treatments-xml/data/03/8A/87/038A8789FFACFFD516FE5DCCC054FCB6.xml

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<mods:title id="CC821964387E36DC575346E2337B01C8">Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)</mods:title>
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<mods:namePart id="A733C99224AA7758B23AF1A4D52698CA">Xu, Xin</mods:namePart>
<mods:affiliation id="DAB1015B00C9143F7FA21227CD673B84">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China &amp; School of Life and Environmental Sciences, University of Sydney, Sydney, NSW, Australia &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering &amp; Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<mods:affiliation id="3B03A6059F2DE440C789EF87B8320C81">State Key Laboratory of Biocatalysis and Enzyme Engineering &amp; Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<subSubSection id="C3396514FFACFFCA16FE5DCCC69AFE65" box="[151,765,427,454]" pageId="14" pageNumber="145" type="nomenclature">
<paragraph id="8B9C369FFFACFFCA16FE5DCCC69AFE65" blockId="14.[151,765,427,490]" box="[151,765,427,454]" pageId="14" pageNumber="145">
<heading id="D0D481F3FFACFFCA16FE5DCCC69AFE65" bold="true" box="[151,765,427,454]" fontSize="11" level="1" pageId="14" pageNumber="145" reason="1">
<emphasis id="B957EA8DFFACFFCA16FE5DCCC69AFE65" bold="true" box="[151,765,427,454]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5DCCC60EFE65" authority="(Schenkel, 1953)" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[151,617,427,454]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis" status="comb. nov.">
<emphasis id="B957EA8DFFACFFCA16FE5DCCC5F4FE65" bold="true" box="[151,403,427,454]" italics="true" pageId="14" pageNumber="145">Luthela schensiensis</emphasis>
(
<bibRefCitation id="EFB24B6EFFACFFCA17CB5DCBC607FE65" author="Schenkel, E." box="[418,608,428,454]" pageId="14" pageNumber="145" pagination="1 - 108" refId="ref14178" refString="Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108." type="journal article" year="1953">Schenkel, 1953</bibRefCitation>
)
</taxonomicName>
, 
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</emphasis>
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<subSubSection id="C3396514FFACFFCA16FE5DB7C5E9FE49" box="[151,398,464,490]" pageId="14" pageNumber="145" type="description">
<paragraph id="8B9C369FFFACFFCA16FE5DB7C5E9FE49" blockId="14.[151,765,427,490]" box="[151,398,464,490]" pageId="14" pageNumber="145">
(
<figureCitation id="13182A1AFFACFFCA16F65DB7C54BFE49" box="[159,300,464,490]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Figs 10A–K</figureCitation>
, 
<figureCitation id="13182A1AFFACFFCA17515DB7C5E1FE49" box="[312,390,464,490]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="14" pageNumber="145">11A–P</figureCitation>
)
</paragraph>
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<subSubSection id="C3396514FFACFFCA16FE5E73C654FC18" pageId="14" pageNumber="145" type="reference_group">
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<treatmentCitationGroup id="AB3311B1FFACFFCA16FE5E73C1FAFD8F" box="[151,1437,532,557]" pageId="14" pageNumber="145">
<treatmentCitation id="0A82108EFFACFFCA16FE5E73C6E6FD8E" author="Schenkel, E." box="[151,641,532,557]" page="1" pageId="14" pageNumber="145" year="1953">
<taxonomicName id="4C234D1CFFACFFCA16FE5E73C6E6FD8E" authority="Schenkel, 1953: 1" authorityName="Schenkel" authorityPageNumber="1" authorityYear="1953" box="[151,641,532,557]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="subSpecies" species="sinensis" subSpecies="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5E73C5A9FD8F" box="[151,462,532,556]" italics="true" pageId="14" pageNumber="145">Liphistius sinensis schensiensis</emphasis>
<bibRefCitation id="EFB24B6EFFACFFCA17B85E73C6E6FD8E" author="Schenkel, E." box="[465,641,532,557]" pageId="14" pageNumber="145" pagination="1 - 108" refId="ref14178" refString="Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108." type="journal article" year="1953">Schenkel,1953: 1</bibRefCitation>
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(the subspecies was described based on 
<specimenCount id="9D25FD16FFACFFCA126A5E72C008FD8F" box="[1027,1135,532,556]" pageId="14" pageNumber="145" type="female">one female</specimenCount>
collected from Tongyuanfang,
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Gaoling, 
<collectingRegion id="49E7F87DFFACFFCA174C5E54C5BDFDE8" box="[293,474,563,587]" country="China" name="Shaanxi" pageId="14" pageNumber="145">Shaanxi Province</collectingRegion>
, 
<date id="FF9D105FFFACFFCA178A5E54C6E5FDE8" box="[483,642,563,587]" pageId="14" pageNumber="145" value="1916-10-08">
<collectingDate id="EFD9E9B7FFACFFCA178A5E54C6E5FDE8" box="[483,642,563,587]" pageId="14" pageNumber="145" value="1916-10-08">8 October 1916</collectingDate>
</date>
, deposited at Museum Hoangho-Peiho in 
<collectingCountry id="F334760FFFACFFCA124E5E54C01EFDE8" box="[1063,1145,563,587]" name="China" pageId="14" pageNumber="145">Tientsin</collectingCountry>
[corresponding to Huanghebaihe in 
<collectingCountry id="F334760FFFACFFCA17775E35C501FDC9" box="[286,358,594,618]" name="China" pageId="14" pageNumber="145">Tianjin</collectingCountry>
and Yellow River-Haihe River in 
<collectingCountry id="F334760FFFACFFCA14A95E35C76CFDC9" box="[704,779,594,618]" name="China" pageId="14" pageNumber="145">Tianjin</collectingCountry>
, in Chinese and English, respectively], lost).
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5E17C633FD2B" blockId="14.[151,1437,532,955]" box="[151,596,624,648]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5E17C637FD2B" authority=": Gertsch, 1967: 115" authorityName="Gertsch" authorityPageNumber="115" authorityYear="1967" box="[151,592,624,648]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5E17C519FD2B" box="[151,382,624,648]" italics="true" pageId="14" pageNumber="145">Liphistius schensiensis</emphasis>
: Gertsch, 1967: 115
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5EE8C6CFFD04" blockId="14.[151,1437,532,955]" box="[151,680,655,679]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5EE8C6C3FD04" authority=": Haupt, 1983: 285" authorityName="Haupt" authorityPageNumber="285" authorityYear="1983" box="[151,676,655,679]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="subSpecies" species="sinensis" subSpecies="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5EE8C58CFD04" box="[151,491,655,679]" italics="true" pageId="14" pageNumber="145">Heptathela sinensis schensiensis:</emphasis>
<bibRefCitation id="EFB24B6EFFACFFCA17985EF7C6C3FD04" author="Haupt, J." box="[497,676,655,679]" pageId="14" pageNumber="145" pagination="275 - 293" refId="ref14030" refString="Haupt, J. (1983) Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur Zoologische Systematik und Evolutionsforschung, 21 (4), 275 - 293. https: // doi. org / 10.1111 / j. 1439 - 0469.1983. tb 00296. x" type="journal article" year="1983">Haupt, 1983: 285</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5EC9C71DFD65" blockId="14.[151,1437,532,955]" box="[151,890,686,710]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5EC9C614FD65" authority=": Zhu &amp; Wang, 1983: 131" authorityName="Zhu &amp; Wang" authorityPageNumber="131" authorityYear="1983" box="[151,627,686,710]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="xianensis">
<emphasis id="B957EA8DFFACFFCA16FE5EC9C50BFD66" box="[151,364,686,710]" italics="true" pageId="14" pageNumber="145">Heptathela xianensis</emphasis>
: Zhu &amp; Wang, 1983: 131
</taxonomicName>
; Wang &amp; 
<bibRefCitation id="EFB24B6EFFACFFCA14BE5EC9C711FD65" author="Zhu, C. D. &amp; Wang, Y. W." box="[727,886,686,710]" pageId="14" pageNumber="145" pagination="251 - 253" refId="ref15094" refString="Zhu, C. D. &amp; Wang, Y. W. (1984) A new species of genus Liphistius (Araneae: Liphistiidae). Journal of the Bethune Medical University, 10, 251 - 253." type="journal article" year="1984">Zhu, 1984: 403</bibRefCitation>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5EABC171FD46" blockId="14.[151,1437,532,955]" box="[151,1302,716,741]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5EABC6ABFD46" authority=": Platnick &amp; Sedgwick, 1984: 4" authorityName="Platnick &amp; Sedgwick" authorityPageNumber="4" authorityYear="1984" box="[151,716,716,741]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5EABC5ECFD47" box="[151,395,716,740]" italics="true" pageId="14" pageNumber="145">Heptathela schensiensis</emphasis>
: Platnick &amp; Sedgwick, 1984: 4
</taxonomicName>
; 
<bibRefCitation id="EFB24B6EFFACFFCA14BE5EAAC7B8FD46" author="Song, D. X. &amp; Haupt, J." box="[727,991,717,741]" pageId="14" pageNumber="145" pagination="443 - 451" refId="ref14273" refString="Song, D. X. &amp; Haupt, J. (1984) Comparative morphology and phylogeny of Liphistiomorph spiders (Araneae: Mesothelae). 2. Revision of new Chinese Heptathelid species. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 27, 443 - 451." type="journal article" year="1984">Song &amp; Haupt, 1984: 447</bibRefCitation>
; Song, Zhu &amp; Chen, 1999: 33.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5E8CC159FC82" blockId="14.[151,1437,532,955]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5E8CC61FFCA0" authority="Zhu &amp; Wang, 1984: 251" authorityName="Zhu &amp; Wang" authorityPageNumber="251" authorityYear="1984" box="[151,632,747,772]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFACFFCA16FE5E8CC51AFCA0" box="[151,381,747,771]" italics="true" pageId="14" pageNumber="145">Liphistius heyangensis</emphasis>
<bibRefCitation id="EFB24B6EFFACFFCA17EA5E8CC61FFCA0" author="Zhu, C. D. &amp; Wang, Y. W." box="[387,632,747,772]" pageId="14" pageNumber="145" pagination="251 - 253" refId="ref15094" refString="Zhu, C. D. &amp; Wang, Y. W. (1984) A new species of genus Liphistius (Araneae: Liphistiidae). Journal of the Bethune Medical University, 10, 251 - 253." type="journal article" year="1984">Zhu &amp; Wang, 1984: 251</bibRefCitation>
</taxonomicName>
(holotype: female, from Heyang, 
<collectingRegion id="49E7F87DFFACFFCA15B85E8CC0E1FCA0" box="[977,1158,747,771]" country="China" name="Shaanxi" pageId="14" pageNumber="145">Shaanxi Province</collectingRegion>
, 
<date id="FF9D105FFFACFFCA12E65E8CC17FFCA0" box="[1167,1304,747,771]" pageId="14" pageNumber="145" value="1982-06-30">30 June 1982</date>
, deposited at Bethune Medical University, now College of Life Sciences, 
<collectingRegion id="49E7F87DFFACFFCA15435F6DC731FC81" box="[810,854,778,802]" country="China" name="Jilin" pageId="14" pageNumber="145">Jilin</collectingRegion>
University, type presumed to be lost). Syn. nov.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5F4FC653FCE3" blockId="14.[151,1437,532,955]" box="[151,564,808,833]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5F4FC657FCE3" authority=": Ono, 2000: 150" authorityName="Ono" authorityPageNumber="150" authorityYear="2000" box="[151,560,808,833]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5F4FC5E6FCE3" box="[151,385,808,832]" italics="true" pageId="14" pageNumber="145">Abcathela schensiensis</emphasis>
: 
<bibRefCitation id="EFB24B6EFFACFFCA17E65F4EC657FCE3" author="Ono, H." box="[399,560,808,833]" pageId="14" pageNumber="145" pagination="145 - 151" refId="ref14140" refString="Ono, H. (2000) Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae). Memoirs of the National Science Museum, 33, 145 - 151." type="journal article" year="2000">Ono, 2000: 150</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5F20C656FCFC" blockId="14.[151,1437,532,955]" box="[151,561,839,863]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5F20C51CFCFC" authority="(Schenkel, 1953)" box="[151,379,839,863]" class="Arachnida" family="Liphistiidae" genus="Sinothela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="heyangensi">
<emphasis id="B957EA8DFFACFFCA16FE5F20C508FCFC" box="[151,367,839,863]" italics="true" pageId="14" pageNumber="145">Sinothela heyangensi</emphasis>
