406 lines
53 KiB
XML
406 lines
53 KiB
XML
<document id="45ED342652A26BB7F74713D4C65B5996" ID-CLB-Dataset="2631" ID-DOI="10.11646/zootaxa.5091.1.5" ID-GBIF-Dataset="b8e67802-3d14-4b63-85c6-6ee00ebc55d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840635" ID-ZooBank="7E744DAE-E9E6-4FB5-9897-1CCF03D373BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641987074997" checkinUser="plazi" docAuthor="Xu, Xin, Yu, Li, Liu, Fengxiang & Li, Daiqin" docDate="2022" docId="038A8789FFA8FFCF16FE5EC0C570FCB6" docLanguage="en" docName="zootaxa.5091.1.5.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luthela handan Xu & Yu & Liu & Li 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="142" masterDocId="FFB3FFF1FFA2FFC416695C67C467FFA3" masterDocTitle="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)" masterLastPageNumber="154" masterPageNumber="131" pageNumber="141" updateTime="1699301488680" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5DC42C5A3BDC051D7E1F4D48E1034553">Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)</mods:title>
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<mods:name id="4235AAFF98CDE893C9D4960C079A50FF" type="personal">
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<mods:namePart id="46C7B373BA02A308414F53199F8B453C">Xu, Xin</mods:namePart>
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<mods:affiliation id="93EB7C7126DF7B4FD5A33E6E4E471CB7">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China & School of Life and Environmental Sciences, University of Sydney, Sydney, NSW, Australia & State Key Laboratory of Biocatalysis and Enzyme Engineering & Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<mods:roleTerm id="D093066604843CCADC4CCCD451E9F12D">Author</mods:roleTerm>
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<mods:namePart id="A356EBFDA4FE00DA5357894F7EFE8788">Yu, Li</mods:namePart>
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<mods:affiliation id="F85DA9F121D258E00A90F3E192B599EF">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China</mods:affiliation>
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<mods:name id="C5818C323E4B6F38EEB42766E9F3A75D" type="personal">
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<mods:namePart id="3735AFB27EBC297513D033C4F56873C3">Liu, Fengxiang</mods:namePart>
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<mods:affiliation id="2F25D700A89862D4FF3C74C9C0DEB037">State Key Laboratory of Biocatalysis and Enzyme Engineering & Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<mods:namePart id="6055A4C2BC99B7E04294EAB98371047A">Li, Daiqin</mods:namePart>
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<mods:affiliation id="2763030FC4EA7703DA74C4DFC5F238D2">Department of Biological Sciences, National University of Singapore, Singapore</mods:affiliation>
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<mods:title id="80709B51A18423372748BFAC5DF2F1B9">Zootaxa</mods:title>
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<mods:part id="5CD15831D183B1CB70D5300A2F668D17">
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<mods:date id="D5BE85E398413D4D6A18B2659DFB8DD5">2022</mods:date>
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<mods:number id="9F8309E82BEB15B475F5F172C723B69F">5091</mods:number>
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<treatment id="038A8789FFA8FFCF16FE5EC0C570FCB6" ID-DOI="http://doi.org/10.5281/zenodo.5840633" ID-GBIF-Taxon="192044610" ID-Zenodo-Dep="5840633" LSID="urn:lsid:plazi:treatment:038A8789FFA8FFCF16FE5EC0C570FCB6" httpUri="http://treatment.plazi.org/id/038A8789FFA8FFCF16FE5EC0C570FCB6" lastPageId="11" lastPageNumber="142" pageId="10" pageNumber="141">
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<subSubSection id="C3396514FFA8FFCE16FE5EC0C5D8FD61" box="[151,447,679,706]" pageId="10" pageNumber="141" type="nomenclature">
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<paragraph id="8B9C369FFFA8FFCE16FE5EC0C5D8FD61" blockId="10.[151,447,679,743]" box="[151,447,679,706]" pageId="10" pageNumber="141">
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<emphasis id="B957EA8DFFA8FFCE16FE5EC0C5D8FD61" bold="true" box="[151,447,679,706]" pageId="10" pageNumber="141">
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<taxonomicName id="4C234D1CFFA8FFCE16FE5EC0C53DFD61" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[151,346,679,706]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
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<emphasis id="B957EA8DFFA8FFCE16FE5EC0C53DFD61" bold="true" box="[151,346,679,706]" italics="true" pageId="10" pageNumber="141">Luthela handan</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26457F6FFA8FFCE17085ECFC5D8FD61" box="[353,447,680,706]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C3396514FFA8FFCE16FE5EABC56CFD45" box="[151,267,716,743]" pageId="10" pageNumber="141" type="description">
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<paragraph id="8B9C369FFFA8FFCE16FE5EABC56CFD45" blockId="10.[151,447,679,743]" box="[151,267,716,743]" pageId="10" pageNumber="141">
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(
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<figureCitation id="13182A1AFFA8FFCE16F65EABC48FFD44" box="[159,232,716,743]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Figs 7</figureCitation>
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,
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<figureCitation id="13182A1AFFA8FFCE169D5EABC565FD45" box="[244,258,716,742]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="10" pageNumber="141">8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396514FFA8FFCE16FE5F73C088FBED" pageId="10" pageNumber="141" type="materials_examined">
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<paragraph id="8B9C369FFFA8FFCE16FE5F73C5D0FCD5" blockId="10.[151,1437,788,2039]" pageId="10" pageNumber="141">
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<materialsCitation id="3B4B3CC2FFA8FFCE16FE5F73C5D0FCD5" ID-GBIF-Occurrence="3435942309" collectingDate="2011-11-23" collectingDateMax="2012-07-10" collectingDateMin="2011-11-23" collectorName="H. Z. Li & F. X. Liu & X. Y. Wang & X. Xu" country="China" county="Yongnian County" elevation="58" latitude="36.76" location="Male" longLatPrecision="70" longitude="114.476" municipality="Handan City" pageId="10" pageNumber="141" specimenCount="1" stateProvince="Hebei" typeStatus="holotype">
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<emphasis id="B957EA8DFFA8FFCE16FE5F73C56CFC8D" bold="true" box="[151,267,788,814]" pageId="10" pageNumber="141">
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<typeStatus id="5498883DFFA8FFCE16FE5F73C561FC8D" box="[151,262,788,814]" pageId="10" pageNumber="141" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8EFC6044FFA8FFCE177D5F73C529FC8D" LSID="urn:lsid:plazi:treatment:038A8789FFA8FFCF16FE5EC0C570FCB6:8EFC6044FFA8FFCE177D5F73C529FC8D" box="[276,334,788,814]" country="China" county="Yongnian County" latitude="36.76" longLatPrecision="70" longitude="114.476" municipality="Handan City" name="Male" pageId="10" pageNumber="141" stateProvince="Hebei">Male</location>
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(XUX-2011-214, matured on
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<date id="FF9D105FFFA8FFCE14C35F73C758FC8D" box="[682,831,788,814]" pageId="10" pageNumber="141" value="2012-07-10">
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<collectingDate id="EFD9E9B7FFA8FFCE14C35F73C758FC8D" box="[682,831,788,814]" pageId="10" pageNumber="141" value="2012-07-10">10 July 2012</collectingDate>
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</date>
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at CBEE),
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<location id="8EFC6044FFA8FFCE15AF5F73C042FC8D" LSID="urn:lsid:plazi:treatment:038A8789FFA8FFCF16FE5EC0C570FCB6:8EFC6044FFA8FFCE15AF5F73C042FC8D" box="[966,1061,788,814]" country="China" county="Yongnian County" latitude="36.76" longLatPrecision="70" longitude="114.476" municipality="Handan City" name="Qilidian" pageId="10" pageNumber="141" stateProvince="Hebei">Qilidian</location>
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,
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<collectingCounty id="62FD4E13FFA8FFCE12585F73C09DFC8D" box="[1073,1274,788,814]" pageId="10" pageNumber="141">Yongnian County</collectingCounty>
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,
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<collectingMunicipality id="6BF8ACE5FFA8FFCE136E5F73C1FFFC8D" box="[1287,1432,788,814]" pageId="10" pageNumber="141">Handan City</collectingMunicipality>
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,
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<collectingRegion id="49E7F87DFFA8FFCE16FE5F5FC52EFCF1" box="[151,329,824,850]" country="China" name="Hebei" pageId="10" pageNumber="141">Hebei Province</collectingRegion>
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,
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<collectingCountry id="F334760FFFA8FFCE173E5F5FC5FAFCF1" box="[343,413,824,850]" name="China" pageId="10" pageNumber="141">China</collectingCountry>
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,
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<geoCoordinate id="EE175058FFA8FFCE17C35F5FC67EFCF1" box="[426,537,824,850]" degrees="36.760" direction="north" orientation="latitude" pageId="10" pageNumber="141" precision="55" value="36.76">36.760°N</geoCoordinate>
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,
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<geoCoordinate id="EE175058FFA8FFCE144F5F5FC6C7FCF1" box="[550,672,824,850]" degrees="114.476" direction="east" orientation="longitude" pageId="10" pageNumber="141" precision="55" value="114.476">114.476°E</geoCoordinate>
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,
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<quantity id="4CDB9B7AFFA8FFCE14C45F5FC68DFCF0" box="[685,746,824,851]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="10" pageNumber="141" unit="m" value="58.0">
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<elevation id="000ED1ACFFA8FFCE14C45F5FC68DFCF0" box="[685,746,824,851]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="10" pageNumber="141" unit="m" value="58.0">58 m</elevation>
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</quantity>
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, collected on
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<date id="FF9D105FFFA8FFCE15E25F5FC001FCF1" box="[907,1126,824,850]" pageId="10" pageNumber="141" value="2011-11-23">
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<collectingDate id="EFD9E9B7FFA8FFCE15E25F5FC001FCF1" box="[907,1126,824,850]" pageId="10" pageNumber="141" value="2011-11-23">23 November 2011</collectingDate>
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</date>
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by
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<collectorName id="26D65349FFA8FFCE12FD5F5EC08BFCF1" box="[1172,1260,824,851]" pageId="10" pageNumber="141">H.Z. Li</collectorName>
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,
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<collectorName id="26D65349FFA8FFCE12935F5EC13BFCF1" box="[1274,1372,824,851]" pageId="10" pageNumber="141">F.X. Liu</collectorName>
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,
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||
<collectorName id="26D65349FFA8FFCE13005F5EC4BCFCD4" pageId="10" pageNumber="141">X.Y. Wang</collectorName>
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||
, and
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||
<collectorName id="26D65349FFA8FFCE177C5F3AC53BFCD4" box="[277,348,861,887]" pageId="10" pageNumber="141">X. Xu</collectorName>
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, CBEE.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9C369FFFA8FFCE16AE5FE7C088FBED" blockId="10.[151,1437,788,2039]" pageId="10" pageNumber="141">
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<emphasis id="B957EA8DFFA8FFCE16AE5FE7C521FC39" bold="true" box="[199,326,896,922]" pageId="10" pageNumber="141">
|
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<typeStatus id="5498883DFFA8FFCE16AE5FE7C526FC39" box="[199,321,896,922]" pageId="10" pageNumber="141" type="paratype">Paratypes</typeStatus>
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.
|
||
</emphasis>
|
||
<materialsCitation id="3B4B3CC2FFA8FFCE17265FE7C0E2FC1D" ID-GBIF-Occurrence="3435942323" collectingDate="2013-06-12" collectingDateMax="2012-07-10" collectingDateMin="2011-11-23" collectorName="H. Z. Li & F. X. Liu & X. Y. Wang & X. Xu" country="China" county="Yongnian County" elevation="58" latitude="36.76" location="Male" longLatPrecision="70" longitude="114.476" municipality="Handan City" pageId="10" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Hebei" typeStatus="paratype">
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<specimenCount id="9D25FD16FFA8FFCE17265FE7C5DAFC39" box="[335,445,896,922]" pageId="10" pageNumber="141" type="female">4 females</specimenCount>
|
||
(XUX-2011-206/207/208/215, 208 and 215 matured on
|
||
<date id="FF9D105FFFA8FFCE122E5FE7C0BBFC39" box="[1095,1244,896,922]" pageId="10" pageNumber="141" value="2012-07-10">10 July 2012</date>
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at CBEE) and
|
||
<specimenCount id="9D25FD16FFA8FFCE13E75FE7C4AAFC1D" pageId="10" pageNumber="141" type="male">1 male</specimenCount>
|
||
(XUX-2011-209, matured on
|
||
<date id="FF9D105FFFA8FFCE14465FC3C6ACFC1D" box="[559,715,932,959]" pageId="10" pageNumber="141" value="2013-06-12">
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||
<collectingDate id="EFD9E9B7FFA8FFCE14465FC3C6ACFC1D" box="[559,715,932,959]" pageId="10" pageNumber="141" value="2013-06-12">12 June 2013</collectingDate>
|
||
</date>
|
||
at CBEE), same data as for holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA8FFCE12FA5FC3C73CFBA5" ID-GBIF-Occurrence="3435942314" collectingDate="2011-11-23" country="China" county="Yongnian County" elevation="66" latitude="36.752" location="University of Engineering" longLatPrecision="70" longitude="114.471" municipality="Handan City" pageId="10" pageNumber="141" specimenCount="3" specimenCount-female="3" stateProvince="Hebei" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA8FFCE12FA5FC3C165FC1D" box="[1171,1282,932,958]" pageId="10" pageNumber="141" type="female">3 females</specimenCount>
|
||
(XUX-2011- 202/203/205), nearby
|
||
