageId="1" pageNumber="372">
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(
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<figureCitation id="13182A46FFA9FFB180EEFC81FEEFFC67" box="[159,279,873,899]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–14" captionStart-3="FIGURES 15–20" captionStart-4="FIGURES 21–23" captionStart-5="FIGURES 24–26" captionStart-6="FIGURES 27–29" captionStart-7="FIGURES 30–32" captionStart-8="FIGURE 33" captionStartId-0="2.[151,264,1942,1967]" captionStartId-1="3.[151,264,1927,1952]" captionStartId-2="7.[151,264,1890,1915]" captionStartId-3="8.[151,264,1856,1881]" captionStartId-4="9.[151,264,1082,1107]" captionStartId-5="9.[151,264,1935,1960]" captionStartId-6="10.[151,264,936,961]" captionStartId-7="10.[151,264,1762,1787]" captionStartId-8="11.[151,250,1744,1769]" captionTargetBox-0="[253,1334,170,1922]" captionTargetBox-1="[151,1429,196,1890]" captionTargetBox-2="[151,1436,793,1871]" captionTargetBox-3="[151,1436,776,1817]" captionTargetBox-4="[151,1425,344,1040]" captionTargetBox-5="[151,1425,1185,1885]" captionTargetBox-6="[151,1425,181,905]" captionTargetBox-7="[151,1436,1019,1743]" captionTargetBox-8="[151,1436,373,1695]" captionTargetId-0="figure@2.[239,1349,166,1922]" captionTargetId-1="figure@3.[151,1436,196,1890]" captionTargetId-2="figure@7.[151,1436,793,1871]" captionTargetId-3="figure@8.[151,1436,776,1817]" captionTargetId-4="figure@9.[151,1436,344,1051]" captionTargetId-5="figure@9.[151,1436,1178,1896]" captionTargetId-6="figure@10.[151,1436,181,916]" captionTargetId-7="figure@10.[151,1436,1019,1743]" captionTargetId-8="figure@11.[151,1436,373,1695]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIGURES 1–4. Habitus photographs of species of Peripus Chatzimanolis & Hightower. 1. The holotype of P. brunkei Chatzimanolis & Hightower. 2. Peripus madrededios Chatzimanolis & Hightower. 3. Peripus didontus Chatzimanolis & Hightower. 4. Peripus monodontus Chatzimanolis & Hightower. Not to scale." captionText-1="FIGURES 5–8. Heads and pronota of Peripus Chatzimanolis & Hightower. 5. Peripus brunkei Chatzimanolis & Hightower. 6. Peripus didontus Chatzimanolis & Hightower. 7. Peripus madrededios Chatzimanolis & Hightower. 8. Peripus monodontus Chatzimanolis & Hightower. Not to scale." captionText-2="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." captionText-3="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." captionText-4="FIGURES 21–23. Aedeagus of P. brunkei Chatzimanolis & Hightower. 21. Dorsal view. 22. Lateral view. 23. Detail of paramere, ventral view." captionText-5="FIGURES 24–26. Aedeagus of P. didontus Chatzimanolis & Hightower. 24. Dorsal view. 25. Lateral view. 26. Detail of paramere, ventral view." captionText-6="FIGURES 27–29. Aedeagus of P. madrededios Chatzimanolis & Hightower. 27. Dorsal view. 28. Lateral view. 29. Detail of paramere, ventral view." captionText-7="FIGURES 30–32. Aedeagus of P. monodontus Chatzimanolis & Hightower. 30. Dorsal view. 31. Lateral view. 32. Detail of paramere, ventral view." captionText-8="FIGURE 33. Distribution map of Peripus Chatzimanolis & Hightower: P. brunkei Chatzimanolis & Hightower (squares); P. didontus Chatzimanolis & Hightower (stars); P. madrededios Chatzimanolis & Hightower (circles); and P. monodontus Chatzimanolis & Hightower (triangles)." httpUri-0="https://zenodo.org/record/3354778/files/figure.png" httpUri-1="https://zenodo.org/record/3354780/files/figure.png" httpUri-2="https://zenodo.org/record/3354782/files/figure.png" httpUri-3="https://zenodo.org/record/3354784/files/figure.png" httpUri-4="https://zenodo.org/record/3354786/files/figure.png" httpUri-5="https://zenodo.org/record/3354788/files/figure.png" httpUri-6="https://zenodo.org/record/3354792/files/figure.png" httpUri-7="https://zenodo.org/record/3354794/files/figure.png" httpUri-8="https://zenodo.org/record/3354796/files/figure.png" pageId="1" pageNumber="372">Figs. 1–33</figureCitation>
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)
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</paragraph>
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<paragraph id="8B9C36C3FFA9FFB180E6FC65FC2FFC43" blockId="1.[151,983,837,935]" box="[151,983,909,935]" pageId="1" pageNumber="372">
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<uri id="FFB23AC1FFA9FFB180E6FC65FC2FFC43" box="[151,983,909,935]" pageId="1" pageNumber="372">
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urn:lsid:zoobank.org:act:
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<uuid id="FF850C16FFA9FFB181DDFC65FC2FFC43" box="[428,983,909,935]" pageId="1" pageNumber="372">722D927C-3FB4-4A2A-AFE1-B189B8BD623A</uuid>
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</uri>
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<subSubSection id="C3396548FFA9FFB480E6FC39FF2DF907" lastPageId="4" lastPageNumber="375" pageId="1" pageNumber="372" type="materials_examined">
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<paragraph id="8B9C36C3FFA9FFB180E6FC39FCCBFC0E" blockId="1.[151,819,977,1002]" box="[151,819,977,1002]" pageId="1" pageNumber="372">
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<emphasis id="B957EAD1FFA9FFB180E6FC39FEDBFC0E" bold="true" box="[151,291,977,1002]" pageId="1" pageNumber="372">
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<typeStatus id="54988861FFA9FFB180E6FC39FF34FC0D" box="[151,204,977,1001]" pageId="1" pageNumber="372">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C234D40FFA9FFB1815BFC3AFE03FC0D" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis & Hightower. Not" authorityYear="2019" box="[298,507,977,1001]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="monodontus" status="sp. nov.">
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<emphasis id="B957EAD1FFA9FFB1815BFC3AFE03FC0D" box="[298,507,977,1001]" italics="true" pageId="1" pageNumber="372">Peripus monodontus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26457AAFFA9FFB18276FC3AFD7CFC0E" box="[519,644,977,1002]" pageId="1" pageNumber="372" rank="species">new species</taxonomicNameLabel>
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, here designated.
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</paragraph>
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<paragraph id="8B9C36C3FFA9FFB180E6FBF9FD6AF9C1" blockId="1.[151,1437,1041,2041]" pageId="1" pageNumber="372">
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<emphasis id="B957EAD1FFA9FFB180E6FBF9FEEAFBC8" bold="true" box="[151,274,1041,1068]" pageId="1" pageNumber="372">Diagnosis.</emphasis>
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<taxonomicName id="4C234D40FFA9FFB1816DFBFBFE8BFBC8" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[284,371,1043,1068]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B957EAD1FFA9FFB1816DFBFBFE8BFBC8" box="[284,371,1043,1068]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
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</taxonomicName>
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belongs to the
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<taxonomicName id="4C234D40FFA9FFB1825DFBFBFD69FBC8" box="[556,657,1043,1068]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1FFA9FFB1825DFBFBFD69FBC8" box="[556,657,1043,1068]" italics="true" pageId="1" pageNumber="372">Isanopus</emphasis>
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</taxonomicName>
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group of genera of Xanthopygina (
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<bibRefCitation id="EFB24B32FFA9FFB18441FBFAFA6DFBC8" author="Chatzimanolis, S. & Brunke, A. J." box="[1072,1429,1042,1068]" pageId="1" pageNumber="372" pagination="494 - 506" refId="ref7073" refString="Chatzimanolis, S. & Brunke, A. J. (2019) A phylogeny of Xanthopygina (Insecta: Coleoptera) reveals major lineages and the origin of myrmecophily. Zoologica Scripta, 48, 494 - 506. https: // doi. org / 10.1111 / zsc. 12358" type="journal article" year="2019">Chatzimanolis & Brunke 2019</bibRefCitation>
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) based on the following characteristics: antennomeres 8–10 elongate or quadrate (
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<figureCitation id="13182A46FFA9FFB1846AFBDEFB96FBB4" box="[1051,1134,1078,1104]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="1" pageNumber="372">Fig. 12</figureCitation>
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); basal transverse carina of sternum 3 (
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<figureCitation id="13182A46FFA9FFB18167FBB2FE90FB90" box="[278,360,1114,1140]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Fig. 17</figureCitation>
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) acutely pointed medially; mesocoxae moderately to strongly separated (
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<figureCitation id="13182A46FFA9FFB184EEFBB2FB0AFB91" box="[1183,1266,1114,1141]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Fig. 16</figureCitation>
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); lack of dense meshed microsculpture anteriolaterally on sterna 5–7 (
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<figureCitation id="13182A46FFA9FFB18377FB96FC69FB7C" box="[774,913,1150,1176]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Figs. 18–19</figureCitation>
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). The sister group relationship between
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<taxonomicName id="4C234D40FFA9FFB1852DFB96FF1BFB58" authorityName="Chatzimanolis" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1FFA9FFB1852DFB96FF1BFB58" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C234D40FFA9FFB1816AFB4BFE8AFB58" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[283,370,1187,1212]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B957EAD1FFA9FFB1816AFB4BFE8AFB58" box="[283,370,1187,1212]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
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</taxonomicName>
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is supported by these characteristics: anterior margin of postclypeus slightly emarginate; basal transverse carina of sternum 4 (
|
||
<figureCitation id="13182A46FFA9FFB18189FB2EFDB2FB04" box="[504,586,1222,1248]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Fig. 17</figureCitation>
|
||
) produced medially; sterna 5–7 (
|
||
<figureCitation id="13182A46FFA9FFB183CCFB2EFBBBFB04" box="[957,1091,1222,1248]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Figs. 18–19</figureCitation>
|
||
) anteriorly with longitudinally elongate punctures and posteriorly with rounded punctures; labial P
|
||
<subScript id="17A73486FFA9FFB183E1FB14FC60FAEE" attach="left" box="[912,920,1276,1290]" fontSize="6" pageId="1" pageNumber="372">3</subScript>
|
||
dilated (securiform). The following two characteristics can distinguish
|
||
<taxonomicName id="4C234D40FFA9FFB181CFFAE7FDEDFACC" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[446,533,1295,1320]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFA9FFB181CFFAE7FDEDFACC" box="[446,533,1295,1320]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C234D40FFA9FFB1822BFAE6FD25FACC" authorityName="Chatzimanolis" authorityYear="2012" box="[602,733,1294,1320]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB1822BFAE6FD25FACC" box="[602,733,1294,1320]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
and are putative synapomorphies for
|
||
<taxonomicName id="4C234D40FFA9FFB184F5FAE7FB23FACC" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[1156,1243,1295,1320]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFA9FFB184F5FAE7FB23FACC" box="[1156,1243,1295,1320]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
|
||
</taxonomicName>
|
||
: basal transverse carina of sternum 5 (
|
||
<figureCitation id="13182A46FFA9FFB181F1FADAFE2AFAA8" box="[384,466,1330,1356]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="1" pageNumber="372">Fig. 18</figureCitation>
|
||
) medially produced (not seen in the species of
|
||
<taxonomicName id="4C234D40FFA9FFB18395FADAFB9FFAA8" authorityName="Chatzimanolis" authorityYear="2012" box="[996,1127,1330,1356]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18395FADAFB9FFAA8" box="[996,1127,1330,1356]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
examined); and securiform labial P
|
||
<subScript id="17A73486FFA9FFB1809CFA80FF0DFA92" attach="left" box="[237,245,1384,1398]" fontSize="6" pageId="1" pageNumber="372">3</subScript>
|
||
with unique sensory setae orientation: distally with 2–3 rows of setae (as in
|
||
<taxonomicName id="4C234D40FFA9FFB18410FABEFB1CFA94" authorityName="Chatzimanolis" authorityYear="2012" box="[1121,1252,1366,1392]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18410FABEFB1CFA94" box="[1121,1252,1366,1392]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
) but proximally expanding to 4–6 rows (
|
||
<figureCitation id="13182A46FFA9FFB181DBFA92FE04FA70" box="[426,508,1402,1428]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="1" pageNumber="372">Fig. 11</figureCitation>
|
||
). Additionally, male
|
||
<taxonomicName id="4C234D40FFA9FFB18298FA92FC94FA70" authorityName="Chatzimanolis" authorityYear="2012" box="[745,876,1402,1428]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18298FA92FC94FA70" box="[745,876,1402,1428]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
specimens can be easily distinguished from male
|
||
<taxonomicName id="4C234D40FFA9FFB180E6FA77FF16FA5C" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[151,238,1439,1464]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFA9FFB180E6FA77FF16FA5C" box="[151,238,1439,1464]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
|
||
</taxonomicName>
|
||
specimens based on the elongate 11 antennomere seen in
|
||
<taxonomicName id="4C234D40FFA9FFB18308FA76FC04FA5C" authorityName="Chatzimanolis" authorityYear="2012" box="[889,1020,1438,1464]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18308FA76FC04FA5C" box="[889,1020,1438,1464]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
. Distinguishing female specimens of
|
||
<taxonomicName id="4C234D40FFA9FFB180E6FA2BFF16FA38" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[151,238,1475,1500]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFA9FFB180E6FA2BFF16FA38" box="[151,238,1475,1500]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C234D40FFA9FFB18157FA2AFE51FA38" authorityName="Chatzimanolis" authorityYear="2012" box="[294,425,1474,1500]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18157FA2AFE51FA38" box="[294,425,1474,1500]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
can be slightly more challenging, especially from
|
||
<collectingCountry id="F3347653FFA9FFB18397FA2BFBE4FA39" box="[998,1052,1475,1501]" name="Peru" pageId="1" pageNumber="372">Peru</collectingCountry>
|
||
, where species belonging in these genera tend to have the same coloration. Besides the characters given above,
|
||
<taxonomicName id="4C234D40FFA9FFB18382FA0EFB8EF9E4" authorityName="Chatzimanolis" authorityYear="2012" box="[1011,1142,1510,1536]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFA9FFB18382FA0EFB8EF9E4" box="[1011,1142,1510,1536]" italics="true" pageId="1" pageNumber="372">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
species tend to be slightly larger and more robust than those of
|
||
<taxonomicName id="4C234D40FFA9FFB18245F9E3FD73F9C0" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[564,651,1547,1572]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFA9FFB18245F9E3FD73F9C0" box="[564,651,1547,1572]" italics="true" pageId="1" pageNumber="372">Peripus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFA9FFB180B6F9C5FDB4F974" blockId="1.[151,1437,1041,2041]" pageId="1" pageNumber="372">
|
||
<emphasis id="B957EAD1FFA9FFB180B6F9C5FEA1F9AC" bold="true" box="[199,345,1581,1608]" pageId="1" pageNumber="372">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="13182A46FFA9FFB18184F9C6FD98F9AC" box="[501,608,1582,1608]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1942,1967]" captionTargetBox="[253,1334,170,1922]" captionTargetId="figure@2.[239,1349,166,1922]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus photographs of species of Peripus Chatzimanolis & Hightower. 1. The holotype of P. brunkei Chatzimanolis & Hightower. 2. Peripus madrededios Chatzimanolis & Hightower. 3. Peripus didontus Chatzimanolis & Hightower. 4. Peripus monodontus Chatzimanolis & Hightower. Not to scale." httpUri="https://zenodo.org/record/3354778/files/figure.png" pageId="1" pageNumber="372">Figs. 1–4</figureCitation>
|
||
. Body medium-sized, forebody
|
||
<quantity id="4CDB9B26FFA9FFB183B5F9C5FBB2F9AD" box="[964,1098,1581,1609]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="1" pageNumber="372" unit="mm" value="4.85" valueMax="5.5" valueMin="4.2">4.2–5.5 mm</quantity>
|
||
long; without long bristle-like setae. Coloration of head and pronotum dark metallic blue purple or brown-blue purple; elytra metallic brown-green or blue-green; ventral forebody brown.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFA9FFB180B6F972FA9EF8A1" blockId="1.[151,1437,1041,2041]" pageId="1" pageNumber="372">
|
||
Head (
|
||
<figureCitation id="13182A46FFA9FFB18160F972FE83F950" box="[273,379,1689,1716]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[151,1429,196,1890]" captionTargetId="figure@3.[151,1436,196,1890]" captionTargetPageId="3" captionText="FIGURES 5–8. Heads and pronota of Peripus Chatzimanolis & Hightower. 5. Peripus brunkei Chatzimanolis & Hightower. 6. Peripus didontus Chatzimanolis & Hightower. 7. Peripus madrededios Chatzimanolis & Hightower. 8. Peripus monodontus Chatzimanolis & Hightower. Not to scale." httpUri="https://zenodo.org/record/3354780/files/figure.png" pageId="1" pageNumber="372">Figs. 5–8</figureCitation>
|
||
) shape rectangular; head length in comparison to pronotum shorter to subequal. Eye size large, more than 3/4 length of head. Postclypeus, in comparison to frons not deflexed; anterior margin slightly emarginate. Middle of epicranium impunctate but with microsculpture. Postmandibular ridge laterally; with deep punctures demarcating raised postmandibular ridge dorsolaterally present. Gular sutures not joined before neck, extended close to each other at base of head capsule. Nuchal ridge incomplete dorsally. Neck disc punctures absent or sparse.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFA9FFB180B6F8A6FC02F81C" blockId="1.[151,1437,1041,2041]" pageId="1" pageNumber="372">
|
||
Antennae (
|
||
<figureCitation id="13182A46FFA9FFB18132F8A6FE6EF88C" box="[323,406,1870,1896]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="1" pageNumber="372">Fig. 12</figureCitation>
|
||
) with antennomere 1 same width or slightly wider than 2. Antennomeres 3: three times as long as wide; antennomere 4 with tomentose pubescence; antennomeres 4–10 cylindrical in shape; antennomeres 4–7 longer than wide; antennomeres 8–10 symmetrical in shape, quadrate to elongate (longer than wide); antennomere 6 with curved, distinctly longer and thicker subapical setae than other macrosetae, forming circlet. Antennomeres 5–10 without club; antennomere
|
||
<quantity id="4CDB9B26FFA9FFB1821EF836FD5FF81C" box="[623,679,2014,2040]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="372" unit="in" value="11.0">11 in</quantity>
|
||
males slightly longer than 10.
|
||
</paragraph>
|
||
<caption id="DF5C664BFFAAFFB280E6F87EFCCAF813" httpUri="https://zenodo.org/record/3354778/files/figure.png" pageId="2" pageNumber="373" startId="2.[151,264,1942,1967]" targetBox="[253,1334,170,1922]" targetPageId="2">
|
||
<paragraph id="8B9C36C3FFAAFFB280E6F87EFCCAF813" blockId="2.[151,1437,1942,2039]" pageId="2" pageNumber="373">
|
||
<emphasis id="B957EAD1FFAAFFB280E6F87EFEC3F84B" bold="true" box="[151,315,1942,1967]" pageId="2" pageNumber="373">FIGURES 1–4.</emphasis>
|
||
Habitus photographs of species of
|
||
<taxonomicName id="4C234D40FFAAFFB282D3F87FFD09F84A" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[674,753,1943,1966]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFAAFFB282D3F87FFD09F84A" box="[674,753,1943,1966]" italics="true" pageId="2" pageNumber="373">Peripus</emphasis>
|
||
</taxonomicName>
|
||
Chatzimanolis & Hightower. 1. The holotype of
|
||
<taxonomicName id="4C234D40FFAAFFB28494F87FFE84F837" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="brunkei">
|
||
<emphasis id="B957EAD1FFAAFFB28494F87FFAB3F84B" box="[1253,1355,1943,1967]" italics="true" pageId="2" pageNumber="373">P. brunkei</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
2.
|
||
<taxonomicName id="4C234D40FFAAFFB281EDF853FC5AF837" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." box="[412,930,1978,2003]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="madrededios">
|
||
<emphasis id="B957EAD1FFAAFFB281EDF853FD8CF837" box="[412,628,1979,2003]" italics="true" pageId="2" pageNumber="373">Peripus madrededios</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
3.
|
||
<taxonomicName id="4C234D40FFAAFFB283B3F853FA65F837" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." box="[962,1437,1978,2003]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="didontus">
|
||
<emphasis id="B957EAD1FFAAFFB283B3F853FB97F837" box="[962,1135,1979,2003]" italics="true" pageId="2" pageNumber="373">Peripus didontus</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
4.
|
||
<taxonomicName id="4C234D40FFAAFFB280C0F837FD20F812" authority="Chatzimanolis & Hightower. Not" authorityName="Chatzimanolis & Hightower. Not" authorityYear="2019" box="[177,728,2014,2039]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="monodontus">
|
||
<emphasis id="B957EAD1FFAAFFB280C0F837FE7AF813" box="[177,386,2015,2039]" italics="true" pageId="2" pageNumber="373">Peripus monodontus</emphasis>
|
||
Chatzimanolis & Hightower. Not
|
||
</taxonomicName>
|
||
to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5C664BFFABFFB380E6F86FFDBAF80C" httpUri="https://zenodo.org/record/3354780/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,264,1927,1952]" targetBox="[151,1429,196,1890]" targetPageId="3">
|
||
<paragraph id="8B9C36C3FFABFFB380E6F86FFDBAF80C" blockId="3.[151,1437,1927,2024]" pageId="3" pageNumber="374">
|
||
<emphasis id="B957EAD1FFABFFB380E6F86FFEC4F844" bold="true" box="[151,316,1927,1952]" pageId="3" pageNumber="374">FIGURES 5–8.</emphasis>
|
||
Heads and pronota of
|
||
<taxonomicName id="4C234D40FFABFFB38258F860FD80F87B" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[553,632,1928,1951]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFABFFB38258F860FD80F87B" box="[553,632,1928,1951]" italics="true" pageId="3" pageNumber="374">Peripus</emphasis>
|
||
</taxonomicName>
|
||
Chatzimanolis & Hightower. 5.
|
||
<taxonomicName id="4C234D40FFABFFB383B8F860FA65F844" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." box="[969,1437,1927,1952]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="brunkei">
|
||
<emphasis id="B957EAD1FFABFFB383B8F860FB95F87B" box="[969,1133,1927,1951]" italics="true" pageId="3" pageNumber="374">Peripus brunkei</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
6.
|
||
<taxonomicName id="4C234D40FFABFFB380C0F844FD71F820" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." box="[177,649,1963,1988]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="didontus">
|
||
<emphasis id="B957EAD1FFABFFB380C0F844FEA5F827" box="[177,349,1963,1987]" italics="true" pageId="3" pageNumber="374">Peripus didontus</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
7.
|
||
<taxonomicName id="4C234D40FFABFFB382D8F844FB54F820" authority="Chatzimanolis & Hightower." authorityName="Chatzimanolis & Hightower." box="[681,1196,1963,1988]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="madrededios">
|
||
<emphasis id="B957EAD1FFABFFB382D8F844FC78F827" box="[681,896,1963,1987]" italics="true" pageId="3" pageNumber="374">Peripus madrededios</emphasis>
|
||
Chatzimanolis & Hightower.
|
||
</taxonomicName>
|
||
8.
|
||
<taxonomicName id="4C234D40FFABFFB384BAF844FE10F803" authority="Chatzimanolis & Hightower. Not" authorityName="Chatzimanolis & Hightower. Not" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="monodontus">
|
||
<emphasis id="B957EAD1FFABFFB384BAF844FA64F827" box="[1227,1436,1963,1987]" italics="true" pageId="3" pageNumber="374">Peripus monodontus</emphasis>
|
||
Chatzimanolis & Hightower. Not
|
||
</taxonomicName>
|
||
to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FF71FDC0FE4B" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Mouthparts with labrum (
|
||
<figureCitation id="13182A46FFACFFB4819DFF71FDC7FF57" box="[492,575,153,179]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="4" pageNumber="375">Fig. 14</figureCitation>
|
||
) having broad U-shaped emargination, lobes strongly separated. Mandibles (
|
||
<figureCitation id="13182A46FFACFFB480EEFF55FEE1FF33" box="[159,281,189,215]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="4" pageNumber="375">Figs. 9–10</figureCitation>
|
||
) with relative length typical (i.e. closed mandible not extending beyond lateral margin of head); without asymmetrical torsion. Mandibles in dorsal view curved from apical half; in lateral view straight; left and right mandibles each with one tooth. Maxilla (
|
||
<figureCitation id="13182A46FFACFFB4821FFEEDFD38FEFB" box="[622,704,261,287]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="4" pageNumber="375">Fig. 13</figureCitation>
|
||
) with galea much shorter than palpus; maxillary palpus with P
|
||
<subScript id="17A73486FFACFFB485E5FEFEFA64FEC0" attach="left" box="[1428,1436,278,292]" fontSize="6" pageId="4" pageNumber="375">3</subScript>
|
||
distinctly shorter than P
|
||
<subScript id="17A73486FFACFFB481D5FED2FE54FEAC" attach="left" box="[420,428,314,328]" fontSize="6" pageId="4" pageNumber="375">2</subScript>
|
||
; P
|
||
<subScript id="17A73486FFACFFB481BDFED2FE2CFEAC" attach="left" box="[460,468,314,328]" fontSize="6" pageId="4" pageNumber="375">4</subScript>
|
||
distinctly longer than P
|
||
<subScript id="17A73486FFACFFB48293FED2FD12FEAC" attach="left" box="[738,746,314,328]" fontSize="6" pageId="4" pageNumber="375">3</subScript>
|
||
; P
|
||
<subScript id="17A73486FFACFFB4837BFED2FCEAFEAC" attach="left" box="[778,786,314,328]" fontSize="6" pageId="4" pageNumber="375">4</subScript>
|
||
not dilated. Hypopharynx and labial palpus as in
|
||
<figureCitation id="13182A46FFACFFB48534FEC1FA6EFEA7" box="[1349,1430,297,323]" captionStart="FIGURES 9–14" captionStartId="7.[151,264,1890,1915]" captionTargetBox="[151,1436,793,1871]" captionTargetId="figure@7.[151,1436,793,1871]" captionTargetPageId="7" captionText="FIGURES 9–14. Peripus monodontus Chatzimanolis & Hightower. 9. Dorsal view of left mandible, scale bar = 0.5 mm. 10. Dorsal view of right mandible, scale bar = 0.5 mm. 11. Hypopharynx and labial palps, scale bar = 0.35 mm. 12. Antenna, scale bar = 0.57 mm. 13. Maxilla, scale bar = 0.32 mm. 14. Labrum, scale bar = 0.4 mm." httpUri="https://zenodo.org/record/3354782/files/figure.png" pageId="4" pageNumber="375">Fig. 11</figureCitation>
|
||
; labial palpus with P
|
||
<subScript id="17A73486FFACFFB48108FEB6FE79FE88" attach="left" box="[377,385,350,364]" fontSize="6" pageId="4" pageNumber="375">3</subScript>
|
||
securiform, starting with 2–3 rows of setae as in
|
||
<taxonomicName id="4C234D40FFACFFB483C8FEA5FBC4FE83" authorityName="Chatzimanolis" authorityYear="2012" box="[953,1084,333,359]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFACFFB483C8FEA5FBC4FE83" box="[953,1084,333,359]" italics="true" pageId="4" pageNumber="375">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
but expanding to 4–6 rows; P
|
||
<subScript id="17A73486FFACFFB485E5FEB6FA64FE88" attach="left" box="[1428,1436,350,364]" fontSize="6" pageId="4" pageNumber="375">3</subScript>
|
||
without long dense setae on lateral sides. Ligula shape small, entire. Mentum with alpha seta present; hypostomal cavity present, moderately delimited.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FE51FC1CFE13" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Pronotum (
|
||
<figureCitation id="13182A46FFACFFB48136FE51FE4CFE37" box="[327,436,440,467]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[151,1429,196,1890]" captionTargetId="figure@3.[151,1436,196,1890]" captionTargetPageId="3" captionText="FIGURES 5–8. Heads and pronota of Peripus Chatzimanolis & Hightower. 5. Peripus brunkei Chatzimanolis & Hightower. 6. Peripus didontus Chatzimanolis & Hightower. 7. Peripus madrededios Chatzimanolis & Hightower. 8. Peripus monodontus Chatzimanolis & Hightower. Not to scale." httpUri="https://zenodo.org/record/3354780/files/figure.png" pageId="4" pageNumber="375">Figs. 5–8</figureCitation>
|
||
) with shape of lateral margins in dorsal view posteriad of midpoint straight to sinuate; anterior angles in dorsal view not strongly acuminate and produced laterad.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FDE9FBE9FCF3" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Pronotum near anterolateral angles without raised impunctate spots; anterolateral corners with punctation; disc of pronotum with punctation beyond midlength; punctation various but with multiple rows of large punctures; with microsculpture; without coarse punctures impressed in flange at posterior angle of pronotum; impunctate area in middle expanded. Hypomeron (
|
||
<figureCitation id="13182A46FFACFFB48271FD85FDABFD63" box="[512,595,620,647]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Fig. 15</figureCitation>
|
||
) with superior marginal line continuous to anterior margin; superior marginal line without distinct deflection under anterior angles in ventral view; inferior marginal line of hypomeron continued as separate entity beyond anterior pronotal angles and curving around them. Postcoxal process absent. Basisternum slightly longer than furcasternum; basisternum with pair of macrosetae situated far from anterior margin of prosternum; furcasternum with medial elongate carina pointed vertically.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FCC9FB9FFC2F" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Elytra (
|
||
<figureCitation id="13182A46FFACFFB4816FFCC9FE8AFCDF" box="[286,370,801,827]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Fig. 20</figureCitation>
|
||
) without contiguous polygon-shaped meshed microsculpture or with patches of white setae. Elytral setae not reduced, easily seen at low magnification. Mesoscutellum with dense cluster of punctures medially. Mesoventrite (
|
||
<figureCitation id="13182A46FFACFFB4814BFC81FE74FC60" box="[314,396,873,900]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Fig. 16</figureCitation>
|
||
) with anterior margin forming “lip”; with median carina and mesoventral process triangular; process extended distally to distance about 2/5 between mesocoxae. Metaventrite (
|
||
<figureCitation id="13182A46FFACFFB48439FC65FB63FC4C" box="[1096,1179,909,936]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Fig. 16</figureCitation>
|
||
) with large punctures; metaventral processes, small, rounded, triangular, extended to beginning of metacoxae.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FC3DFCC7FB47" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Legs with tarsal segmentation 5-5-5; profemora without lateroventral apical spine; protarsi with modified pale (adhesive) setae ventrally; tarsomeres 1–4 of protarsi dorsoventrally flattened. Mesocoxae (
|
||
<figureCitation id="13182A46FFACFFB484EDFC11FB16FBF0" box="[1180,1262,1017,1044]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Fig. 16</figureCitation>
|
||
) strongly separated, intercoxal area on approximately same plane as both meso and metaventrital processes; metatibiae with spines. Metacoxae without coxal shield; metatibia without thick and long apical spurs but smaller spurs present; tarsomeres 3–5 of metatarsi with chaetotaxy developed only at margins of dorsal surface, dorsal surface of tarsomeres glabrous along midline. Pretarsal claws with empodial setae.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FB45FD0BFA03" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Abdomen (
|
||
<figureCitation id="13182A46FFACFFB48137FB45FE36FB23" box="[326,462,1197,1223]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[151,1436,776,1817]" captionTargetId="figure@8.[151,1436,776,1817]" captionTargetPageId="8" captionText="FIGURES 15–20. Peripus monodontus Chatzimanolis & Hightower. 15. Oblique view of prosternum and pronotal hypomeron, scale bar = 0.78 mm. 16. Meso- and metaventrite, scale bar = 1.19 mm. 17. Abdominal terga 3 and 4, scale bar = 1.15 mm. 18. Abdominal terga 5 and 6, scale bar = 1.06 mm. 19. Abdominal terga 7 and 8, scale bar = 0.90 mm. 20. Elytron, scale bar = 1.25 mm." httpUri="https://zenodo.org/record/3354784/files/figure.png" pageId="4" pageNumber="375">Figs. 17–19</figureCitation>
|
||
), with protergal glands having well-developed acetabula. Anterior basal transverse carina on terga 3 and 4 without pair of accessory ridges; tergum 3 without posterior basal transverse carina and without curved (arch-like) carina on disc; tergum 5 without pair of accessory ridges on anterior basal transverse carina; center of tergum 5 punctate; posterior half of tergum
|
||
<quantity id="4CDB9B26FFACFFB48293FAF0FCF4FAD7" box="[738,780,1304,1331]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="375" unit="in" value="5.0">5 in</quantity>
|
||
lateral view not appearing bulging. Sternum 3 with acutely pointed basal transverse carina medially; basal transverse carina laterally not sinuate. Basal transverse carina point- ed medially on sterna 4 and 5. Sterna 5–7 anteriorly with longitudinally elongate punctures and posteriorly with rounded punctures; polygon-shaped microsculpture present in grooves of longitudinally elongate punctures. Males with secondary sexual structures (emargination medially) on sterna 7 and 8; without porose structure on sternum 7. Females without obvious secondary sexual structures.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6FA19FEE0F9B7" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
Aedeagus as
|
||
<figureCitation id="13182A46FFACFFB48128FA19FE18F9EF" box="[345,480,1521,1547]" captionStart-0="FIGURES 21–23" captionStart-1="FIGURES 24–26" captionStart-2="FIGURES 27–29" captionStart-3="FIGURES 30–32" captionStartId-0="9.[151,264,1082,1107]" captionStartId-1="9.[151,264,1935,1960]" captionStartId-2="10.[151,264,936,961]" captionStartId-3="10.[151,264,1762,1787]" captionTargetBox-0="[151,1425,344,1040]" captionTargetBox-1="[151,1425,1185,1885]" captionTargetBox-2="[151,1425,181,905]" captionTargetBox-3="[151,1436,1019,1743]" captionTargetId-0="figure@9.[151,1436,344,1051]" captionTargetId-1="figure@9.[151,1436,1178,1896]" captionTargetId-2="figure@10.[151,1436,181,916]" captionTargetId-3="figure@10.[151,1436,1019,1743]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="10" captionText-0="FIGURES 21–23. Aedeagus of P. brunkei Chatzimanolis & Hightower. 21. Dorsal view. 22. Lateral view. 23. Detail of paramere, ventral view." captionText-1="FIGURES 24–26. Aedeagus of P. didontus Chatzimanolis & Hightower. 24. Dorsal view. 25. Lateral view. 26. Detail of paramere, ventral view." captionText-2="FIGURES 27–29. Aedeagus of P. madrededios Chatzimanolis & Hightower. 27. Dorsal view. 28. Lateral view. 29. Detail of paramere, ventral view." captionText-3="FIGURES 30–32. Aedeagus of P. monodontus Chatzimanolis & Hightower. 30. Dorsal view. 31. Lateral view. 32. Detail of paramere, ventral view." httpUri-0="https://zenodo.org/record/3354786/files/figure.png" httpUri-1="https://zenodo.org/record/3354788/files/figure.png" httpUri-2="https://zenodo.org/record/3354792/files/figure.png" httpUri-3="https://zenodo.org/record/3354794/files/figure.png" pageId="4" pageNumber="375">Figs. 21–32</figureCitation>
|
||
; with long median lobe and single paramere; paramere with sensory peg setae; median lobe with single or pair of subapical teeth; median lobe without apical tooth, carina or paired apex. Spermatheca not sclerotized.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6F9B5FA69F97F" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
<emphasis id="B957EAD1FFACFFB480B6F9B5FEB6F993" bold="true" box="[199,334,1629,1655]" pageId="4" pageNumber="375">Etymology.</emphasis>
|
||
The name is derived from the Greek word “περίπου” (about), and refers to the fact that the first author of this paper thought for a long time that
|
||
<taxonomicName id="4C234D40FFACFFB482C2F96AFCF2F97F" authority="Chatzimanolis, 2019" authorityName="Chatzimanolis" authorityYear="2019" box="[691,778,1666,1691]" class="Insecta" family="Staphylinidae" genus="Peripus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B957EAD1FFACFFB482C2F96AFCF2F97F" box="[691,778,1666,1691]" italics="true" pageId="4" pageNumber="375">Peripus</emphasis>
|
||
</taxonomicName>
|
||
is about the same as
|
||
<taxonomicName id="4C234D40FFACFFB48389F969FB83F97F" authorityName="Chatzimanolis" authorityYear="2012" box="[1016,1147,1665,1691]" class="Insecta" family="Staphylinidae" genus="Zackfalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAD1FFACFFB48389F969FB83F97F" box="[1016,1147,1665,1691]" italics="true" pageId="4" pageNumber="375">Zackfalinus</emphasis>
|
||
</taxonomicName>
|
||
. The name is masculine.
|
||
</paragraph>
|
||
<paragraph id="8B9C36C3FFACFFB480B6F94CFF2DF907" blockId="4.[151,1437,153,1763]" pageId="4" pageNumber="375">
|
||
<emphasis id="B957EAD1FFACFFB480B6F94CFED2F95B" bold="true" box="[199,298,1700,1727]" pageId="4" pageNumber="375">Habitat.</emphasis>
|
||
Collected in lowland tropical rainforests (elevations
|
||
<quantity id="4CDB9B26FFACFFB483EBF94CFBFBF95B" box="[922,1027,1700,1727]" metricMagnitude="2" metricUnit="m" metricValue="3.025" metricValueMax="6.0" metricValueMin="0.05" pageId="4" pageNumber="375" unit="m" value="302.5" valueMax="600.0" valueMin="5.0">5–600 m</quantity>
|
||
) using flight intercept and malaise traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |