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<mods:title id="256A5A33FB07BF204C5A39B5B6C11507">Five new species of the genus Philydrodes Bernhauer, 1929 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="60520268B8EC4D7D262D7E6E556BCF4E">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="B980DAFD33EE4C14A0DE4D3A237203DB">2017</mods:date>
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<treatment id="038A87ABFFE2B93AFF415AA3869299F2" ID-DOI="http://doi.org/10.5281/zenodo.6029490" ID-GBIF-Taxon="127694604" ID-Zenodo-Dep="6029490" LSID="urn:lsid:plazi:treatment:038A87ABFFE2B93AFF415AA3869299F2" httpUri="http://treatment.plazi.org/id/038A87ABFFE2B93AFF415AA3869299F2" lastPageId="13" lastPageNumber="182" pageId="11" pageNumber="180">
<subSubSection id="C3396536FFE2B93CFF415AA387449EC3" pageId="11" pageNumber="180" type="nomenclature">
<paragraph id="8B9C36BDFFE2B93CFF415AA386169EE2" blockId="11.[151,841,1473,1533]" box="[151,841,1473,1500]" pageId="11" pageNumber="180">
<heading id="D0D481D1FFE2B93CFF415AA386169EE2" bold="true" box="[151,841,1473,1500]" fontSize="11" level="1" pageId="11" pageNumber="180" reason="1">
<emphasis id="B957EAAFFFE2B93CFF415AA386169EE2" bold="true" box="[151,841,1473,1500]" pageId="11" pageNumber="180">
<taxonomicName id="4C234D3EFFE2B93CFF415AA3865E9EE2" ID-CoL="4GDP3" authority="Shavrin" authorityName="Shavrin" box="[151,769,1473,1500]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov." subGenus="Minyphilydrodes">
<emphasis id="B957EAAFFFE2B93CFF415AA384799EE5" bold="true" box="[151,294,1473,1499]" italics="true" pageId="11" pageNumber="180">Philydrodes</emphasis>
(
<emphasis id="B957EAAFFFE2B93CFEE15AA3875D9EE5" bold="true" box="[311,514,1473,1499]" italics="true" pageId="11" pageNumber="180">Minyphilydrodes</emphasis>
)
<emphasis id="B957EAAFFFE2B93CFDC25AA387CA9EE5" bold="true" box="[532,661,1473,1499]" italics="true" pageId="11" pageNumber="180">reticulatus</emphasis>
Shavrin
</taxonomicName>
,
<taxonomicNameLabel id="A26457D4FFE2B93CFCDB5AA086169EE2" box="[781,841,1474,1500]" pageId="11" pageNumber="180" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36BDFFE2B93CFF415A8687449EC3" blockId="11.[151,841,1473,1533]" box="[151,539,1508,1533]" pageId="11" pageNumber="180">
(
<figureCitation id="13182A38FFE2B93CFF495A8685B29EC3" box="[159,237,1508,1533]" captionStart="FIGURES 1 5" captionStartId="4.[151,264,1968,1991]" captionTargetBox="[167,1428,711,1924]" captionTargetId="figure@4.[150,1436,697,1947]" captionTargetPageId="4" captionText="FIGURES 1 5. Habitus of Philydrodes: 1 — Ph. tridentatus, holotype, 2 — Ph. michaeli, paratype, 3 — Ph. reticulatus, paratype, 4 — Ph. tibetanus, holotype, 5 — Ph. wrasei, paratype. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/291271/files/figure.png" pageId="11" pageNumber="180">Figs. 3</figureCitation>
,
<figureCitation id="13182A38FFE2B93CFF2C5A87841C9EC3" box="[250,323,1509,1533]" captionStart="FIGURES 36 41" captionStartId="9.[151,264,1968,1991]" captionTargetBox="[259,1317,200,1934]" captionTargetId="figure@9.[237,1334,193,1945]" captionTargetPageId="9" captionText="FIGURES 36 41. Aedeagus of Philydrodes michaeli (36, 39: holotype), Ph. reticulatus (37, 40: paratype) and Ph. wrasei (38, 41: paratype): 36 ‾ 38 — parameral view, 39 ‾ 41 — lateral view. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291797/files/figure.png" pageId="11" pageNumber="180">37, 40</figureCitation>
, 4445, 5253, 58)
</paragraph>
</subSubSection>
<subSubSection id="C3396536FFE2B93BFF41594E814E9F8A" lastPageId="12" lastPageNumber="181" pageId="11" pageNumber="180" type="materials_examined">
<paragraph id="8B9C36BDFFE2B93CFF41594E84209DC7" blockId="11.[151,1437,1580,2001]" pageId="11" pageNumber="180">
<materialsCitation id="3B4B3CE0FFE2B93CFF41594E80599DB3" ID-GBIF-Occurrence="1427603795" collectorName="Mountain W &amp; S Xian" country="China" elevation="2400" latitude="33.51" location="Qin Ling Shan" longLatPrecision="723" longitude="108.47" municipality="Autorote" pageId="11" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="SHAANXI PROVINCE" typeStatus="holotype">
<emphasis id="B957EAAFFFE2B93CFF41594E84909D7B" bold="true" box="[151,463,1580,1605]" pageId="11" pageNumber="180">
<typeStatus id="5498881FFFE2B93CFF41594E858D9D7B" box="[151,210,1580,1605]" pageId="11" pageNumber="180">Type</typeStatus>
material examined:
</emphasis>
<typeStatus id="5498881FFFE2B93CFE0A594E871B9D7B" box="[476,580,1580,1605]" pageId="11" pageNumber="180" type="holotype">Holotype</typeStatus>
<specimenCount id="9D25FD34FFE2B93CFD85594F87379D7A" box="[595,616,1581,1604]" pageId="11" pageNumber="180" type="male"></specimenCount>
[The preparation of the aedeagus and apical abdominal segment in Canadian balsam was attached under the card with beetle]:
<emphasis id="B957EAAFFFE2B93CFCFB5932817D9D57" bold="true" box="[813,1058,1616,1641]" pageId="11" pageNumber="180">
<collectingRegion id="49E7F85FFFE2B93CFCFB593281479D59" box="[813,1048,1616,1641]" country="China" name="Shaanxi" pageId="11" pageNumber="180">SHAANXI PROVINCE</collectingRegion>
:
</emphasis>
<collectingCountry id="F334762DFFE2B93CFBFB593281269D57" box="[1069,1145,1616,1641]" name="China" pageId="11" pageNumber="180">China</collectingCountry>
:
<collectingRegion id="49E7F85FFFE2B93CFB55593281BF9D57" box="[1155,1248,1616,1641]" country="China" name="Shaanxi" pageId="11" pageNumber="180">Shaanxi</collectingRegion>
,
<location id="8EFC6066FFE2B93CFB3D593280CF9D57" LSID="urn:lsid:plazi:treatment:038A87ABFFE2B93AFF415AA3869299F2:8EFC6066FFE2B93CFB3D593280CF9D57" box="[1259,1424,1616,1641]" country="China" latitude="33.51" longLatPrecision="723" longitude="108.47" municipality="Autorote" name="Qin Ling Shan" pageId="11" pageNumber="180" stateProvince="SHAANXI PROVINCE">Qin Ling Shan</location>
|
<geoCoordinate id="EE17507AFFE2B93CFF415917845F9DB2" box="[151,256,1653,1677]" direction="east" orientation="longitude" pageId="11" pageNumber="180" precision="555" value="108.47">108.47 E</geoCoordinate>
,
<geoCoordinate id="EE17507AFFE2B93CFEDA591684349DB2" box="[268,363,1652,1677]" direction="north" orientation="latitude" pageId="11" pageNumber="180" precision="555" value="33.51">33.51 N</geoCoordinate>
,
<collectorName id="26D6536BFFE2B93CFEA1591687599DB2" box="[375,518,1652,1677]" pageId="11" pageNumber="180">Mountain W</collectorName>
| pass at
<collectingMunicipality id="6BF8ACC7FFE2B93CFDA75917878A9DB3" box="[625,725,1653,1677]" pageId="11" pageNumber="180">Autorote</collectingMunicipality>
km 70, 47 km |
<collectorName id="26D6536BFFE2B93CFC44591786BD9DB3" box="[914,994,1652,1677]" pageId="11" pageNumber="180">S Xian</collectorName>
,
<elevation id="000ED18EFFE2B93CFC39591681D99DB3" box="[1007,1158,1652,1677]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="11" pageNumber="180" unit="m" value="2400.0" valueMax="2500.0" valueMin="2300.0">
<quantity id="4CDB9B58FFE2B93CFC39591681D99DB3" box="[1007,1158,1652,1677]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="11" pageNumber="180" unit="m" value="2400.0" valueMax="2500.0" valueMin="2300.0">23002500m</quantity>
</elevation>
, sifted | 26.
</materialsCitation>
<date id="FF9D107DFFE2B93CFAC2591680C79DB3" box="[1300,1432,1652,1677]" pageId="11" pageNumber="180" value="1995-08-30">30.08.1995</date>
, leg. M. Schülke &lt;rectangular label, printed&gt;, Sammlung | M. Schülke | Berlin &lt;green rectangular label, printed&gt;,
<emphasis id="B957EAAFFFE2B93CFF7659DE84609DEB" bold="true" box="[160,319,1724,1749]" pageId="11" pageNumber="180">HOLOTYPE</emphasis>
|
<taxonomicName id="4C234D3EFFE2B93CFE8559DE87EF9DEB" authority="(Minyphilydrodes)" baseAuthorityName="Minyphilydrodes" box="[339,688,1724,1749]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAAFFFE2B93CFE8559DE84869DEB" box="[339,473,1724,1749]" italics="true" pageId="11" pageNumber="180">Philydrodes</emphasis>
(
<emphasis id="B957EAAFFFE2B93CFE3C59DE87F99DEB" box="[490,678,1724,1749]" italics="true" pageId="11" pageNumber="180">Minyphilydrodes</emphasis>
)
</taxonomicName>
|
<taxonomicName id="4C234D3EFFE2B93CFD1259DE86649DEB" box="[708,827,1724,1749]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="B957EAAFFFE2B93CFD1259DE86649DEB" box="[708,827,1724,1749]" italics="true" pageId="11" pageNumber="180">reticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457D4FFE2B93CFC9459DF86299DEA" box="[834,886,1725,1748]" pageId="11" pageNumber="180" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. des. 2016 &lt;red rectangular label, printed&gt; (MNHUB).
</paragraph>
<paragraph id="8B9C36BDFFE2B93CFF11586786619CB7" blockId="11.[151,1437,1580,2001]" pageId="11" pageNumber="180">
<typeStatus id="5498881FFFE2B93CFF11586784669C23" box="[199,313,1797,1821]" pageId="11" pageNumber="180" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D25FD34FFE2B93CFE925867842F9C22" box="[324,368,1797,1821]" pageId="11" pageNumber="180" type="male">1 ♂</specimenCount>
[The preparation of aedeagus, abdominal tergite VIII and sternite VIII, and apical abdominal segement in Canadian balsam was attached under the card with beetle],
<specimenCount id="9D25FD34FFE2B93CFC15584A86B29C7E" box="[963,1005,1832,1857]" pageId="11" pageNumber="180" type="female">5 ♀</specimenCount>
[The preparation of abdominal tergite VIII, sternite VIII and apical apical abdominal segment in Canadian balsam was attached under the card with beetle]: same data as the
<typeStatus id="5498881FFFE2B93CFE795812874E9CB7" box="[431,529,1904,1929]" pageId="11" pageNumber="180" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D25FD34FFE2B93CFDF7581387119CB6" box="[545,590,1905,1929]" pageId="11" pageNumber="180" type="female">4 ♀</specimenCount>
: cSch;
<specimenCount id="9D25FD34FFE2B93CFD76581387929CB6" box="[672,717,1905,1929]" pageId="11" pageNumber="180" type="male">1 ♂</specimenCount>
;
<specimenCount id="9D25FD34FFE2B93CFD0F5813865A9CB6" box="[729,773,1905,1929]" pageId="11" pageNumber="180" type="female">1 ♀</specimenCount>
: cS).
</paragraph>
<paragraph id="8B9C36BDFFE2B93CFF1158F6864A9CEF" blockId="11.[151,1437,1580,2001]" pageId="11" pageNumber="180">
All
<typeStatus id="5498881FFFE2B93CFF2058F7843D9C93" box="[246,354,1941,1965]" pageId="11" pageNumber="180" type="paratype">paratypes</typeStatus>
with my additional label:
<typeStatus id="5498881FFFE2B93CFD7058F686659C93" box="[678,826,1940,1965]" pageId="11" pageNumber="180" type="paratype">
<emphasis id="B957EAAFFFE2B93CFD7058F686659C93" bold="true" box="[678,826,1940,1965]" pageId="11" pageNumber="180">PARATYPE</emphasis>
</typeStatus>
|
<taxonomicName id="4C234D3EFFE2B93CFC8158F681E89C93" authority="(Minyphilydrodes)" baseAuthorityName="Minyphilydrodes" box="[855,1207,1940,1965]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAAFFFE2B93CFC8158F686829C93" box="[855,989,1940,1965]" italics="true" pageId="11" pageNumber="180">Philydrodes</emphasis>
(
<emphasis id="B957EAAFFFE2B93CFC2458F681F19C93" box="[1010,1198,1940,1965]" italics="true" pageId="11" pageNumber="180">Minyphilydrodes</emphasis>
)
</taxonomicName>
|
<taxonomicName id="4C234D3EFFE2B93CFB0258F680149C93" box="[1236,1355,1940,1965]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="B957EAAFFFE2B93CFB0258F680149C93" box="[1236,1355,1940,1965]" italics="true" pageId="11" pageNumber="180">reticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457D4FFE2B93CFA8058F780D49C92" box="[1366,1419,1941,1964]" pageId="11" pageNumber="180" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. des. 2016 &lt;red rectangular label, printed&gt;.
</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF415B5C81D59F4D" blockId="12.[151,1437,1086,1140]" pageId="12" pageNumber="181">
<emphasis id="B957EAAFFFE5B93BFF415B5C84089F6A" bold="true" box="[151,343,1086,1109]" pageId="12" pageNumber="181">FIGURES 5057.</emphasis>
Male (50, 52, 54, 56) and female (51, 53, 55, 57) abdominal sternites VIII of
<taxonomicName id="4C234D3EFFE5B93BFB405B5D80509F6B" box="[1174,1295,1087,1109]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAAFFFE5B93BFB405B5D80509F6B" box="[1174,1295,1087,1109]" italics="true" pageId="12" pageNumber="181">Philydrodes</emphasis>
</taxonomicName>
: 50‾51—
<taxonomicName id="4C234D3EFFE5B93BFAAF5B5D85B19F4D" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="181" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B957EAAFFFE5B93BFAAF5B5D85B19F4D" italics="true" pageId="12" pageNumber="181">
<collectingCountry id="F334762DFFE5B93BFAAF5B5D80C79F6B" box="[1401,1432,1087,1109]" name="Philippines" pageId="12" pageNumber="181">Ph</collectingCountry>
. michaeli
</emphasis>
</taxonomicName>
, 52‾53—
<taxonomicName id="4C234D3EFFE5B93BFE8D5B3F84AF9F4D" box="[347,496,1117,1139]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="181" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B957EAAFFFE5B93BFE8D5B3F84AF9F4D" box="[347,496,1117,1139]" italics="true" pageId="12" pageNumber="181">
<collectingCountry id="F334762DFFE5B93BFE8D5B3F84259F4D" box="[347,378,1117,1139]" name="Philippines" pageId="12" pageNumber="181">Ph</collectingCountry>
. reticulatus
</emphasis>
</taxonomicName>
, 54‾55—
<taxonomicName id="4C234D3EFFE5B93BFD8A5B3F87BD9F4D" box="[604,738,1117,1139]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="181" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="B957EAAFFFE5B93BFD8A5B3F87BD9F4D" box="[604,738,1117,1139]" italics="true" pageId="12" pageNumber="181">
<collectingCountry id="F334762DFFE5B93BFD8A5B3F87249F4D" box="[604,635,1117,1139]" name="Philippines" pageId="12" pageNumber="181">Ph</collectingCountry>
. tibetanus
</emphasis>
</taxonomicName>
, 56‾57—
<taxonomicName id="4C234D3EFFE5B93BFC995B3F86E39F4A" box="[847,956,1117,1140]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="181" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="B957EAAFFFE5B93BFC995B3F86E39F4A" box="[847,956,1117,1140]" italics="true" pageId="12" pageNumber="181">
<collectingCountry id="F334762DFFE5B93BFC995B3F86319F4D" box="[847,878,1117,1139]" name="Philippines" pageId="12" pageNumber="181">Ph</collectingCountry>
. wrasei
</emphasis>
</taxonomicName>
. Scale bars
<quantity id="4CDB9B58FFE5B93BFBE05B3F81D89F4D" box="[1078,1159,1117,1139]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="181" unit="mm" value="0.1">0.1 mm</quantity>
.
</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF115BF9814E9F8A" blockId="12.[151,1437,1179,2035]" box="[199,1041,1179,1204]" pageId="12" pageNumber="181">One specimen without abdomen was not included in the type series (cSch).</paragraph>
</subSubSection>
<subSubSection id="C3396536FFE5B93AFF115BDD84F99A92" lastPageId="13" lastPageNumber="182" pageId="12" pageNumber="181" type="description">
<paragraph id="8B9C36BDFFE5B93BFF115BDD84959E7A" blockId="12.[151,1437,1179,2035]" pageId="12" pageNumber="181">
<emphasis id="B957EAAFFFE5B93BFF115BDD84069FE6" bold="true" box="[199,345,1215,1240]" pageId="12" pageNumber="181">Description.</emphasis>
Measurements (n=7): WH: 0.760.85; LH: 0.550.67; LA (
<typeStatus id="5498881FFFE5B93BFBD25BDD81329FE6" box="[1028,1133,1215,1240]" pageId="12" pageNumber="181" type="holotype">holotype</typeStatus>
): 2.60; LE: 0.20¯0.25; LT: 0.20; LPM/WPM (
<typeStatus id="5498881FFFE5B93BFEBD5B81848B9FC2" box="[363,468,1251,1276]" pageId="12" pageNumber="181" type="holotype">holotype</typeStatus>
): III: 0.20 × 0.07, IV: 0.10 × 0.03; LP: 0.650.78; WPMax: 0.680.80; WPMin: 0.55 0.65; LEl: 1.461.62; WE: 1.451.75; WA: 1.45¯1.75; LMTb: 1.201.22; LMTr: 0.370.44; LAed: 0.600.62; TL: 4.205.15 (
<typeStatus id="5498881FFFE5B93BFEC05A4984219E7A" box="[278,382,1323,1348]" pageId="12" pageNumber="181" type="holotype">holotype</typeStatus>
: 4.35).
</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF115A2D867B9D5D" blockId="12.[151,1437,1179,2035]" pageId="12" pageNumber="181">
Head and abdomen brown, pronotum and elytra reddish brown; mouthparts, antennae and legs yellow brown; bases of antennomeres 1¯3 and ocelli yellow to yellow brown. Punctation of head regular, small and indistinct; punctation of pronotum denser and deeper than that on head, interspaces between punctures in median part of disc as diameter of 23 punctures, with small elongate impunctate area in medio-basal portion; punctation of elytra deeper and larger than that on pronotum, interspaces between punctures on median portion of each elytron as diameter of 12 punctures; abdomen with very dense, fine regular punctation. Microsculpture of pronotum finer than that on head with distinctly visible punctation on background of it; narrow band along basal margin with coarser and smaller ground sculpture. Habitus as in
<figureCitation id="13182A38FFE5B93BFD0A592986409D5D" box="[732,799,1611,1636]" captionStart="FIGURES 1 5" captionStartId="4.[151,264,1968,1991]" captionTargetBox="[167,1428,711,1924]" captionTargetId="figure@4.[150,1436,697,1947]" captionTargetPageId="4" captionText="FIGURES 1 5. Habitus of Philydrodes: 1 — Ph. tridentatus, holotype, 2 — Ph. michaeli, paratype, 3 — Ph. reticulatus, paratype, 4 — Ph. tibetanus, holotype, 5 — Ph. wrasei, paratype. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/291271/files/figure.png" pageId="12" pageNumber="181">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF11590D86479DEC" blockId="12.[151,1437,1179,2035]" pageId="12" pageNumber="181">
Head 1.2¯1.3 times as broad as long. Eyes as long as temples or slightly longer. Maxillary palp segment 3 twice as long as apical (4th) segment. Antennomeres with lengths widths (
<typeStatus id="5498881FFFE5B93BFC0C59F7811C9D90" box="[986,1091,1685,1710]" pageId="12" pageNumber="181" type="holotype">holotype</typeStatus>
): 1: 0.27 0.10; 2: 0.20 0.07; 3: 0.25 0.07; 48: 0.23 0.07; 910: 0.23 0.08; 11: 0.27 0.08.
</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF1159BD864B9C25" blockId="12.[151,1437,1179,2035]" pageId="12" pageNumber="181">Pronotum slightly narrower than head, slightly broader than long. Elytra slightly more than twice longer than pronotum, as long as broad or slightly broader than long.</paragraph>
<paragraph id="8B9C36BDFFE5B93BFF115847875E9CCC" blockId="12.[151,1437,1179,2035]" pageId="12" pageNumber="181">
Male. Apical margin of abdominal tergite VIII rounded (Fig. 44). Apical margin of abdominal sternite VIII with narrow deep emargination (Fig. 52). Aedeagus (
<figureCitation id="13182A38FFE5B93BFCD9582886399C5C" box="[783,870,1866,1891]" captionStart="FIGURES 36 41" captionStartId="9.[151,264,1968,1991]" captionTargetBox="[259,1317,200,1934]" captionTargetId="figure@9.[237,1334,193,1945]" captionTargetPageId="9" captionText="FIGURES 36 41. Aedeagus of Philydrodes michaeli (36, 39: holotype), Ph. reticulatus (37, 40: paratype) and Ph. wrasei (38, 41: paratype): 36 ‾ 38 — parameral view, 39 ‾ 41 — lateral view. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291797/files/figure.png" pageId="12" pageNumber="181">Fig. 37</figureCitation>
) elongated, with small phallobase, median lobe narrowing after median part toward rounded apex; parameres thin, exceeding apex of aedeagus, with two apical and two postero-apical short setae; aedeagus with strongly sclerotized narrow dorsal lobe not reaching apex of aedeagus, with very thin long flagellum within it; endophallus with two elongated areas of thorns in apical third. Aedeagus laterally as in
<figureCitation id="13182A38FFE5B93BFE7C58B884A19CCC" box="[426,510,2010,2035]" captionStart="FIGURES 36 41" captionStartId="9.[151,264,1968,1991]" captionTargetBox="[259,1317,200,1934]" captionTargetId="figure@9.[237,1334,193,1945]" captionTargetPageId="9" captionText="FIGURES 36 41. Aedeagus of Philydrodes michaeli (36, 39: holotype), Ph. reticulatus (37, 40: paratype) and Ph. wrasei (38, 41: paratype): 36 ‾ 38 — parameral view, 39 ‾ 41 — lateral view. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291797/files/figure.png" pageId="12" pageNumber="181">Fig. 40</figureCitation>
.
</paragraph>
<paragraph id="8B9C36BDFFE4B93AFF115FF584B49BEB" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="182">Female. Apical margin of abdominal tergite VIII indistinctly rounded (Fig. 45). Apical margin of abdominal sternite VIII straight (Fig. 53).</paragraph>
<paragraph id="8B9C36BDFFE4B93AFF115FBD84F99A92" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFF115FBD84EE9BC6" bold="true" box="[199,433,223,248]" pageId="13" pageNumber="182">Comparative notes.</emphasis>
Based on the shape of narrow aedeagus with very small phallobase,
<taxonomicName id="4C234D3EFFE4B93AFB6E5FBD80029BC6" box="[1208,1373,223,248]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="182" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<collectingCountry id="F334762DFFE4B93AFB6E5FBD81889BC6" box="[1208,1239,223,248]" name="Philippines" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFB6E5FBD81889BC6" box="[1208,1239,223,248]" italics="true" pageId="13" pageNumber="182">Ph</emphasis>
</collectingCountry>
.
<emphasis id="B957EAAFFFE4B93AFB305FBD80029BC6" box="[1254,1373,223,248]" italics="true" pageId="13" pageNumber="182">reticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457D4FFE4B93AFAB25FBD80C49BC6" box="[1380,1435,223,248]" pageId="13" pageNumber="182" rank="species">
<emphasis id="B957EAAFFFE4B93AFAB25FBD80C49BC6" bold="true" box="[1380,1435,223,248]" pageId="13" pageNumber="182">sp.n.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C234D3EFFE4B93AFEF75E6684F19A23" box="[289,430,260,285]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="182" phylum="Arthropoda" rank="species" species="michaeli" status="sp. nov.">
<collectingCountry id="F334762DFFE4B93AFEF75E66841F9A23" box="[289,320,260,285]" name="Philippines" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFEF75E66841F9A23" box="[289,320,260,285]" italics="true" pageId="13" pageNumber="182">Ph</emphasis>
</collectingCountry>
.
<emphasis id="B957EAAFFFE4B93AFE985E6684F19A23" box="[334,430,260,285]" italics="true" pageId="13" pageNumber="182">michaeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457D4FFE4B93AFE635E6684B39A23" box="[437,492,260,285]" pageId="13" pageNumber="182" rank="species">
<emphasis id="B957EAAFFFE4B93AFE635E6684B39A23" bold="true" box="[437,492,260,285]" pageId="13" pageNumber="182">sp.n.</emphasis>
</taxonomicNameLabel>
, from which it differs by slightly wider median lobe and parameres, by the narrower apex of aedeagus, by the arrangment of parameral setae (all parameral setae of
<taxonomicName id="4C234D3EFFE4B93AFBEA5E4581929A7E" box="[1084,1229,295,320]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="182" phylum="Arthropoda" rank="species" species="michaeli" status="sp. nov.">
<collectingCountry id="F334762DFFE4B93AFBEA5E4581049A7E" box="[1084,1115,295,320]" name="Philippines" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFBEA5E4581049A7E" box="[1084,1115,295,320]" italics="true" pageId="13" pageNumber="182">Ph</emphasis>
</collectingCountry>
.
<emphasis id="B957EAAFFFE4B93AFBBB5E4581929A7E" box="[1133,1229,295,320]" italics="true" pageId="13" pageNumber="182">michaeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457D4FFE4B93AFB0E5E45804F9A7E" box="[1240,1296,295,320]" pageId="13" pageNumber="182" rank="species">
<emphasis id="B957EAAFFFE4B93AFB0E5E45804F9A7E" bold="true" box="[1240,1296,295,320]" pageId="13" pageNumber="182">sp.n.</emphasis>
</taxonomicNameLabel>
are apical); furthermore it differs from other Chinese species by the more indistict punctation of head, by the longer eyes, by the narrower pronotum, by the slightly shorter elytra and by the shape of apical emargination of male abdominal tergite and sternite VIII.
</paragraph>
</subSubSection>
<subSubSection id="C3396536FFE4B93AFF115ED587A89ACA" pageId="13" pageNumber="182" type="etymology">
<paragraph id="8B9C36BDFFE4B93AFF115ED587A89ACA" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFF115ED584119AEE" bold="true" box="[199,334,439,464]" pageId="13" pageNumber="182">Etymology.</emphasis>
The specific epithet is a Latin adjective
<taxonomicName id="4C234D3EFFE4B93AFCC35ED586D39AEE" box="[789,908,439,464]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="182" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B957EAAFFFE4B93AFCC35ED586D39AEE" box="[789,908,439,464]" italics="true" pageId="13" pageNumber="182">reticulatus</emphasis>
</taxonomicName>
(reticulate, meshy) and alludes to the presence of moderately strong microsculpture on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C3396536FFE4B93AFF115E9D869299F2" pageId="13" pageNumber="182" type="distribution">
<paragraph id="8B9C36BDFFE4B93AFF115E9D84159903" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFF115E9D843F9926" bold="true" box="[199,352,511,536]" pageId="13" pageNumber="182">Distribution.</emphasis>
The species is known from the
<typeStatus id="5498881FFFE4B93AFD1A5D6287A39926" box="[716,764,512,536]" pageId="13" pageNumber="182">type</typeStatus>
locality (
<figureCitation id="13182A38FFE4B93AFCBE5E9D86E59926" box="[872,954,511,536]" captionStart="FIGURE 58" captionStartId="13.[151,250,1553,1575]" captionTargetBox="[179,1408,764,1532]" captionTargetId="figure@13.[179,1408,764,1532]" captionTargetPageId="13" captionText="FIGURE 58. Distribution of Philydrodes in China: Ph. tridentatus (black circles), Ph. michaeli (black star), Ph. reticulatus (black square), Ph. tibetanus (black rhomb), Ph. wrasei (black triangle)." httpUri="https://zenodo.org/record/291277/files/figure.png" pageId="13" pageNumber="182">Fig. 58</figureCitation>
) in high altitudes of Qin Ling Shan range,
<collectingRegion id="49E7F85FFFE4B93AFF415D4685AB9903" box="[151,244,548,573]" country="China" name="Shaanxi" pageId="13" pageNumber="182">Shaanxi</collectingRegion>
,
<collectingCountry id="F334762DFFE4B93AFF295D4684199903" box="[255,326,548,573]" name="China" pageId="13" pageNumber="182">China</collectingCountry>
.
</paragraph>
<paragraph id="8B9C36BDFFE4B93AFF115D25869299F2" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="182">
<emphasis id="B957EAAFFFE4B93AFF115D258415995E" bold="true" box="[199,330,583,608]" pageId="13" pageNumber="182">Bionomics.</emphasis>
According to the personal communication of M. Schülke, the specimens were collected in a place (
<quantity id="4CDB9B58FFE4B93AFF485D0E846499BA" box="[158,315,620,645]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="13" pageNumber="182" unit="m" value="2400.0" valueMax="2500.0" valueMin="2300.0">23002500 m</quantity>
a.s.l.) with streams and large rocks overgrown with grass, bushes and small trees (mostly pines); the specimens were collected running on the ground, only at the surface of places forming small swampy areas with small patches of
<taxonomicName id="4C234D3EFFE4B93AFE825DD684CB99F3" box="[340,404,692,717]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="182" phylum="Tracheophyta" rank="genus">
<emphasis id="B957EAAFFFE4B93AFE825DD684CB99F3" box="[340,404,692,717]" italics="true" pageId="13" pageNumber="182">Alnus</emphasis>
</taxonomicName>
forest by sifting and near the water turning stones.
</paragraph>
</subSubSection>
</treatment>
</document>