s
</taxonomicName>
: 
<bibRefCitation id="EFB24B6EFFACFFCA17EF5F2FC64AFCFC" author="Haupt, J." box="[390,557,839,863]" pageId="14" pageNumber="145" pagination="1 - 102" refId="ref14095" refString="Haupt, J. (2003) The Mesothelae - a monograph of an exceptional group of spiders (Araneae: Mesothelae) (Morphology, behaviour, ecology, taxonomy, distribution and phylogeny). Zoologica, 154, 1 - 102." type="journal article" year="2003">Haupt, 2003: 71</bibRefCitation>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5F01C772FCDD" blockId="14.[151,1437,532,955]" box="[151,789,870,894]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5F01C649FCDD" authority=": Haupt, 2003: 71" authorityName=": Haupt" authorityPageNumber="71" authorityYear="2003" box="[151,558,870,894]" class="Arachnida" family="Liphistiidae" genus="Sinothela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA16FE5F01C51DFCDD" box="[151,378,870,894]" italics="true" pageId="14" pageNumber="145">Sinothela schensiensis</emphasis>
: 
<bibRefCitation id="EFB24B6EFFACFFCA17E15F01C649FCDD" author="Haupt, J." box="[392,558,870,894]" pageId="14" pageNumber="145" pagination="1 - 102" refId="ref14095" refString="Haupt, J. (2003) The Mesothelae - a monograph of an exceptional group of spiders (Araneae: Mesothelae) (Morphology, behaviour, ecology, taxonomy, distribution and phylogeny). Zoologica, 154, 1 - 102." type="journal article" year="2003">Haupt, 2003: 71</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="EFB24B6EFFACFFCA14505F01C776FCDD" author="Xu, X. &amp; Liu, F. &amp; Chen, J. &amp; Ono, H. &amp; Li, D. &amp; Kuntner, M." box="[569,785,870,894]" pageId="14" pageNumber="145" pagination="121 - 151" refId="ref14523" refString="Xu, X., Liu, F., Chen, J., Ono, H., Li, D. &amp; Kuntner, M. (2015 a) A genus-level taxonomic review of primitively segmented spiders (Mesothelae, Liphistiidae). Zookeys, 488, 121 - 151. https: // doi. org / 10.3897 / zookeys. 488.8726" type="journal article" year="2015">Xu et al., 2015a: 139</bibRefCitation>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5FE3C630FC3F" blockId="14.[151,1437,532,955]" box="[151,599,900,925]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5FE3C634FC3F" authority=": Platnick, 1989: 57" authorityName="Platnick" authorityPageNumber="57" authorityYear="1989" box="[151,595,900,925]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFACFFCA16FE5FE3C5EDFC3F" box="[151,394,900,924]" italics="true" pageId="14" pageNumber="145">Heptathela heyangensis</emphasis>
: Platnick, 1989: 57
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C369FFFACFFCA16FE5FC4C654FC18" blockId="14.[151,1437,532,955]" box="[151,563,931,955]" pageId="14" pageNumber="145">
<taxonomicName id="4C234D1CFFACFFCA16FE5FC4C648FC18" authority=": Ono, 2000: 150" authorityName=": Ono" authorityPageNumber="150" authorityYear="2000" box="[151,559,931,955]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFACFFCA16FE5FC4C5E7FC18" box="[151,384,931,955]" italics="true" pageId="14" pageNumber="145">Abcathela heyangensis</emphasis>
: 
<bibRefCitation id="EFB24B6EFFACFFCA17E45FC4C648FC18" author="Ono, H." box="[397,559,931,955]" pageId="14" pageNumber="145" pagination="145 - 151" refId="ref14140" refString="Ono, H. (2000) Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae). Memoirs of the National Science Museum, 33, 145 - 151." type="journal article" year="2000">Ono, 2000: 150</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFACFFCA16FE5F8EC0E8FB5C" pageId="14" pageNumber="145" type="materials_examined">
<paragraph id="8B9C369FFFACFFCA16FE5F8EC0E8FB5C" blockId="14.[151,1437,1001,2036]" pageId="14" pageNumber="145">
<emphasis id="B957EA8DFFACFFCA16FE5F8EC5E6FBA0" bold="true" box="[151,385,1001,1027]" pageId="14" pageNumber="145">Material examined.</emphasis>
<materialsCitation id="3B4B3CC2FFACFFCA17E05F8EC6ACFBE8" ID-GBIF-Occurrence="3435942327" pageId="14" pageNumber="145">
<specimenCount id="9D25FD16FFACFFCA17E05F8EC5B8FBA0" box="[393,479,1001,1027]" pageId="14" pageNumber="145" type="male">3 males</specimenCount>
(XUX-2011-241/242/247, matured on 
<date id="FF9D105FFFACFFCA15F35F8EC04BFBA0" box="[922,1068,1001,1027]" pageId="14" pageNumber="145" value="2012-07-10">10 July 2012</date>
at CBEE) and 
<specimenCount id="9D25FD16FFACFFCA12B15F8DC122FBA0" box="[1240,1349,1001,1028]" pageId="14" pageNumber="145" type="female">7 females</specimenCount>
(XUX- 2011-243/245/248/249/251/253/254), 
<location id="8EFC6044FFACFFCA1420586AC77AFB84" LSID="urn:lsid:plazi:treatment:038A8789FFACFFD516FE5DCCC054FCB6:8EFC6044FFACFFCA1420586AC77AFB84" box="[585,797,1037,1063]" country="China" county="Heyang County" latitude="35.242" longLatPrecision="71" longitude="110.155" name="Kangjiapo Village" pageId="14" pageNumber="145" stateProvince="Shaanxi">Kangjiapo Village</location>
, 
<collectingCounty id="62FD4E13FFACFFCA15475869C782FB84" box="[814,997,1037,1064]" pageId="14" pageNumber="145">Heyang County</collectingCounty>
, 
<collectingRegion id="49E7F87DFFACFFCA159C586AC0A5FB84" box="[1013,1218,1037,1063]" country="China" name="Shaanxi" pageId="14" pageNumber="145">Shaanxi Province</collectingRegion>
, 
<collectingCountry id="F334760FFFACFFCA12BA586AC17EFB84" box="[1235,1305,1037,1063]" name="China" pageId="14" pageNumber="145">China</collectingCountry>
, 
<geoCoordinate id="EE175058FFACFFCA1340586AC1FFFB8B" box="[1321,1432,1037,1064]" degrees="35.242" direction="north" orientation="latitude" pageId="14" pageNumber="145" precision="55" value="35.242">35.242°N</geoCoordinate>
, 
<geoCoordinate id="EE175058FFACFFCA16FE5856C561FBEF" box="[151,262,1073,1100]" degrees="110.155" direction="east" orientation="longitude" pageId="14" pageNumber="145" precision="55" value="110.155">110.155E</geoCoordinate>
, 
<quantity id="4CDB9B7AFFACFFCA17785856C53BFBEF" box="[273,348,1073,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="145" unit="m" value="730.0">
<elevation id="000ED1ACFFACFFCA17785856C53BFBEF" box="[273,348,1073,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="145" unit="m" value="730.0">730 m</elevation>
</quantity>
, collected on 1 
<collectorName id="26D65349FFACFFCA14795856C6E2FBE8" box="[528,645,1073,1099]" pageId="14" pageNumber="145">December</collectorName>
, 2011
</materialsCitation>
; 
<materialsCitation id="3B4B3CC2FFACFFCA14B15856C1F0FB30" ID-GBIF-Occurrence="3435942307" pageId="14" pageNumber="145">
<specimenCount id="9D25FD16FFACFFCA14B15856C757FBE8" box="[728,816,1073,1099]" pageId="14" pageNumber="145" type="male">3 males</specimenCount>
(XUX-2011-255/264 matured on 
<date id="FF9D105FFFACFFCA12DD5856C12DFBE8" box="[1204,1354,1073,1099]" pageId="14" pageNumber="145" value="2012-07-10">10 July 2012</date>
, XUX- 2011-263 matured on 
<date id="FF9D105FFFACFFCA17E25832C646FBCC" box="[395,545,1109,1136]" pageId="14" pageNumber="145" value="2013-06-13">13 June 2013</date>
, at CBEE) and 
<specimenCount id="9D25FD16FFACFFCA14A35832C724FBCC" box="[714,835,1109,1135]" pageId="14" pageNumber="145" type="female">10 females</specimenCount>
(XUX-2011-256–262/265/266/269), 
<location id="8EFC6044FFACFFCA128F5832C1FFFBCC" LSID="urn:lsid:plazi:treatment:038A8789FFACFFD516FE5DCCC054FCB6:8EFC6044FFACFFCA128F5832C1FFFBCC" box="[1254,1432,1109,1135]" country="China" county="Gaoling County" latitude="35.546" longLatPrecision="71" longitude="109.04" municipality="Tongyuan" name="Quyang Village" pageId="14" pageNumber="145" stateProvince="Shaanxi">Quyang Village</location>
, 
<collectingMunicipality id="6BF8ACE5FFACFFCA16FE581DC56DFB37" box="[151,266,1146,1172]" pageId="14" pageNumber="145">Tongyuan</collectingMunicipality>
, 
<collectingCounty id="62FD4E13FFACFFCA177D581EC5AEFB30" box="[276,457,1145,1171]" pageId="14" pageNumber="145">Gaoling County</collectingCounty>
, 
<collectingRegion id="49E7F87DFFACFFCA17BA581EC6FDFB30" box="[467,666,1145,1171]" country="China" name="Shaanxi" pageId="14" pageNumber="145">Shaanxi Province</collectingRegion>
, 
<collectingCountry id="F334760FFFACFFCA14CD581EC68DFB30" box="[676,746,1145,1171]" name="China" pageId="14" pageNumber="145">China</collectingCountry>
, 
<geoCoordinate id="EE175058FFACFFCA149C581EC703FB37" box="[757,868,1145,1172]" degrees="35.546" direction="north" orientation="latitude" pageId="14" pageNumber="145" precision="55" value="35.546">35.546°N</geoCoordinate>
, 
<geoCoordinate id="EE175058FFACFFCA1506581EC7B9FB30" box="[879,990,1145,1171]" degrees="109.040" direction="east" orientation="longitude" pageId="14" pageNumber="145" precision="55" value="109.04">109.040E</geoCoordinate>
, 
<quantity id="4CDB9B7AFFACFFCA1581581EC056FB37" box="[1000,1073,1145,1172]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="145" unit="m" value="390.0">
<elevation id="000ED1ACFFACFFCA1581581EC056FB37" box="[1000,1073,1145,1172]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="145" unit="m" value="390.0">390 m</elevation>
</quantity>
, collected on 2 
<collectorName id="26D65349FFACFFCA12B7581EC134FB30" box="[1246,1363,1145,1171]" pageId="14" pageNumber="145">December</collectorName>
, 2011
</materialsCitation>
; 
<materialsCitation id="3B4B3CC2FFACFFCA16FE58FAC0B0FB14" ID-GBIF-Occurrence="3435942310" box="[151,1239,1181,1208]" pageId="14" pageNumber="145">
<specimenCount id="9D25FD16FFACFFCA16FE58FAC483FB14" box="[151,228,1181,1207]" pageId="14" pageNumber="145" type="male">1 male</specimenCount>
(XUX-2011-273, matured on 
<date id="FF9D105FFFACFFCA142858FAC6B1FB14" box="[577,726,1181,1207]" pageId="14" pageNumber="145" value="2012-07-10">10 July 2012</date>
at CBEE), 
<geoCoordinate id="EE175058FFACFFCA153458FAC7ABFB14" box="[861,972,1181,1207]" degrees="34.493" direction="north" orientation="latitude" pageId="14" pageNumber="145" precision="55" value="34.493">34.493°N</geoCoordinate>
, 
<geoCoordinate id="EE175058FFACFFCA15B358FAC02EFB14" box="[986,1097,1181,1207]" degrees="108.806" direction="east" orientation="longitude" pageId="14" pageNumber="145" precision="55" value="108.806">108.806E</geoCoordinate>
, 
<quantity id="4CDB9B7AFFACFFCA123F58FAC0C6FB1B" box="[1110,1185,1181,1208]" metricMagnitude="2" metricUnit="m" metricValue="3.7399999999999998" pageId="14" pageNumber="145" unit="m" value="374.0">374 m</quantity>
, and
</materialsCitation>
<materialsCitation id="3B4B3CC2FFACFFCA128858FAC0ECFB5C" ID-GBIF-Occurrence="3435942326" pageId="14" pageNumber="145">
<specimenCount id="9D25FD16FFACFFCA128858FAC123FB14" box="[1249,1348,1181,1207]" pageId="14" pageNumber="145" type="female">1 female</specimenCount>
(XUX- 2011-270), 
<geoCoordinate id="EE175058FFACFFCA177E58A6C5E1FB78" box="[279,390,1217,1243]" degrees="34.498" direction="north" orientation="latitude" pageId="14" pageNumber="145" precision="55" value="34.498">34.498°N</geoCoordinate>
, 
<geoCoordinate id="EE175058FFACFFCA17F958A6C667FB78" box="[400,512,1217,1243]" degrees="108.786" direction="east" orientation="longitude" pageId="14" pageNumber="145" precision="55" value="108.786">108.786E</geoCoordinate>
, 
<quantity id="4CDB9B7AFFACFFCA146058A6C637FB7F" box="[521,592,1217,1244]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="14" pageNumber="145" unit="m" value="359.0">
<elevation id="000ED1ACFFACFFCA146058A6C637FB7F" box="[521,592,1217,1244]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="14" pageNumber="145" unit="m" value="359.0">359 m</elevation>
</quantity>
, 
<location id="8EFC6044FFACFFCA143358A6C763FB78" LSID="urn:lsid:plazi:treatment:038A8789FFACFFD516FE5DCCC054FCB6:8EFC6044FFACFFCA143358A6C763FB78" box="[602,772,1217,1243]" country="China" county="Jingyang County" latitude="34.498" longLatPrecision="71" longitude="108.786" municipality="Xianyang City" name="Taiping village" pageId="14" pageNumber="145" stateProvince="Shaanxi">Taiping village</location>
, 
<collectingCounty id="62FD4E13FFACFFCA156758A6C7ABFB78" box="[782,972,1217,1243]" pageId="14" pageNumber="145">Jingyang County</collectingCounty>
, 
<collectingMunicipality id="6BF8ACE5FFACFFCA15BC58A6C01EFB78" box="[981,1145,1217,1243]" pageId="14" pageNumber="145">Xianyang City</collectingMunicipality>
, 
<collectingRegion id="49E7F87DFFACFFCA12EA58A6C12FFB78" box="[1155,1352,1217,1243]" country="China" name="Shaanxi" pageId="14" pageNumber="145">Shaanxi Province</collectingRegion>
, 
<collectingCountry id="F334760FFFACFFCA133B58A6C1FFFB78" box="[1362,1432,1217,1243]" name="China" pageId="14" pageNumber="145">China</collectingCountry>
, collected on 3 
<collectorName id="26D65349FFACFFCA17525882C5D7FB5C" box="[315,432,1253,1279]" pageId="14" pageNumber="145">December</collectorName>
, 2011, by 
<collectorName id="26D65349FFACFFCA144A5881C61FFB5C" box="[547,632,1253,1280]" pageId="14" pageNumber="145">H.Z. Li</collectorName>
, 
<collectorName id="26D65349FFACFFCA14ED5881C682FB5C" box="[644,741,1253,1280]" pageId="14" pageNumber="145">F.X. Liu</collectorName>
, 
<collectorName id="26D65349FFACFFCA14995881C70AFAA3" box="[752,877,1254,1280]" pageId="14" pageNumber="145">X.Y. Wang</collectorName>
, and X. 
<collectorName id="26D65349FFACFFCA15A05881C07BFB5C" box="[969,1052,1253,1280]" pageId="14" pageNumber="145">Xu. All</collectorName>
in CBEE
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFACFFCA16AE596EC669F977" pageId="14" pageNumber="145" type="diagnosis">
<paragraph id="8B9C369FFFACFFCA16AE596EC669F977" blockId="14.[151,1437,1001,2036]" pageId="14" pageNumber="145">
<emphasis id="B957EA8DFFACFFCA16AE596EC525FA80" bold="true" box="[199,322,1289,1315]" pageId="14" pageNumber="145">Diagnosis.</emphasis>
Males of 
<taxonomicName id="4C234D1CFFACFFCA17DA596DC63CFA80" authorityName=": Haupt" authorityYear="2003" box="[435,603,1289,1315]" class="Arachnida" family="Liphistiidae" genus="Sinothela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA17DA596DC63CFA80" box="[435,603,1289,1315]" italics="true" pageId="14" pageNumber="145">L. schensiensis</emphasis>
</taxonomicName>
resemble those of 
<taxonomicName id="4C234D1CFFACFFCA1559596DC7FEFA80" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[816,921,1290,1315]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA1559596DC7FEFA80" box="[816,921,1290,1315]" italics="true" pageId="14" pageNumber="145">L. yiyuan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA15F6596EC79EFA80" bold="true" box="[927,1017,1289,1315]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA15F6596EC79EFA80" box="[927,1017,1289,1315]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C234D1CFFACFFCA1259596DC0DEFA80" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1072,1209,1289,1315]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA1259596DC0DEFA80" box="[1072,1209,1289,1315]" italics="true" pageId="14" pageNumber="145">L. yuncheng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA12A8596EC17DFA80" bold="true" box="[1217,1306,1289,1315]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA12A8596EC17DFA80" box="[1217,1306,1289,1315]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished from those of 
<taxonomicName id="4C234D1CFFACFFCA17B05949C624FAE4" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[473,579,1326,1351]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA17B05949C624FAE4" box="[473,579,1326,1351]" italics="true" pageId="14" pageNumber="145">L. yiyuan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA1425594AC6C0FAE4" bold="true" box="[588,679,1325,1351]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA1425594AC6C0FAE4" box="[588,679,1325,1351]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the tegulum having a slightly smaller terminal apophysis (
<figureCitation id="13182A1AFFACFFCA1306594AC564FAC8" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10F, 10G</figureCitation>
), and the embolus with more longitudinal ribs retrolaterally (
<figureCitation id="13182A1AFFACFFCA15DC5936C03AFAC8" box="[949,1117,1361,1387]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10G, 10K</figureCitation>
); from those of 
<taxonomicName id="4C234D1CFFACFFCA137A5935C1FBFAC8" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1299,1436,1361,1387]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA137A5935C1FBFAC8" box="[1299,1436,1361,1387]" italics="true" pageId="14" pageNumber="145">L. yuncheng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA16FE5912C497FA2C" bold="true" box="[151,240,1397,1423]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA16FE5912C497FA2C" box="[151,240,1397,1423]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the marginal apophysis and the dorsal extension of the terminal apophysis of the tegulum with irregular serrations (
<figureCitation id="13182A1AFFACFFCA177C59FEC5A7FA10" box="[277,448,1433,1459]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10G, 10K</figureCitation>
); from those of 
<taxonomicName id="4C234D1CFFACFFCA141259FDC765FA10" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[635,770,1433,1459]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA141259FDC765FA10" box="[635,770,1433,1459]" italics="true" pageId="14" pageNumber="145">L. dengfeng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA156359FEC701FA10" bold="true" box="[778,870,1433,1459]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA156359FEC701FA10" box="[778,870,1433,1459]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the conductor with a longer lower spine (
<figureCitation id="13182A1AFFACFFCA130659FEC56FFA74" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10G, 10K</figureCitation>
); from those of 
<taxonomicName id="4C234D1CFFACFFCA17DE59D9C64FFA74" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[439,552,1469,1495]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA17DE59D9C64FFA74" box="[439,552,1469,1495]" italics="true" pageId="14" pageNumber="145">L. handan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA144759DAC6E1FA74" bold="true" box="[558,646,1469,1495]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA144759DAC6E1FA74" box="[558,646,1469,1495]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the contrategulum margin with slightly smaller teeth, and the dorsal extension of the terminal apophysis of the tegulum with irregular serrations (
<figureCitation id="13182A1AFFACFFCA15905986C0C6FA58" box="[1017,1185,1505,1531]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10G, 10K</figureCitation>
); and from those of 
<taxonomicName id="4C234D1CFFACFFCA13EF5985C48CF9BC" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA13EF5985C48CF9BC" italics="true" pageId="14" pageNumber="145">L. badong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457F6FFACFFCA169D5A61C54CF983" box="[244,299,1542,1568]" pageId="14" pageNumber="145" rank="species">sp. n</taxonomicNameLabel>
and 
<taxonomicName id="4C234D1CFFACFFCA170C5A61C5D9F9BC" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[357,446,1542,1567]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA170C5A61C5D9F9BC" box="[357,446,1542,1567]" italics="true" pageId="14" pageNumber="145">L. taian</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA17AF5A62C645F9BC" bold="true" box="[454,546,1541,1567]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA17AF5A62C645F9BC" box="[454,546,1541,1567]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the conductor with small middle spines (
<figureCitation id="13182A1AFFACFFCA12495A62C14DF9BC" box="[1056,1322,1541,1567]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="14" pageNumber="145">Fig. 10A–E, 10H, 10K</figureCitation>
). Females of 
<taxonomicName id="4C234D1CFFACFFCA16DF5A4DC538F9E0" authorityName=": Haupt" authorityYear="2003" box="[182,351,1577,1603]" class="Arachnida" family="Liphistiidae" genus="Sinothela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFACFFCA16DF5A4DC538F9E0" box="[182,351,1577,1603]" italics="true" pageId="14" pageNumber="145">L. schensiensis</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="4C234D1CFFACFFCA14245A4DC6A7F9E0" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[589,704,1577,1603]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA14245A4DC6A7F9E0" box="[589,704,1577,1603]" italics="true" pageId="14" pageNumber="145">L. badong</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA14A15A4EC745F9E0" bold="true" box="[712,802,1577,1603]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA14A15A4EC745F9E0" box="[712,802,1577,1603]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the middle pair of receptacular clusters with slightly thicker and shorter stalks; from those of 
<taxonomicName id="4C234D1CFFACFFCA14375A29C6A0F9C4" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[606,711,1614,1639]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA14375A29C6A0F9C4" box="[606,711,1614,1639]" italics="true" pageId="14" pageNumber="145">L. yiyuan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA14A75A2AC74FF9C4" bold="true" box="[718,808,1613,1639]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA14A75A2AC74FF9C4" box="[718,808,1613,1639]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the middle pair of receptacular clusters with slightly longer stalks (
<figureCitation id="13182A1AFFACFFCA175A5A16C66CF928" box="[307,523,1649,1675]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="14" pageNumber="145">Fig. 11A–D, 11I–L</figureCitation>
); from those of 
<taxonomicName id="4C234D1CFFACFFCA14D05A15C74EF928" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[697,809,1649,1675]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
<emphasis id="B957EA8DFFACFFCA14D05A15C74EF928" box="[697,809,1649,1675]" italics="true" pageId="14" pageNumber="145">L. handan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFACFFCA15465A16C7E0F928" bold="true" box="[815,903,1649,1675]" pageId="14" pageNumber="145">
<taxonomicNameLabel id="A26457F6FFACFFCA15465A16C7E0F928" box="[815,903,1649,1675]" pageId="14" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by the middle pair of receptacular clusters with slightly thicker stalks (
<figureCitation id="13182A1AFFACFFCA17FC5AF2C678F90C" box="[405,543,1685,1711]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="14" pageNumber="145">Fig. 11A–D</figureCitation>
); from those of the other 
<taxonomicName id="4C234D1CFFACFFCA15505AF2C781F90C" box="[825,998,1685,1711]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B957EA8DFFACFFCA15505AF2C7E8F90C" box="[825,911,1685,1711]" italics="true" pageId="14" pageNumber="145">Luthela</emphasis>
species
</taxonomicName>
by the bases of the middle stalks being close to each other (
<figureCitation id="13182A1AFFACFFCA17115ADEC665F977" box="[376,514,1721,1748]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="14" pageNumber="145">Fig. 11A–D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFACFFD416AE5ABAC11CF939" lastPageId="16" lastPageNumber="147" pageId="14" pageNumber="145" type="description">
<paragraph id="8B9C369FFFACFFCA16AE5ABAC67AF873" blockId="14.[151,1437,1001,2036]" pageId="14" pageNumber="145">
<emphasis id="B957EA8DFFACFFCA16AE5ABAC512F954" bold="true" box="[199,373,1757,1783]" pageId="14" pageNumber="145">Redescription.</emphasis>
Male (XUX-2011-273). Carapace reddish brown; opisthosoma brown, with dark brown tergites; sternum narrow, much longer than wide; few long pointed hairs running over ocular mound in longitudinal row; chelicerae robust with promargin of groove with 11 denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, second to fifth larger than others and the others small; 8 spinnerets. Measurements: BL 14.95, CL 7.05, CW 6.60, OL 7.15, OW 4.90; ALE&gt; PLE&gt; PME&gt; AME; leg I 20.65 (6.05 + 2.75 + 4.15 + 5.05 + 2.65), leg II 20.85 (5.95 + 2.65 + 3.95 + 5.25 + 3.05), leg III 21.30 (5.65 + 2.55 + 3.75 + 6.40 + 2.95), leg IV 29.00 (7.20 + 2.95 + 5.35 + 8.95 + 4.55).
</paragraph>
<paragraph id="8B9C369FFFACFFCB16AE5BBEC0F6FEC6" blockId="14.[151,1437,1001,2036]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="146" pageId="14" pageNumber="145">
Palp: Prolateral paracymbium unpigmented and unsclerotized, numerous setae and spines at tip of paracymbium (
<figureCitation id="13182A1AFFADFFCB16C95CF0C54DFF12" box="[160,298,151,177]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146">Fig. 10A–C</figureCitation>
). Contrategulum with 7–10 teeth along margin, and distal tooth serrated (
<figureCitation id="13182A1AFFADFFCB12195CF0C1E8FF12" box="[1136,1423,151,177]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146">Fig. 10A, 10D, 10H, 10J</figureCitation>
). Tegulum with an irregular dentate marginal apophysis and an irregular dorsal extension of terminal apophysis, as well as smooth terminal apophysis (
<figureCitation id="13182A1AFFADFFCB144F5CB8C731FF5A" box="[550,854,223,249]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146">Fig. 10C, 10F, 10G, 10I–K</figureCitation>
). Conductor situated ventro-proximally on embolus, with two long spines, upper one parallel to the furrow of embolus distally, lower one pointed to contrategulum, and one or two small teeth situated at between two long spines (
<figureCitation id="13182A1AFFADFFCB15165D40C0ABFEE2" box="[895,1228,295,321]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146">Fig. 10A–C, 10D, 10H, 10K</figureCitation>
). Embolus largely sclerotized, with a flat opening and several longitudinal ribs retrolaterally (
<figureCitation id="13182A1AFFADFFCB15B45D2CC0E3FEC6" box="[989,1156,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146">Fig. 10H, 10K</figureCitation>
).
</paragraph>
<caption id="DF5C6617FFADFFCB16FE5BF0C62CF854" ID-DOI="http://doi.org/10.5281/zenodo.5840670" ID-Zenodo-Dep="5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="15" pageNumber="146" startId="15.[151,250,1943,1967]" targetBox="[151,1436,387,1916]" targetPageId="15">
<paragraph id="8B9C369FFFADFFCB16FE5BF0C62CF854" blockId="15.[151,1437,1943,2039]" pageId="15" pageNumber="146">
<emphasis id="B957EA8DFFADFFCB16FE5BF0C578F80C" bold="true" box="[151,287,1943,1967]" pageId="15" pageNumber="146">FIGURE 10.</emphasis>
Male genital anatomy of 
<taxonomicName id="4C234D1CFFADFFCB144A5BF0C7C1F80C" authority="(Schenkel, 1953)" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[547,934,1943,1967]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="schensiensis" status="comb. nov.">
<emphasis id="B957EA8DFFADFFCB144A5BF0C694F80C" box="[547,755,1943,1967]" italics="true" pageId="15" pageNumber="146">Luthela schensiensis</emphasis>
(
<bibRefCitation id="EFB24B6EFFADFFCB15695BF0C7F8F80C" author="Schenkel, E." box="[768,927,1943,1967]" pageId="15" pageNumber="146" pagination="1 - 108" refId="ref14178" refString="Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108." type="journal article" year="1953">Schenkel, 1953</bibRefCitation>
)
</taxonomicName>
<emphasis id="B957EA8DFFADFFCB15C55BF0C078F80C" bold="true" box="[940,1055,1943,1967]" pageId="15" pageNumber="146">
<taxonomicNameLabel id="A26457F6FFADFFCB15C55BF0C078F80C" box="[940,1055,1943,1967]" pageId="15" pageNumber="146" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF5C6617FFB2FFD416FE58B6C0DEFA92" ID-DOI="http://doi.org/10.5281/zenodo.5840672" ID-Zenodo-Dep="5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="16" pageNumber="147" startId="16.[151,250,1233,1257]" targetBox="[151,1436,185,1208]" targetPageId="16">
<paragraph id="8B9C369FFFB2FFD416FE58B6C0DEFA92" blockId="16.[151,1436,1233,1329]" pageId="16" pageNumber="147">
<emphasis id="B957EA8DFFB2FFD416FE58B6C579FB4A" bold="true" box="[151,286,1233,1257]" pageId="16" pageNumber="147">FIGURE 11.</emphasis>
Female genital anatomy of 
<taxonomicName id="4C234D1CFFB2FFD4145158B6C7DBFB4A" authority="(Schenkel, 1953)" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[568,956,1233,1257]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="147" phylum="Arthropoda" rank="species" species="schensiensis" status="comb. nov.">
<emphasis id="B957EA8DFFB2FFD4145158B6C76EFB4A" box="[568,777,1233,1257]" italics="true" pageId="16" pageNumber="147">Luthela schensiensis</emphasis>
(
<bibRefCitation id="EFB24B6EFFB2FFD4157F58B6C7D2FB4A" author="Schenkel, E." box="[790,949,1233,1257]" pageId="16" pageNumber="147" pagination="1 - 108" refId="ref14178" refString="Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108." type="journal article" year="1953">Schenkel, 1953</bibRefCitation>
)
</taxonomicName>
<emphasis id="B957EA8DFFB2FFD415AB58B6C052FB4A" bold="true" box="[962,1077,1233,1257]" pageId="16" pageNumber="147">
<taxonomicNameLabel id="A26457F6FFB2FFD415AB58B6C052FB4A" box="[962,1077,1233,1257]" pageId="16" pageNumber="147" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9C369FFFB2FFD416AE5938C558F98D" blockId="16.[151,1437,1375,2014]" pageId="16" pageNumber="147">Female (XUX-2011-243). Carapace and opisthosoma similar to male; chelicerae robust with promargin of groove with 10 strong denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, similar to male; 8 spinnerets. Measurements: BL 25.25, CL 10.35, CW 9.25, OL 14.75, OW 12.75; ALE&gt; PLE&gt; PME&gt; AME; palp 16.30 (5.70 + 2.75 + 3.35 + 4.50), leg I 17.45 (4.85 + 3.05 + 3.55 + 3.75 + 2.25), leg II 19.15 (6.15 + 3.05 + 3.55 + 3.95 + 2.45), leg III 19.85 (6.05 + 3.15 + 3.45 + 4.45 + 2.75), leg IV 28.80 (7.65 + 3.75 + 5.35 + 8.00 + 4.05).</paragraph>
<paragraph id="8B9C369FFFB2FFD416AE5A50C11CF939" blockId="16.[151,1437,1375,2014]" pageId="16" pageNumber="147">
Genitalia: Posterior margin of genital sternite slightly incurved, two pairs of receptacular clusters, middle ones slightly smaller than lateral ones with stalks, close to each other, situated at anterior margin of bursa copulatrix, lateral ones situated at slightly dorsolateral position of bursa copulatrix with inconspicuous stalks (
<figureCitation id="13182A1AFFB2FFD412805A18C109F939" box="[1257,1390,1663,1690]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="16" pageNumber="147">Fig. 11A–P</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFB2FFD416AE5AC4C71EF91D" box="[199,889,1699,1726]" pageId="16" pageNumber="147" type="distribution">
<paragraph id="8B9C369FFFB2FFD416AE5AC4C71EF91D" blockId="16.[151,1437,1375,2014]" box="[199,889,1699,1726]" pageId="16" pageNumber="147">
<emphasis id="B957EA8DFFB2FFD416AE5AC4C506F91E" bold="true" box="[199,353,1699,1725]" pageId="16" pageNumber="147">Distribution.</emphasis>
<collectingRegion id="49E7F87DFFB2FFD4170E5AC4C64AF91E" box="[359,557,1699,1725]" country="China" name="Shaanxi" pageId="16" pageNumber="147">Shaanxi Province</collectingRegion>
(Heyang, Gaoling, Jingyang)
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFB2FFD516AE5AAFC054FCB6" lastPageId="17" lastPageNumber="148" pageId="16" pageNumber="147" type="discussion">
<paragraph id="8B9C369FFFB2FFD416AE5AAFC535F889" blockId="16.[151,1437,1375,2014]" pageId="16" pageNumber="147">
<emphasis id="B957EA8DFFB2FFD416AE5AAFC55CF941" bold="true" box="[199,315,1736,1762]" pageId="16" pageNumber="147">Remarks.</emphasis>
Males and females vary in body size: males (
<emphasis id="B957EA8DFFB2FFD4155C5AAFC72FF942" box="[821,840,1736,1761]" italics="true" pageId="16" pageNumber="147">N</emphasis>
= 7): BL 14.95–18.55, CL 6.75–7.85, CW 6.25–7.30, OL 7.15–10.45, OW 4.90–7.55; females (
<emphasis id="B957EA8DFFB2FFD4140C5A8BC61FF8A6" box="[613,632,1772,1797]" italics="true" pageId="16" pageNumber="147">N</emphasis>
= 18): BL 17.85–25.65, CL 7.55–10.25, CW 6.90–9.10, OL 9.05–16.65, OW 5.90–13.25.
</paragraph>
<paragraph id="8B9C369FFFB2FFD516AE5B54C6CDFF76" blockId="16.[151,1437,1375,2014]" lastBlockId="17.[151,1437,151,789]" lastPageId="17" lastPageNumber="148" pageId="16" pageNumber="147">
There is intraspecific variation in male and female genital morphology and molecular markers. Specifically, for the male genitalia, there was considerable intraspecific variation in the number of the small middle spines of the conductor, either one (
<figureCitation id="13182A1AFFB2FFD417D65B1CC641F835" box="[447,550,1915,1942]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10A</figureCitation>
) or two spines (
<figureCitation id="13182A1AFFB2FFD414B45B1CC7E2F835" box="[733,901,1915,1942]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10D, 10K</figureCitation>
); in the position of the small spine, either close to the upper spine (
<figureCitation id="13182A1AFFB2FFD417065BF8C5B0F819" box="[367,471,1951,1978]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10K</figureCitation>
), close to the lower spine (
<figureCitation id="13182A1AFFB2FFD4156D5BF8C70BF819" box="[772,876,1951,1978]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10D</figureCitation>
), or located at the middle of two long spines (
<figureCitation id="13182A1AFFB2FFD413065BF8C4ACF87D" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10A</figureCitation>
); in the tip shape of lower spine of the conductor, bifurcated (
<figureCitation id="13182A1AFFB2FFD4151F5BA4C79EF87D" box="[886,1017,1987,2014]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10D–F</figureCitation>
) or not (
<figureCitation id="13182A1AFFB2FFD412315BA4C09AF87D" box="[1112,1277,1987,2014]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="16" pageNumber="147">Fig. 10A, 10H</figureCitation>
). Intraspecific variation was noted in the number of teeth (7–10) along the contrategulum margin, and the different shape of the serrated distal tooth (
<figureCitation id="13182A1AFFB3FFD517EA5CDCC6FBFF76" box="[387,668,187,213]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[151,1436,387,1916]" captionTargetId="figure-117@15.[151,1436,381,1920]" captionTargetPageId="15" captionText="FIGURE 10. Male genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view.A–C, XUX-2011-241 (holotype). D–G, XUX-2011-263. H–K, XUX-2011-264. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840670" httpUri="https://zenodo.org/record/5840670/files/figure.png" pageId="17" pageNumber="148">Fig. 10A, 10D, 10H, 10J</figureCitation>
).
</paragraph>
<paragraph id="8B9C369FFFB3FFD516AE5CB8C126FEE2" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="148">
For intraspecific variation in female genitalia, the posterior margin of the genital sternite may be incurved (
<figureCitation id="13182A1AFFB3FFD513065CB8C53EFEBE" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="17" pageNumber="148">Fig. 11C, 11D, 11I–K</figureCitation>
) or straight (
<figureCitation id="13182A1AFFB3FFD517825D64C6EBFEBE" box="[491,652,259,285]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="17" pageNumber="148">Fig. 11A, 11L</figureCitation>
), the middle receptacular clusters were situated at the anterior margin of the bursa copulatrix (
<figureCitation id="13182A1AFFB3FFD517CA5D40C63BFEE2" box="[419,604,295,321]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="17" pageNumber="148">Fig. 11A–J, 11L</figureCitation>
), or at slightly dorsal part of the bursa copulatrix (
<figureCitation id="13182A1AFFB3FFD512F95D40C153FEE2" box="[1168,1332,295,321]" captionStart="FIGURE 11" captionStartId="16.[151,250,1233,1257]" captionTargetBox="[151,1436,185,1208]" captionTargetId="figure-28@16.[151,1436,181,1209]" captionTargetPageId="16" captionText="FIGURE 11. Female genital anatomy of Luthela schensiensis (Schenkel, 1953) comb. nov. A–D, I–L, vulva dorsal view. E–H, M–P, vulva ventral view. A, E, XUX-2011-243. B, F, XUX-2011-256. C, G, XUX-2011-253. D, H, XUX-2011-254A. I, M, XUX-2011-248. J, N, XUX-2011-249. K, O, XUX-2011-269. L, P, XUX-2011-270. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840672" httpUri="https://zenodo.org/record/5840672/files/figure.png" pageId="17" pageNumber="148">Fig. 11K, 11O</figureCitation>
).
</paragraph>
<paragraph id="8B9C369FFFB3FFD516A45D2CC511FE2A" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="148">
Intraspecific genetic distance ranges from 0% to 1.8% based on the K2P substitution model and 
<emphasis id="B957EA8DFFB3FFD5129B5D2CC145FEC6" box="[1266,1314,331,357]" italics="true" pageId="17" pageNumber="148">COI</emphasis>
nucleotide sequences (
<emphasis id="B957EA8DFFB3FFD517705D08C54BFE2B" box="[281,300,367,392]" italics="true" pageId="17" pageNumber="148">N</emphasis>
= 18).
</paragraph>
<paragraph id="8B9C369FFFB3FFD516AE5DF4C054FCB6" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="148">
We treated 
<taxonomicName id="4C234D1CFFB3FFD517225DF4C592FE0E" authorityName=": Ono" authorityYear="2000" box="[331,501,403,429]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD517225DF4C592FE0E" box="[331,501,403,429]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="4C234D1CFFB3FFD5157A5DF4C7DAFE0E" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[787,957,403,429]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFB3FFD5157A5DF4C7DAFE0E" box="[787,957,403,429]" italics="true" pageId="17" pageNumber="148">L. schensiensis</emphasis>
</taxonomicName>
for several reasons. First, 
<taxonomicName id="4C234D1CFFB3FFD5129A5DF4C1FBFE0E" authorityName=": Ono" authorityYear="2000" box="[1267,1436,403,429]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD5129A5DF4C1FBFE0E" box="[1267,1436,403,429]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
was diagnosed and described based exclusively on 
<specimenCount id="9D25FD16FFB3FFD514B65DD0C73AFE72" box="[735,861,439,465]" pageId="17" pageNumber="148" type="female">one female</specimenCount>
(
<bibRefCitation id="EFB24B6EFFB3FFD515045DD0C020FE72" author="Zhu, C. D. &amp; Wang, Y. W." box="[877,1095,439,465]" pageId="17" pageNumber="148" pagination="251 - 253" refId="ref15094" refString="Zhu, C. D. &amp; Wang, Y. W. (1984) A new species of genus Liphistius (Araneae: Liphistiidae). Journal of the Bethune Medical University, 10, 251 - 253." type="journal article" year="1984">Zhu &amp; Wang 1984</bibRefCitation>
). As liphistiid females exhibit considerable intraspecific variation in female genitalia (
<bibRefCitation id="EFB24B6EFFB3FFD515615DBCC7C9FE56" author="Xu, X. &amp; Liu, F. &amp; Ono, H. &amp; Chen, J. &amp; Kuntner, M. &amp; Li, D." box="[776,942,475,501]" pageId="17" pageNumber="148" pagination="1 - 43" refId="ref14775" refString="Xu, X., Liu, F., Ono, H., Chen, J., Kuntner, M. &amp; Li, D. (2017) Targeted sampling in Ryukyus facilitates species delimitation of the primitively segmented spider genus Ryuthela (Araneae: Mesothelae: Liphistiidae). Zoological Journal of the Linnean Society, 181, 1 - 43. https: // doi. org / 10.1093 / zoolinnean / zlx 024" type="journal article" year="2017">
Xu 
<emphasis id="B957EA8DFFB3FFD5155A5DBCC702FE56" box="[819,869,475,501]" italics="true" pageId="17" pageNumber="148">et al</emphasis>
. 2017
</bibRefCitation>
, 
<bibRefCitation id="EFB24B6EFFB3FFD515D05DBCC792FE56" author="Xu, X. &amp; Ono, H. &amp; Kuntner, M. &amp; Liu, F. &amp; Li, D." box="[953,1013,475,501]" pageId="17" pageNumber="148" pagination="1 - 50" refId="ref14855" refString="Xu, X., Ono, H., Kuntner, M., Liu, F. &amp; Li, D. (2019) A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. Zookeys, 888, 1 - 50. https: // doi. org / 10.3897 / zookeys. 888.34494" type="journal article" year="2019">2019</bibRefCitation>
), it is difficult to diagnose the species based exclusively on female characters of a single specimen. Second, we collected intensively from the 
<typeStatus id="5498883DFFB3FFD5136B5D98C155FDBA" box="[1282,1330,511,537]" pageId="17" pageNumber="148">type</typeStatus>
localities of both 
<taxonomicName id="4C234D1CFFB3FFD516875E44C5F3FD9E" authorityName=": Ono" authorityYear="2000" box="[238,404,547,573]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD516875E44C5F3FD9E" box="[238,404,547,573]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C234D1CFFB3FFD517A35E44C616FD9E" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[458,625,547,573]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFB3FFD517A35E44C616FD9E" box="[458,625,547,573]" italics="true" pageId="17" pageNumber="148">L. schensiensis</emphasis>
</taxonomicName>
, and we examined and compared the newly collected males and females with the original descriptions and illustrations of both 
<taxonomicName id="4C234D1CFFB3FFD514935E20C7C6FDC2" authorityName=": Ono" authorityYear="2000" box="[762,929,583,609]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD514935E20C7C6FDC2" box="[762,929,583,609]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C234D1CFFB3FFD515B15E20C018FDC2" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[984,1151,583,609]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFB3FFD515B15E20C018FDC2" box="[984,1151,583,609]" italics="true" pageId="17" pageNumber="148">L. schensiensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB24B6EFFB3FFD512E65E20C15EFDC2" author="Schenkel, E." box="[1167,1337,582,609]" pageId="17" pageNumber="148" pagination="1 - 108" refId="ref14178" refString="Schenkel, E. (1953) Chinesische Arachnoidea aus dem Museum Hoangho-Peiho in Tientsin. Boletim do Museu Nacional do Rio de Janeiro, 119, 1 - 108." type="journal article" year="1953">Schenkel 1953</bibRefCitation>
; 
<bibRefCitation id="EFB24B6EFFB3FFD5132F5E20C578FD26" author="Song, D. X. &amp; Haupt, J." pageId="17" pageNumber="148" pagination="443 - 451" refId="ref14273" refString="Song, D. X. &amp; Haupt, J. (1984) Comparative morphology and phylogeny of Liphistiomorph spiders (Araneae: Mesothelae). 2. Revision of new Chinese Heptathelid species. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 27, 443 - 451." type="journal article" year="1984">Song &amp; Haupt 1984</bibRefCitation>
; 
<bibRefCitation id="EFB24B6EFFB3FFD517455E0CC663FD26" author="Zhu, C. D. &amp; Wang, Y. W." box="[300,516,619,645]" pageId="17" pageNumber="148" pagination="251 - 253" refId="ref15094" refString="Zhu, C. D. &amp; Wang, Y. W. (1984) A new species of genus Liphistius (Araneae: Liphistiidae). Journal of the Bethune Medical University, 10, 251 - 253." type="journal article" year="1984">Zhu &amp; Wang 1984</bibRefCitation>
). All lines of evidence support the 
<typeStatus id="5498883DFFB3FFD515FA5E0CC7A4FD26" box="[915,963,619,645]" pageId="17" pageNumber="148">type</typeStatus>
female of 
<taxonomicName id="4C234D1CFFB3FFD512565E0CC080FD26" authorityName=": Ono" authorityYear="2000" box="[1087,1255,619,645]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD512565E0CC080FD26" box="[1087,1255,619,645]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
belonging to 
<taxonomicName id="4C234D1CFFB3FFD513EF5E0CC546FD0A" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFB3FFD513EF5E0CC546FD0A" italics="true" pageId="17" pageNumber="148">L. schensiensis</emphasis>
</taxonomicName>
. Third, there is very low genetic variation (1.8%) among the newly collected specimens from the 
<typeStatus id="5498883DFFB3FFD513055EE8C1FBFD0A" box="[1388,1436,655,681]" pageId="17" pageNumber="148">type</typeStatus>
localities of both 
<taxonomicName id="4C234D1CFFB3FFD517325ED4C665FD6E" authorityName=": Ono" authorityYear="2000" box="[347,514,691,717]" class="Arachnida" family="Liphistiidae" genus="Abcathela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="heyangensis">
<emphasis id="B957EA8DFFB3FFD517325ED4C665FD6E" box="[347,514,691,717]" italics="true" pageId="17" pageNumber="148">L. heyangensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C234D1CFFB3FFD514505ED4C686FD6E" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[569,737,691,717]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFB3FFD514505ED4C686FD6E" box="[569,737,691,717]" italics="true" pageId="17" pageNumber="148">L. schensiensis</emphasis>
</taxonomicName>
. Thus, the specimens collected from Heyang County, 
<collectingRegion id="49E7F87DFFB3FFD5132B5ED4C49CFD52" country="China" name="Shaanxi" pageId="17" pageNumber="148">Shaanxi Province</collectingRegion>
should be considered the same species as those collected from both Gaoling and Jingyang Counties, 
<collectingRegion id="49E7F87DFFB3FFD516FE5E9CC538FCB6" box="[151,351,763,789]" country="China" name="Shaanxi" pageId="17" pageNumber="148">Shaanxi Province</collectingRegion>
. The GenBank accession code of XUX-2011-273 is 
<accessionNumber id="9470AB7CFFB3FFD515C35E9CC048FCB6" box="[938,1071,763,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH172701" pageId="17" pageNumber="148">MH172701</accessionNumber>
.
</paragraph>
</subSubSection>
</treatment>
</document>