<collectingRegion id="49E7F87DFFA8FFCE17FC5FAFC5BFFC41" box="[405,472,968,994]" country="China" name="Hebei" pageId="10" pageNumber="141">Hebei</collectingRegion>
|
||
<location id="8EFC6044FFA8FFCE17885FAFC769FC41" LSID="urn:lsid:plazi:treatment:038A8789FFA8FFCF16FE5EC0C570FCB6:8EFC6044FFA8FFCE17885FAFC769FC41" box="[481,782,968,994]" country="China" county="Yongnian County" latitude="36.752" longLatPrecision="70" longitude="114.471" municipality="Handan City" name="University of Engineering" pageId="10" pageNumber="141" stateProvince="Hebei">University of Engineering</location>
|
||
,
|
||
<collectingCounty id="62FD4E13FFA8FFCE15725FAFC783FC41" box="[795,996,968,994]" pageId="10" pageNumber="141">Yongnian County</collectingCounty>
|
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,
|
||
<collectingMunicipality id="6BF8ACE5FFA8FFCE15985FAFC0E4FC41" box="[1009,1155,968,994]" pageId="10" pageNumber="141">Handan City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA8FFCE12F85FAFC123FC41" box="[1169,1348,968,994]" country="China" name="Hebei" pageId="10" pageNumber="141">Hebei Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA8FFCE133B5FAFC1FFFC41" box="[1362,1432,968,994]" name="China" pageId="10" pageNumber="141">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA8FFCE16FE5F8BC561FBA4" box="[151,262,1004,1031]" degrees="36.752" direction="north" orientation="latitude" pageId="10" pageNumber="141" precision="55" value="36.752">36.752°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA8FFCE177D5F8BC5EAFBA5" box="[276,397,1004,1030]" degrees="114.471" direction="east" orientation="longitude" pageId="10" pageNumber="141" precision="55" value="114.471">114.471°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA8FFCE17F25F8BC5BEFBA4" box="[411,473,1004,1031]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="141" unit="m" value="66.0">
|
||
<elevation id="000ED1ACFFA8FFCE17F25F8BC5BEFBA4" box="[411,473,1004,1031]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="141" unit="m" value="66.0">66 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA8FFCE14155F8BC73CFBA5" box="[636,859,1004,1030]" pageId="10" pageNumber="141" value="2011-11-23">
|
||
<collectingDate id="EFD9E9B7FFA8FFCE14155F8BC73CFBA5" box="[636,859,1004,1030]" pageId="10" pageNumber="141" value="2011-11-23">23 November 2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA8FFCE15035F8BC08CFBED" ID-GBIF-Occurrence="3435942306" collectingDate="2011-11-25" collectorName="H. Z. Li & F. X. Liu & X. Y. Wang & X. Xu. All" country="China" county="Luquan County" elevation="160" latitude="38.074" location="Dongtumen Village" longLatPrecision="70" longitude="114.282" municipality="Shijiazhang City" pageId="10" pageNumber="141" specimenCount="9" specimenCount-female="9" stateProvince="Hebei" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA8FFCE15035F8BC7BFFBA5" box="[874,984,1004,1030]" pageId="10" pageNumber="141" type="female">9 females</specimenCount>
|
||
(XUX-2011-217/218/219/220/221/222/ 223/224/227)
|
||
<location id="8EFC6044FFA8FFCE175D5876C675FB89" LSID="urn:lsid:plazi:treatment:038A8789FFA8FFCF16FE5EC0C570FCB6:8EFC6044FFA8FFCE175D5876C675FB89" box="[308,530,1040,1067]" country="China" county="Luquan County" latitude="38.074" longLatPrecision="70" longitude="114.282" municipality="Shijiazhang City" name="Dongtumen Village" pageId="10" pageNumber="141" stateProvince="Hebei">Dongtumen Village</location>
|
||
,
|
||
<collectingCounty id="62FD4E13FFA8FFCE14755876C6ABFB89" box="[540,716,1040,1067]" pageId="10" pageNumber="141">Luquan County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BF8ACE5FFA8FFCE14BF5877C7F6FB89" box="[726,913,1040,1066]" pageId="10" pageNumber="141">Shijiazhang City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA8FFCE15F55877C02CFB89" box="[924,1099,1040,1066]" country="China" name="Hebei" pageId="10" pageNumber="141">Hebei Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA8FFCE123C5877C0FCFB89" box="[1109,1179,1040,1066]" name="China" pageId="10" pageNumber="141">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA8FFCE12CC5877C173FB89" box="[1189,1300,1040,1066]" degrees="38.074" direction="north" orientation="latitude" pageId="10" pageNumber="141" precision="55" value="38.074">38.074°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA8FFCE13765877C1FFFB89" box="[1311,1432,1040,1066]" degrees="114.282" direction="east" orientation="longitude" pageId="10" pageNumber="141" precision="55" value="114.282">114.282°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA8FFCE16FE5853C487FBEC" box="[151,224,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="141" unit="m" value="160.0">
|
||
<elevation id="000ED1ACFFA8FFCE16FE5853C487FBEC" box="[151,224,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="141" unit="m" value="160.0">160 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA8FFCE17125853C632FBED" box="[379,597,1076,1103]" pageId="10" pageNumber="141" value="2011-11-25">
|
||
<collectingDate id="EFD9E9B7FFA8FFCE17125853C632FBED" box="[379,597,1076,1103]" pageId="10" pageNumber="141" value="2011-11-25">25 November 2011</collectingDate>
|
||
</date>
|
||
, by
|
||
<collectorName id="26D65349FFA8FFCE14EA5852C6BFFBED" box="[643,728,1076,1103]" pageId="10" pageNumber="141">H.Z. Li</collectorName>
|
||
,
|
||
<collectorName id="26D65349FFA8FFCE148D5852C722FBED" box="[740,837,1076,1103]" pageId="10" pageNumber="141">F.X. Liu</collectorName>
|
||
,
|
||
<collectorName id="26D65349FFA8FFCE15395852C7AAFBEC" box="[848,973,1077,1103]" pageId="10" pageNumber="141">X.Y. Wang</collectorName>
|
||
, and
|
||
<collectorName id="26D65349FFA8FFCE126E5852C01BFBED" box="[1031,1148,1076,1103]" pageId="10" pageNumber="141">X. Xu. All</collectorName>
|
||
in CBEE
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA8FFCE16AE583FC55DFA31" pageId="10" pageNumber="141" type="diagnosis">
|
||
<paragraph id="8B9C369FFFA8FFCE16AE583FC55DFA31" blockId="10.[151,1437,788,2039]" pageId="10" pageNumber="141">
|
||
<emphasis id="B957EA8DFFA8FFCE16AE583FC525FBD1" bold="true" box="[199,322,1112,1138]" pageId="10" pageNumber="141">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C234D1CFFA8FFCE17C7583EC679FBD1" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[430,542,1112,1138]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE17C7583EC679FBD1" box="[430,542,1112,1138]" italics="true" pageId="10" pageNumber="141">L. handan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE144A583FC61CFBD1" bold="true" box="[547,635,1112,1138]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE144A583FC61CFBD1" box="[547,635,1112,1138]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from those of
|
||
<taxonomicName id="4C234D1CFFA8FFCE126B583EC0E2FBD1" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1026,1157,1112,1138]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE126B583EC0E2FBD1" box="[1026,1157,1112,1138]" italics="true" pageId="10" pageNumber="141">L. dengfeng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE12E3583FC085FBD1" bold="true" box="[1162,1250,1112,1138]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE12E3583FC085FBD1" box="[1162,1250,1112,1138]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C234D1CFFA8FFCE1286583EC1F2FBD1" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[1263,1429,1112,1138]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="schensiensis">
|
||
<emphasis id="B957EA8DFFA8FFCE1286583EC1F2FBD1" box="[1263,1429,1112,1138]" italics="true" pageId="10" pageNumber="141">L. schensiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D1CFFA8FFCE16FE581AC566FB35" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[151,257,1149,1174]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE16FE581AC566FB35" box="[151,257,1149,1174]" italics="true" pageId="10" pageNumber="141">L. yiyuan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE1761581BC504FB35" bold="true" box="[264,355,1148,1174]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE1761581BC504FB35" box="[264,355,1148,1174]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C234D1CFFA8FFCE17CB581AC64CFB35" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[418,555,1148,1174]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE17CB581AC64CFB35" box="[418,555,1148,1174]" italics="true" pageId="10" pageNumber="141">L. yuncheng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE145A581BC6E9FB35" bold="true" box="[563,654,1148,1174]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE145A581BC6E9FB35" box="[563,654,1148,1174]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the conductor with a longer lower spine, and the dorsal extension of terminal apophysis of tegulum with regular serrations (
|
||
<figureCitation id="13182A1AFFA8FFCE157158C7C7F5FB19" box="[792,914,1184,1210]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7E–G</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA8FFCE122958C6C0D6FB19" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1088,1201,1184,1210]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE122958C6C0D6FB19" box="[1088,1201,1184,1210]" italics="true" pageId="10" pageNumber="141">L. badong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE12D158C7C176FB19" bold="true" box="[1208,1297,1184,1210]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE12D158C7C176FB19" box="[1208,1297,1184,1210]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C234D1CFFA8FFCE132F58C6C1FAFB19" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1350,1437,1185,1210]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE132F58C6C1FAFB19" box="[1350,1437,1185,1210]" italics="true" pageId="10" pageNumber="141">L. taian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE16FE58A3C497FB7D" bold="true" box="[151,240,1220,1246]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE16FE58A3C497FB7D" box="[151,240,1220,1246]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the conductor with small middle spines (
|
||
<figureCitation id="13182A1AFFA8FFCE14B458A3C703FB7C" box="[733,868,1220,1247]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7B, 7D</figureCitation>
|
||
), and lower spine thicker and longer (
|
||
<figureCitation id="13182A1AFFA8FFCE136158A3C1E9FB7D" box="[1288,1422,1220,1247]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7E, 7G</figureCitation>
|
||
); Females of
|
||
<taxonomicName id="4C234D1CFFA8FFCE1775588EC5E8FAA1" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[284,399,1256,1282]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE1775588EC5E8FAA1" box="[284,399,1256,1282]" italics="true" pageId="10" pageNumber="141">L. handan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE17FE588FC596FAA1" bold="true" box="[407,497,1256,1282]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE17FE588FC596FAA1" box="[407,497,1256,1282]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from those of
|
||
<taxonomicName id="4C234D1CFFA8FFCE1489588EC734FAA1" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[736,851,1256,1282]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE1489588EC734FAA1" box="[736,851,1256,1282]" italics="true" pageId="10" pageNumber="141">L. badong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE1532588FC7D1FAA1" bold="true" box="[859,950,1256,1282]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE1532588FC7D1FAA1" box="[859,950,1256,1282]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C234D1CFFA8FFCE15AC588EC00CFAA1" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" baseAuthorityName="Yin, Tang, Zhao & Chen" baseAuthorityYear="2002" box="[965,1131,1256,1282]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="luotianensis">
|
||
<emphasis id="B957EA8DFFA8FFCE15AC588EC00CFAA1" box="[965,1131,1256,1282]" italics="true" pageId="10" pageNumber="141">L. luotianensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D1CFFA8FFCE1213588EC083FAA1" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1146,1252,1257,1282]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE1213588EC083FAA1" box="[1146,1252,1257,1282]" italics="true" pageId="10" pageNumber="141">L. yiyuan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE1282588FC121FAA1" bold="true" box="[1259,1350,1256,1282]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE1282588FC121FAA1" box="[1259,1350,1256,1282]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C234D1CFFA8FFCE13EF588EC565FA85" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE13EF588EC565FA85" italics="true" pageId="10" pageNumber="141">L. yuncheng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE1760596BC505FA85" bold="true" box="[265,354,1292,1318]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE1760596BC505FA85" box="[265,354,1292,1318]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the middle receptacular clusters smaller than lateral ones (
|
||
<figureCitation id="13182A1AFFA8FFCE127A596BC0B2FA85" box="[1043,1237,1292,1318]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="10" pageNumber="141">Fig. 8A–C, 8G–I</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA8FFCE13EF596AC546FAE9" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="schensiensis">
|
||
<emphasis id="B957EA8DFFA8FFCE13EF596AC546FAE9" italics="true" pageId="10" pageNumber="141">L. schensiensis</emphasis>
|
||
</taxonomicName>
|
||
by the lateral receptacular clusters with slightly longer stalks (
|
||
<figureCitation id="13182A1AFFA8FFCE158E5957C050FAE9" box="[999,1079,1328,1354]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="10" pageNumber="141">Fig. 8J</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA8FFCE12875956C114FAE9" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1262,1395,1328,1354]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE12875956C114FAE9" box="[1262,1395,1328,1354]" italics="true" pageId="10" pageNumber="141">L. dengfeng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE13125957C4AFFACD" bold="true" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE13125957C4AFFACD" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C234D1CFFA8FFCE16965932C531FACD" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[255,342,1365,1390]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA8FFCE16965932C531FACD" box="[255,342,1365,1390]" italics="true" pageId="10" pageNumber="141">L. taian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA8FFCE17345933C5D1FACD" bold="true" box="[349,438,1364,1390]" pageId="10" pageNumber="141">
|
||
<taxonomicNameLabel id="A26457F6FFA8FFCE17345933C5D1FACD" box="[349,438,1364,1390]" pageId="10" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the middle receptacular clusters with short stalks and close to each other basally (
|
||
<figureCitation id="13182A1AFFA8FFCE13065933C54AFA31" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="10" pageNumber="141">Fig. 8A–F, 8H, 8I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA8FFCF16AE59FBC5B4FE2A" lastPageId="11" lastPageNumber="142" pageId="10" pageNumber="141" type="description">
|
||
<paragraph id="8B9C369FFFA8FFCE16AE59FBC5BBF92C" blockId="10.[151,1437,788,2039]" pageId="10" pageNumber="141">
|
||
<emphasis id="B957EA8DFFA8FFCE16AE59FBC53EFA15" bold="true" box="[199,345,1436,1462]" pageId="10" pageNumber="141">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="5498883DFFA8FFCE17C359FBC67EFA14" box="[426,537,1436,1463]" pageId="10" pageNumber="141" type="holotype">Holotype</typeStatus>
|
||
). Carapace brown; opisthosoma light brown, with dark brown tergites; sternum narrow, much longer than wide; few long pointed hairs running over ocular mound in longitudinal row; chelicerae robust with promargin of groove with 10 denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, the second to fifth larger than others and the others small; 7 spinnerets. Measurements: BL 15.55, CL 7.15, CW 6.85, OL 8.35, OW 5.65; ALE> PLE> PME> AME; leg I 19.95 (5.95 + 2.65 + 4.05 + 4.95 + 2.35), leg II 19.95 (5.80 + 2.60 + 3.55 + 5.05 + 2.95), leg III 20.55 (5.05 + 2.70 + 3.55 + 5.90 + 3.35), leg IV 26.65 (6.65 + 3.00 + 5.00 + 8.05 + 3.95).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA8FFCE16AE5AFFC4ADF80D" blockId="10.[151,1437,788,2039]" pageId="10" pageNumber="141">
|
||
Palp: Prolateral paracymbium unpigmented and unsclerotized, numerous setae and spines at tip of paracymbium (
|
||
<figureCitation id="13182A1AFFA8FFCE16C95ADBC57AF975" box="[160,285,1724,1750]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7A–C</figureCitation>
|
||
). Contrategulum with 6 teeth along margin, and distal one with 3 serrations (
|
||
<figureCitation id="13182A1AFFA8FFCE12FF5ADBC147F974" box="[1174,1312,1724,1751]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7D, 7F</figureCitation>
|
||
). Marginal apophysis of tegulum with few serrations proximally and smooth distally, dorsal extension of terminal apophysis of tegulum with regular serrations, and terminal apophysis of tegulum smooth, thumb-shaped (
|
||
<figureCitation id="13182A1AFFA8FFCE12F35B63C173F8BD" box="[1178,1300,1796,1822]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7E–G</figureCitation>
|
||
). Conductor situated ventro-proximally on embolus, with two long spines, upper one parallel to embolus, lower one pointed to proximal of contrategulum, and one or two inconspicuous, small spines situated at middle between two long spines (
|
||
<figureCitation id="13182A1AFFA8FFCE16C95B17C578F829" box="[160,287,1904,1930]" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7D–G</figureCitation>
|
||
). Embolus largely sclerotized, with flat opening and several longitudinal ribs retrolaterally (
|
||
<figureCitation id="13182A1AFFA8FFCE13545B17C4DAF80D" captionStart="FIGURE 7" captionStartId="11.[151,250,1843,1867]" captionTargetBox="[152,1436,813,1811]" captionTargetId="figure-292@11.[151,1436,813,1820]" captionTargetPageId="11" captionText="FIGURE 7. Male genital anatomy of Luthela handan sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. A–C, XUX-2011-214 (holotype). D–G, XUX-2011-209. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840661" httpUri="https://zenodo.org/record/5840661/files/figure.png" pageId="10" pageNumber="141">Fig. 7D, 7G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA8FFCF16AE5BDFC119FF5A" blockId="10.[151,1437,788,2039]" lastBlockId="11.[151,1437,150,789]" lastPageId="11" lastPageNumber="142" pageId="10" pageNumber="141">Female (XUX-2011-224). Carapace and opisthosoma light brown; chelicerae robust with promargin of groove with 9–12 strong denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, similar to male; 7 spinnerets. Measurements: BL 17.35, CL 8.05, CW 6.95, OL 8.65, OW 6.30; ALE> PLE> PME> AME; palp 13.75 (4.65 + 2.40 + 3.10 + 3.60), leg I 15.85 (5.25 + 2.85 + 3.15 + 3.20 + 1.40), leg II 14.75 (4.70 + 2.85 + 2.60 + 3.25 + 1.40), leg III 16.30 (4.50 + 2.65 + 2.75 + 3.95 + 2.45), leg IV 22.45 (4.85 + 3.35 + 4.35 + 6.45 + 3.45).</paragraph>
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5D64C5B4FE2A" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="142">
|
||
Genitalia: Posterior margin of genital sternite slightly incurved (
|
||
<figureCitation id="13182A1AFFA9FFCF15FF5D64C077FEBE" box="[918,1040,259,285]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8A–L</figureCitation>
|
||
), two pairs of receptacular clusters, middle ones smaller than lateral ones with slender stalks, separated from each other (
|
||
<figureCitation id="13182A1AFFA9FFCF120B5D40C157FEE2" box="[1122,1328,295,321]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8B, 8F, 8G–I</figureCitation>
|
||
), situated at anterior margin of bursa copulatrix, lateral ones situated at slightly dorsolateral position of bursa copulatrix with short stalks (
|
||
<figureCitation id="13182A1AFFA9FFCF174C5D08C5A1FE2A" box="[293,454,367,393]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8C, 8G–I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA9FFCF16AE5DF4C02FFE0E" box="[199,1096,403,429]" pageId="11" pageNumber="142" type="etymology">
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5DF4C02FFE0E" blockId="11.[151,1437,150,789]" box="[199,1096,403,429]" pageId="11" pageNumber="142">
|
||
<emphasis id="B957EA8DFFA9FFCF16AE5DF4C529FE0E" bold="true" box="[199,334,403,429]" pageId="11" pageNumber="142">Etymology.</emphasis>
|
||
The species epithet, a noun in apposition, refers to the
|
||
<typeStatus id="5498883DFFA9FFCF15DE5DF4C780FE0E" box="[951,999,403,429]" pageId="11" pageNumber="142">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA9FFCF16AE5DD1C742FE71" box="[199,805,438,466]" pageId="11" pageNumber="142" type="distribution">
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5DD1C742FE71" blockId="11.[151,1437,150,789]" box="[199,805,438,466]" pageId="11" pageNumber="142">
|
||
<emphasis id="B957EA8DFFA9FFCF16AE5DD1C506FE73" bold="true" box="[199,353,438,464]" pageId="11" pageNumber="142">Distribution.</emphasis>
|
||
<collectingRegion id="49E7F87DFFA9FFCF170E5DD0C672FE72" box="[359,533,439,465]" country="China" name="Hebei" pageId="11" pageNumber="142">Hebei Province</collectingRegion>
|
||
(Handan, Shijiazhuang)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA9FFCF16AE5DBCC570FCB6" pageId="11" pageNumber="142" type="discussion">
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5DBCC535FD9E" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="142">
|
||
<emphasis id="B957EA8DFFA9FFCF16AE5DBCC55CFE56" bold="true" box="[199,315,475,501]" pageId="11" pageNumber="142">Remarks.</emphasis>
|
||
Males and females vary in body size: males (
|
||
<emphasis id="B957EA8DFFA9FFCF155C5DBCC72FFE57" box="[821,840,475,500]" italics="true" pageId="11" pageNumber="142">N</emphasis>
|
||
= 2): BL 14.75–16.20, CL 6.35–7.25, CW 5.50–6.85, OL 7.40–8.55, OW 5.00–6.15; females (
|
||
<emphasis id="B957EA8DFFA9FFCF14355D98C608FDBB" box="[604,623,511,536]" italics="true" pageId="11" pageNumber="142">N</emphasis>
|
||
= 16): BL 17.35–26.95, CL 8.05–10.15, CW 6.95–9.45, OL 8.65–16.16, OW 6.30–15.35.
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5E20C0E0FD0A" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="142">
|
||
Intraspecific variation was found in female genital morphology and molecular markers. For female genitalia, the middle receptacular clusters are sclerotized well (
|
||
<figureCitation id="13182A1AFFA9FFCF149F5E0CC714FD26" box="[758,883,619,645]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8A–C</figureCitation>
|
||
), or weakly (
|
||
<figureCitation id="13182A1AFFA9FFCF12635E0CC019FD26" box="[1034,1150,619,645]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8G–I</figureCitation>
|
||
); the bases of the middle stalks are fused together (
|
||
<figureCitation id="13182A1AFFA9FFCF17DF5EE8C658FD0A" box="[438,575,655,681]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8A, 8D</figureCitation>
|
||
), or separated from each other (
|
||
<figureCitation id="13182A1AFFA9FFCF15CD5EE8C01CFD0A" box="[932,1147,655,681]" captionStart="FIGURE 8" captionStartId="12.[151,250,1584,1608]" captionTargetBox="[151,1436,185,1558]" captionTargetId="figure-28@12.[151,1436,181,1560]" captionTargetPageId="12" captionText="FIGURE 8. Female genital anatomy of Luthela handan sp. nov. A–C, G–I, vulva dorsal view. D–F, J–L, vulva ventral view. A, D, XUX-2011-203. B, E, XUX-2011-218. C, F, XUX-2011-206. G, J, XUX-2011-222. H, K, XUX-2011-224. I, L, XUX-2011- 227. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840664" httpUri="https://zenodo.org/record/5840664/files/figure.png" pageId="11" pageNumber="142">Fig. 8B, 8C, 8G–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA9FFCF16AE5ED4C570FCB6" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="142">
|
||
Intraspecific genetic distance ranges from 0% to 3.5% based on the K2P substitution model and
|
||
<emphasis id="B957EA8DFFA9FFCF12985ED4C146FD6E" box="[1265,1313,691,717]" italics="true" pageId="11" pageNumber="142">COI</emphasis>
|
||
nucleotide sequences (
|
||
<emphasis id="B957EA8DFFA9FFCF17495EB0C554FD53" box="[288,307,727,752]" italics="true" pageId="11" pageNumber="142">N</emphasis>
|
||
= 14, including juveniles). The GenBank accession code of the
|
||
<typeStatus id="5498883DFFA9FFCF12205EB0C0CCFD52" box="[1097,1195,727,753]" pageId="11" pageNumber="142" type="holotype">holotype</typeStatus>
|
||
(XUX-2011-214) is
|
||
<accessionNumber id="9470AB7CFFA9FFCF16FE5E9CC574FCB6" box="[151,275,763,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP229810" pageId="11" pageNumber="142">KP229810</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |