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<mods:title id="F3799268CF0A291A98F520BC06AF1B2E">Hermit crabs (Decapoda: Crustacea) from deep Mauritanian waters (NW Africa) with the description of a new species</mods:title>
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<taxonomicName id="4C2A0FE08614FFBB12E8D0839E56FF17" ID-CoL="632fd023-bb48-4d7c-81c4-b86bf2d10e9c" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[151,534,221,247]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB12E8D0839E56FF17" bold="true" box="[151,534,221,247]" pageId="23" pageNumber="174">
<emphasis id="B95EA8718614FFBB12E8D0839D0EFF17" bold="true" box="[151,334,221,247]" italics="true" pageId="23" pageNumber="174">Pagurus alatus</emphasis>
<bibRefCitation id="EFBB09928614FFBB132AD0839E56FF17" author="Fabricius" box="[341,534,221,247]" pageId="23" pageNumber="174" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, observationibus. Flensburgi et Lipsiae, xxxii, 832 pp." type="book" year="1775">Fabricius, 1775</bibRefCitation>
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(
<figureCitation id="131168E68614FFBB12E0D0A19CB0FEF8" box="[159,240,255,280]" captionStart="FIGURE 15" captionStartId="24.[151,250,1956,1978]" captionTargetBox="[268,1331,363,1937]" captionTargetId="figure@24.[268,1332,363,1938]" captionTargetPageId="24" captionText="FIGURE 15. Pagurus alatus Fabricius, 1775. Adult male, sl: 6.56 mm (MU 57; UVIGOBA 3 04737): A, shield and cephalic appendages. B, chelipeds: left (top), dorsomesial view, right (bottom) lateral view. Scale: A, 1 mm; B, 5 mm." httpUri="https://zenodo.org/record/242206/files/figure.png" pageId="23" pageNumber="174">Fig. 15</figureCitation>
)
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<subSubSection id="C33027E88614FFBB12E8D1189F8DFE06" pageId="23" pageNumber="174" type="reference_group">
<paragraph id="8B9574638614FFBB12E8D1189E4EFEBB" blockId="23.[151,1284,325,486]" box="[151,526,325,347]" pageId="23" pageNumber="174">
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<taxonomicName id="4C2A0FE08614FFBB12E8D1189E4AFEBB" ID-CoL="632fd023-bb48-4d7c-81c4-b86bf2d10e9c" authority="Fabricius, 1775: 411" authorityName="Fabricius" authorityPageNumber="411" authorityYear="1775" box="[151,522,325,347]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB12E8D1189D71FEBB" box="[151,305,325,347]" italics="true" pageId="23" pageNumber="174">Pagurus alatus</emphasis>
<treatmentCitation id="0A8B52728614FFBB1348D11B9E4AFEBB" author="Fabricius" box="[311,522,325,347]" page="411" pageId="23" pageNumber="174" year="1775">
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: 411
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.
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<paragraph id="8B9574638614FFBB12E8D1379F8DFE06" blockId="23.[151,1284,325,486]" pageId="23" pageNumber="174">
<treatmentCitationGroup id="AB3A534D8614FFBB12E8D1379F8DFE06" pageId="23" pageNumber="174">
<taxonomicName id="4C2A0FE08614FFBB12E8D1379EB5FE9E" ID-CoL="53e3866c-74c9-44f3-986e-5b1f9ef2b34e" authority="Milne-Edwards &amp; Bouvier, 1892: 217" authorityName="Milne-Edwards &amp; Bouvier" authorityPageNumber="217" authorityYear="1892" box="[151,757,360,382]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB12E8D1379D2DFE9E" box="[151,365,360,382]" italics="true" pageId="23" pageNumber="174">Eupagurus variabilis</emphasis>
<treatmentCitation id="0A8B52728614FFBB130BD1369EB5FE9E" author="Milne-Edwards" box="[372,757,360,382]" page="217" pageId="23" pageNumber="174" year="1892">
<bibRefCitation id="EFBB09928614FFBB130BD1369E85FE9E" author="Milne-Edwards" box="[372,709,360,382]" pageId="23" pageNumber="174" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1892) Observations preliminaires sur les paguriens recueillis par les expeditions du Travailleur et du Talisman. Annales des Sciences Naturelles, Serie 7 e, 13, 185 - 226." type="journal article" year="1892">Milne-Edwards &amp; Bouvier, 1892</bibRefCitation>
: 217
</treatmentCitation>
</taxonomicName>
.— 
<treatmentCitation id="0A8B52728614FFBB116BD1369FFCFE9E" author="Selbie" box="[788,956,360,382]" page="36" pageId="23" pageNumber="174" year="1921">
<bibRefCitation id="EFBB09928614FFBB116BD1369FD6FE9E" author="Selbie" box="[788,918,360,382]" pageId="23" pageNumber="174" refString="Selbie, C. M. (1921) The Decapoda Reptantia of the coasts of Ireland. Part II: Paguridea. Fisheries Ireland Scientific Investigations, 1, 1 - 68." type="journal article" year="1921">Selbie, 1921</bibRefCitation>
: 36
</treatmentCitation>
, pl. IV figs. 4, 5, pl. V figs. 1–3. 
<emphasis id="B95EA8718614FFBB12E8D1D59D78FE40" box="[151,312,394,416]" italics="true" pageId="23" pageNumber="174">
<taxonomicName id="4C2A0FE08614FFBB12E8D1D59D74FE40" ID-CoL="632fd023-bb48-4d7c-81c4-b86bf2d10e9c" box="[151,308,394,416]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">Pagurus alatus</taxonomicName>
.
</emphasis>
— 
<treatmentCitation id="0A8B52728614FFBB132DD1D49DB7FE40" author="Ingle" box="[338,503,394,416]" page="765" pageId="23" pageNumber="174" year="1985">
<bibRefCitation id="EFBB09928614FFBB132DD1D49D88FE40" author="Ingle" box="[338,456,394,416]" pageId="23" pageNumber="174" refString="Ingle, R. W. (1985) Northeastern Atlantic and Mediterranean hermit crabs (Crustacea: Anomura: Paguroidea: Paguridae). I. The genus Pagurus Fabricius, 1775. Journal of Natural History, 19, 745 - 769. http: // dx. doi. org / 10.1080 / 00222938500770461" type="journal article" year="1985">Ingle, 1985</bibRefCitation>
: 765
</treatmentCitation>
(key); 1993: 33 (key) 136, figs. 109–112 (lit.). not 
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<taxonomicName id="4C2A0FE08614FFBB12C2D1F09D34FE23" box="[189,372,429,451]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">Eupagurus alatus</taxonomicName>
.
</emphasis>
— 
<treatmentCitation id="0A8B52728614FFBB13EED1F09E03FE23" author="Forest" box="[401,579,429,451]" page="110" pageId="23" pageNumber="174" year="1955">
<bibRefCitation id="EFBB09928614FFBB13EED1F09E52FE23" author="Forest" box="[401,530,429,451]" pageId="23" pageNumber="174" refString="Forest, J. (1955) Crustaces Decapodes, Pagurides. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de L'Atlantique Sud (1948 - 1949). Resultats Scientifiques. Institut Royal des Sciences Naturelles de Belge, 3 (4), 23 - 147." type="journal article" year="1955">Forest, 1955</bibRefCitation>
: 110
</treatmentCitation>
, fig. 23, pl. V figs. 2 and 3 (= 
<taxonomicName id="4C2A0FE08614FFBB1109D1F09FB3FE23" ID-CoL="288bc55e-353b-4bfd-a3ad-16ea7fc33adf" box="[886,1011,430,451]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB1109D1F09FB3FE23" box="[886,1011,430,451]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
) not 
<taxonomicName id="4C2A0FE08614FFBB12C2D18F9D18FE06" ID-CoL="632fd023-bb48-4d7c-81c4-b86bf2d10e9c" box="[189,344,464,486]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB12C2D18F9D18FE06" box="[189,344,464,486]" italics="true" pageId="23" pageNumber="174">Pagurus alatus</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0A8B52728614FFBB130AD18E9E0CFE06" author="Zariquiey" box="[373,588,464,486]" page="247" pageId="23" pageNumber="174" year="1968">
<bibRefCitation id="EFBB09928614FFBB130AD18E9E5AFE06" author="Zariquiey" box="[373,538,464,486]" pageId="23" pageNumber="174" refString="Zariquiey Alvarez, R. (1968) Crustaceos Decapodos Ibericos. Investigacion Pesquera, 32, 1 - 510." type="journal article" year="1968">Zariquiey, 1968</bibRefCitation>
: 247
</treatmentCitation>
, figs 89e, 90b, o, 91e (= 
<taxonomicName id="4C2A0FE08614FFBB1137D18F9F85FE06" ID-CoL="288bc55e-353b-4bfd-a3ad-16ea7fc33adf" box="[840,965,465,486]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB1137D18F9F85FE06" box="[840,965,465,486]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
)
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</paragraph>
</subSubSection>
<subSubSection id="C33027E88614FFBB12E8D24B9D3FFD3B" pageId="23" pageNumber="174" type="materials_examined">
<paragraph id="8B9574638614FFBB12E8D24B9962FDB0" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
<emphasis id="B95EA8718614FFBB12E8D24B9D36FDCE" bold="true" box="[151,374,533,558]" pageId="23" pageNumber="174">Material examined</emphasis>
. MU57, 430– 
<quantity id="4CD2D9868614FFBB106DD24B9E19FDCD" box="[530,601,533,558]" metricMagnitude="2" metricUnit="m" metricValue="4.06" pageId="23" pageNumber="174" unit="m" value="406.0">406 m</quantity>
, (1); MU120, 109– 
<quantity id="4CD2D9868614FFBB114ED24B9F38FDCD" box="[817,888,533,558]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="23" pageNumber="174" unit="m" value="105.0">105 m</quantity>
, (1); MU142, 109– 
<quantity id="4CD2D9868614FFBB1631D24898D5FDCD" box="[1102,1173,534,558]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" pageId="23" pageNumber="174" unit="m" value="112.0">112 m</quantity>
, (1); MU158, 80– 
<quantity id="4CD2D9868614FFBB1721D24899D8FDCD" box="[1374,1432,534,558]" metricMagnitude="1" metricUnit="m" metricValue="9.8" pageId="23" pageNumber="174" unit="m" value="98.0">98 m</quantity>
, (2); MU207, 88– 
<quantity id="4CD2D9868614FFBB132ED2669DD8FDAF" box="[337,408,568,592]" metricMagnitude="2" metricUnit="m" metricValue="1.17" pageId="23" pageNumber="174" unit="m" value="117.0">117 m</quantity>
, (1); MU260, 101– 
<quantity id="4CD2D9868614FFBB1015D2669EF3FDAF" box="[618,691,568,592]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="174" unit="m" value="120.0">120 m</quantity>
, (1); 
<collectionCode id="ED3BECA68614FFBB1093D2669F01FDB0" box="[748,833,568,592]" pageId="23" pageNumber="174">MUBV</collectionCode>
<quantity id="4CD2D9868614FFBB113ED2669FEFFDAF" box="[833,943,567,592]" metricMagnitude="4" metricUnit="m" metricValue="2.0155" pageId="23" pageNumber="174" unit="m" value="20155.0">20, 155 m</quantity>
, (5); 
<collectionCode id="ED3BECA68614FFBB1197D266987FFDB0" box="[1000,1087,568,592]" pageId="23" pageNumber="174">MUBV</collectionCode>
21, 107– 
<quantity id="4CD2D9868614FFBB16E4D26698A3FDAF" box="[1179,1251,568,592]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="23" pageNumber="174" unit="m" value="109.0">109 m</quantity>
, (31).
</paragraph>
<paragraph id="8B9574638614FFBB12B8D204980AFD92" blockId="23.[151,1437,533,2014]" box="[199,1098,602,627]" pageId="23" pageNumber="174">
Males: 
<quantity id="4CD2D9868614FFBB1367D2049DFAFD92" box="[280,442,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" metricValueMax="6.56" metricValueMin="3.06" pageId="23" pageNumber="174" unit="mm" value="4.81" valueMax="6.56" valueMin="3.06">3.06–6.56 mm</quantity>
, females: 
<quantity id="4CD2D9868614FFBB1057D2059E8AFD92" box="[552,714,603,627]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="23" pageNumber="174" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.10–3.40 mm</quantity>
, ovigerous females: 
<quantity id="4CD2D9868614FFBB11D4D205980AFD92" box="[939,1098,603,627]" metricMagnitude="-3" metricUnit="m" metricValue="3.52" metricValueMax="4.03" metricValueMin="3.01" pageId="23" pageNumber="174" unit="mm" value="3.52" valueMax="4.03" valueMin="3.01">3.01–4.03 mm</quantity>
</paragraph>
<paragraph id="8B9574638614FFBB12B8D2239D3FFD3B" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
<emphasis id="B95EA8718614FFBB12B8D2239D61FD76" bold="true" box="[199,289,637,662]" pageId="23" pageNumber="174">Habitat</emphasis>
. This species is characteristic of bathyal muds (
<bibRefCitation id="EFBB09928614FFBB1151D223987DFD76" author="d'Udekem" box="[814,1085,637,662]" pageId="23" pageNumber="174" refString="d'Udekem d'Acoz, C. (1999) Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Paris, Museum national d'histoire naturelle, Service du patrimoine naturel, 40, 1 - 383." type="journal article" year="1999">d’Udekem d’Acoz 1999</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1636D2239957FD76" author="Follesa" box="[1097,1303,637,662]" pageId="23" pageNumber="174" refString="Follesa, M. C., Porcu, C., Gastoni, A., Mulas, A., Sabatini, A. &amp; Cau, A. (2009) Community structure of bathyal decapod crustaceans off South-Eastern Sardinian deep-waters (Central-Western Mediterranean). Marine Ecology, 30 (s 1), 188 - 199. http: // dx. doi. org / 10.1111 / j. 1439 - 0485.2009.00323. x" type="journal article" year="2009">
Follesa 
<emphasis id="B95EA8718614FFBB16E0D220988FFD76" box="[1183,1231,637,662]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB175DD2209C92FD58" author="Mura" pageId="23" pageNumber="174" refString="Mura, M., Orru, F. &amp; Cau, A. (2006) Reproduction Strategy of the Deep-sea Hermit Crabs Pagurus alatus and Pagurus excavatus of the Central-Western Mediterranean Sea. Hydrobiologia, 557 (1), 51 - 57. http: // dx. doi. org / 10.1007 / s 10750 - 005 - 1307 - x" type="journal article" year="2006">
Mura 
<emphasis id="B95EA8718614FFBB171AD22099D5FD76" box="[1381,1429,637,662]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2006
</bibRefCitation>
), but is also reported on sand (
<bibRefCitation id="EFBB09928614FFBB1064D2C19F71FD58" author="Pipitone" box="[539,817,671,696]" pageId="23" pageNumber="174" refString="Pipitone, C. &amp; Arculeo, M. (2003) The marine Crustacea Decapoda of Sicily (central Mediterranean Sea): a checklist with remarks on their distribution. Italian Journal of Zoology, 70, 69 - 78. http: // dx. doi. org / 10.1080 / 11250000309356498" type="journal article" year="2003">Pipitone &amp; Arculeo 2003</bibRefCitation>
) and on bottoms of sand, silt, coarse sands and gravel by 
<bibRefCitation id="EFBB09928614FFBB12E8D29D9D3BFD3B" box="[151,379,706,731]" pageId="23" pageNumber="174" refString="Serrano, A., Sanchez, F., Punzon, A., Velasco, F. &amp; Olaso, I. (2011) Deep sea megafaunal assemblages off the northern Iberian slope related to environmental factors. Scientia Marina, 75 (3), 425 - 437. http: // dx. doi. org / 10.3989 / scimar. 2011.75 n 3425" type="journal article">
Serrano 
<emphasis id="B95EA8718614FFBB128CD29A9D62FD3B" box="[243,290,706,731]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. (2011)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33027E88614FFBB12B8D2B89F08FB9B" pageId="23" pageNumber="174" type="reference_group">
<paragraph id="8B9574638614FFBB12B8D2B89C9EFC68" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
<taxonomicName id="4C2A0FE08614FFBB12B8D2B89D2DFD1E" ID-CoL="632fd023-bb48-4d7c-81c4-b86bf2d10e9c" box="[199,365,741,766]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB12B8D2B89D2DFD1E" box="[199,365,741,766]" italics="true" pageId="23" pageNumber="174">Pagurus alatus</emphasis>
</taxonomicName>
was found inhabiting shells of the gastropod species 
<taxonomicName id="4C2A0FE08614FFBB11CFD2BB98BDFD1E" authority="Mohr, 1786" authorityName="Mohr" authorityYear="1786" box="[944,1277,741,766]" class="Gastropoda" family="Buccinidae" genus="Colus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="islandicus">
<emphasis id="B95EA8718614FFBB11CFD2BB9825FD1E" box="[944,1125,741,766]" italics="true" pageId="23" pageNumber="174">Colus islandicus</emphasis>
(Mohr, 1786)
</taxonomicName>
in Cap Breton Canyon (Bay of Biscay) (
<bibRefCitation id="EFBB09928614FFBB13C0D3599EDCFCC0" author="Urzelai" box="[447,668,775,800]" pageId="23" pageNumber="174" refString="Urzelai, A., Elizalde, M., Capellan, T., Esteban, I., Quiroga, A., Zabala, I. &amp; Ibanez, M. (1990) Estudio preliminar de las comunidades de Pagurus alatus Fabricius, 1775 y Parapagurus pilosimanus S. l. Smith, 1879 (Crustacea Decapoda) y Epizoanthus paguriphilus Verrill, 1883 (Anthozoa Zoantarida) de la fosa de Cap Breton (Golfo de Vizcaya). Lurralde, 13, 193 - 206." type="journal article" year="1990">
Urzelai 
<emphasis id="B95EA8718614FFBB1063D3579E0FFCC0" box="[540,591,775,800]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 1990
</bibRefCitation>
). In symbiosis with this crab were the cnidarians 
<taxonomicName id="4C2A0FE08614FFBB16A3D3599D04FCA3" ID-CoL="7QCFK" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" class="Anthozoa" family="Hormathiidae" genus="Adamsia" kingdom="Animalia" order="Actiniaria" pageId="23" pageNumber="174" phylum="Cnidaria" rank="species" species="palliata">
<emphasis id="B95EA8718614FFBB16A3D35999DCFCC0" box="[1244,1436,775,800]" italics="true" pageId="23" pageNumber="174">Adamsia palliata</emphasis>
(Müller, 1776)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB1325D3749F42FCA3" ID-CoL="PLQB" authority="Couch, 1842" authorityName="Couch" authorityYear="1842" box="[346,770,810,835]" class="Anthozoa" family="Hormathiidae" genus="Calliactis" kingdom="Animalia" order="Actiniaria" pageId="23" pageNumber="174" phylum="Cnidaria" rank="species" species="parasitica">
<emphasis id="B95EA8718614FFBB1325D3749E05FCA3" box="[346,581,810,835]" italics="true" pageId="23" pageNumber="174">Calliactis parasitica</emphasis>
(Couch, 1842)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB1167D37498ABFCA3" authority="Verrill, 1883" authorityName="Verrill" authorityYear="1883" box="[792,1259,810,835]" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="23" pageNumber="174" phylum="Cnidaria" rank="species" species="paguriphylus">
<emphasis id="B95EA8718614FFBB1167D3749801FCA3" box="[792,1089,810,835]" italics="true" pageId="23" pageNumber="174">Epizoanthus paguriphylus</emphasis>
Verrill, 1883
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB177ED3749E04FC86" ID-CoL="4CNBJ" authority="Ross and Zamponi, 1982" authorityName="Ross and Zamponi" authorityYear="1982" class="Anthozoa" family="Hormathiidae" genus="Paracalliactis" kingdom="Animalia" order="Actiniaria" pageId="23" pageNumber="174" phylum="Cnidaria" rank="species" species="mediterranea">
<emphasis id="B95EA8718614FFBB177ED3749D6BFC86" italics="true" pageId="23" pageNumber="174">Paracalliactis mediterranea</emphasis>
Ross and Zamponi, 1982
</taxonomicName>
and the polychaete 
<taxonomicName id="4C2A0FE08614FFBB1161D3139C9AFC68" ID-CoL="45Y4D" authority="(Savigny, 1818) (Williams &amp; McDermott 2004)" authorityName="(Savigny, 1818) (Williams &amp; McDermott" authorityYear="2004" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="23" pageNumber="174" phylum="Annelida" rank="species" species="fucata">
<emphasis id="B95EA8718614FFBB1161D3139F8FFC86" box="[798,975,845,870]" italics="true" pageId="23" pageNumber="174">Neanthes fucata</emphasis>
(Savigny, 1818) (
<bibRefCitation id="EFBB09928614FFBB16ECD3139C92FC68" author="Williams" pageId="23" pageNumber="174" refString="Williams, J. D. &amp; McDermott, J. J. (2004) Hermit crab biocoenoses: A worldwide review of the diversity and natural history of hermit crab associates. Journal of experimental marine biology and ecology, 305, 1 - 128. http: // dx. doi. org / 10.1016 / j. jembe. 2004.02.020" type="journal article" year="2004">Williams &amp; McDermott 2004</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B9574638614FFBB12B8D3CD9F08FB9B" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
Our specimens were collected mainly on sandy bottoms, but also on sandy-mud bottoms. They were found inhabiting shells of the gastropods 
<taxonomicName id="4C2A0FE08614FFBB1062D3E89FC2FC2E" ID-CoL="5VTXQ" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[541,898,949,974]" class="Gastropoda" family="Bursidae" genus="Aspa" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="marginata">
<emphasis id="B95EA8718614FFBB1062D3E89E8DFC2D" box="[541,717,950,973]" italics="true" pageId="23" pageNumber="174">Aspa marginata</emphasis>
(Gmelin, 1791)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB11EFD3EB9978FC2E" ID-CoL="6987L" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[912,1336,949,974]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="granulatum">
<emphasis id="B95EA8718614FFBB11EFD3EB98DEFC2E" box="[912,1182,949,974]" italics="true" pageId="23" pageNumber="174">Calliostoma granulatum</emphasis>
(Born, 1778)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB173AD3E89E76FC10" ID-CoL="6HGNM" authority="Marche-Marchad, 1957" authorityName="Marche-Marchad" authorityYear="1957" class="Gastropoda" family="Naticidae" genus="Euspira" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="grossularia">
<emphasis id="B95EA8718614FFBB173AD3E89D54FC10" italics="true" pageId="23" pageNumber="174">Euspira grossularia</emphasis>
(Marche-Marchad, 1957)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB103CD3879F8EFC10" ID-CoL="6HGQ8" authority="Jeffreys, 1885" authorityName="Jeffreys" authorityYear="1885" box="[579,974,983,1008]" class="Gastropoda" family="Naticidae" genus="Euspira" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="subplicata">
<emphasis id="B95EA8718614FFBB103CD3879F55FC10" box="[579,789,983,1008]" italics="true" pageId="23" pageNumber="174">Euspira subplicata</emphasis>
(Jeffreys, 1885)
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB11A4D3879C91FBF3" ID-CoL="45RK8" authority="Rolan &amp; Hernandez, 2005" authorityName="Rolan &amp; Hernandez" authorityYear="2005" class="Gastropoda" family="Nassariidae" genus="Nassarius" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="arcadioi">
<emphasis id="B95EA8718614FFBB11A4D38798EDFC10" box="[987,1197,983,1008]" italics="true" pageId="23" pageNumber="174">Nassarius arcadioi</emphasis>
Rolán &amp; Hernández, 2005
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB12A2D3A29E0FFBF3" ID-CoL="45TJ2" authority="Odhner, 1932" authorityName="Odhner" authorityYear="1932" box="[221,591,1018,1043]" class="Gastropoda" family="Naticidae" genus="Natica" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="canariensis">
<emphasis id="B95EA8718614FFBB12A2D3A29DEDFBF3" box="[221,429,1020,1043]" italics="true" pageId="23" pageNumber="174">Natica canariensis</emphasis>
Odhner, 1932
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB1023D3A49FA3FBF3" ID-CoL="4RF52" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[604,995,1018,1043]" class="Gastropoda" family="Ranellidae" genus="Ranella" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="olearium">
<emphasis id="B95EA8718614FFBB1023D3A49F5DFBF3" box="[604,797,1018,1043]" italics="true" pageId="23" pageNumber="174">Ranella olearium</emphasis>
(Linnaeus, 1758)
</taxonomicName>
and in a shell of the genus 
<taxonomicName id="4C2A0FE08614FFBB1762D3A5992EFBF2" ID-CoL="7P48M" box="[1309,1390,1019,1042]" class="Gastropoda" family="Borsoniidae" genus="Genota" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="174" phylum="Mollusca" rank="genus">
<emphasis id="B95EA8718614FFBB1762D3A5992EFBF2" box="[1309,1390,1019,1042]" italics="true" pageId="23" pageNumber="174">Genota</emphasis>
</taxonomicName>
. As epibionts over gastropod shells we found Demospongia, two different actinian species, some specimens of 
<taxonomicName id="4C2A0FE08614FFBB175DD44399DCFBD6" box="[1314,1436,1053,1078]" class="Bivalvia" family="Anomiidae" kingdom="Animalia" order="Pectinoida" pageId="23" pageNumber="174" phylum="Mollusca" rank="family">Anomiidae</taxonomicName>
bivalves, one acorn barnacle and some tubes of 
<taxonomicName id="4C2A0FE08614FFBB10CBD4619F69FBB8" ID-CoL="84KRW" box="[692,809,1087,1112]" class="Polychaeta" family="Serpulidae" kingdom="Animalia" order="Sabellida" pageId="23" pageNumber="174" phylum="Annelida" rank="family">Serpulidae</taxonomicName>
polychetes. Over the chelipeds and ambulatory legs we identified the hydrozoan 
<taxonomicName id="4C2A0FE08614FFBB13D7D43C9F04FB9B" ID-CoL="723RK" authority="Fleming, 1823" authorityName="Fleming" authorityYear="1823" box="[424,836,1122,1147]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="23" pageNumber="174" phylum="Cnidaria" rank="species" species="octona">
<emphasis id="B95EA8718614FFBB13D7D43C9EC9FB9B" box="[424,649,1122,1147]" italics="true" pageId="23" pageNumber="174">Leuckartiara octona</emphasis>
(Fleming, 1823)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33027E88614FFBB12B8D4DB9C9EFA36" pageId="23" pageNumber="174" type="distribution">
<paragraph id="8B9574638614FFBB12B8D4DB9DBDFAC8" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
<emphasis id="B95EA8718614FFBB12B8D4DB9D16FB7E" bold="true" box="[199,342,1157,1182]" pageId="23" pageNumber="174">Distribution</emphasis>
. Eastern Atlantic, from 
<collectingCountry id="F33D34F38614FFBB1012D4DB9E80FB7E" box="[621,704,1157,1182]" name="Iceland" pageId="23" pageNumber="174">Iceland</collectingCountry>
, 
<collectingCountry id="F33D34F38614FFBB10B1D4D89F67FB7E" box="[718,807,1158,1182]" name="Norway" pageId="23" pageNumber="174">Norway</collectingCountry>
, the 
<collectingCountry id="F33D34F38614FFBB111DD4DB985AFB7E" box="[866,1050,1157,1182]" name="United Kingdom" pageId="23" pageNumber="174">Shetland Islands</collectingCountry>
and the North Sea southwards to 
<collectingCountry id="F33D34F38614FFBB12E8D4F99D52FB20" box="[151,274,1191,1216]" name="Mauritania" pageId="23" pageNumber="174">Mauritania</collectingCountry>
, including the Canary Islands and the Mediterranean Sea (
<bibRefCitation id="EFBB09928614FFBB11F3D4F998DAFB20" author="d'Udekem" box="[908,1178,1191,1216]" pageId="23" pageNumber="174" refString="d'Udekem d'Acoz, C. (1999) Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Paris, Museum national d'histoire naturelle, Service du patrimoine naturel, 40, 1 - 383." type="journal article" year="1999">d’Udekem d’Acoz 1999</bibRefCitation>
). The record from Loos Islands (
<collectingCountry id="F33D34F38614FFBB1286D4949D0BFB03" box="[249,331,1226,1251]" name="Guinea-Bissau" pageId="23" pageNumber="174">Guinea</collectingCountry>
) (
<bibRefCitation id="EFBB09928614FFBB1318D4959DB3FB03" author="Forest" box="[359,499,1226,1251]" pageId="23" pageNumber="174" refString="Forest, J. (1955) Crustaces Decapodes, Pagurides. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de L'Atlantique Sud (1948 - 1949). Resultats Scientifiques. Institut Royal des Sciences Naturelles de Belge, 3 (4), 23 - 147." type="journal article" year="1955">Forest 1955</bibRefCitation>
) belong to 
<taxonomicName id="4C2A0FE08614FFBB10FCD4929F49FB03" box="[643,777,1228,1251]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB10FCD4929F49FB03" box="[643,777,1228,1251]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
(see remarks). Bathymetrical range varies from 
<quantity id="4CD2D9868614FFBB1736D49499DBFB02" box="[1353,1435,1226,1251]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="23" pageNumber="174" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">5–10 m</quantity>
(
<bibRefCitation id="EFBB09928614FFBB12E1D4B39DF7FAE6" author="Pipitone" box="[158,439,1261,1286]" pageId="23" pageNumber="174" refString="Pipitone, C. &amp; Arculeo, M. (2003) The marine Crustacea Decapoda of Sicily (central Mediterranean Sea): a checklist with remarks on their distribution. Italian Journal of Zoology, 70, 69 - 78. http: // dx. doi. org / 10.1080 / 11250000309356498" type="journal article" year="2003">Pipitone &amp; Arculeo 2003</bibRefCitation>
) to 
<quantity id="4CD2D9868614FFBB139CD4B39E3BFAE5" box="[483,635,1261,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.4675" metricValueMax="1.505" metricValueMin="1.43" pageId="23" pageNumber="174" unit="m" value="1467.5" valueMax="1505.0" valueMin="1430.0">1430–1505 m</quantity>
(
<bibRefCitation id="EFBB09928614FFBB10F5D4B09F14FAE6" author="Garcia" box="[650,852,1261,1286]" pageId="23" pageNumber="174" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero-Moroccan waters. Results of Balgim- 84 Expedition. Bulletin of Marine Science, 58 (3), 730 - 752." type="journal article" year="1996">García Raso 1996</bibRefCitation>
), although the species is mainly found around 
<quantity id="4CD2D9868614FFBB1729D4B399DBFAE5" box="[1366,1435,1261,1286]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="174" unit="m" value="150.0">150 m</quantity>
depth (
<bibRefCitation id="EFBB09928614FFBB129DD5519DB0FAC8" author="d'Udekem" box="[226,496,1295,1320]" pageId="23" pageNumber="174" refString="d'Udekem d'Acoz, C. (1999) Inventaire et distribution des crustaces decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Paris, Museum national d'histoire naturelle, Service du patrimoine naturel, 40, 1 - 383." type="journal article" year="1999">d’Udekem d’Acoz 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B9574638614FFBB12E8D56C9C9EFA36" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
Further records were from the Bay of Biscay (
<bibRefCitation id="EFBB09928614FFBB10EAD56C9F27FAAB" author="Urzelai" box="[661,871,1330,1355]" pageId="23" pageNumber="174" refString="Urzelai, A., Elizalde, M., Capellan, T., Esteban, I., Quiroga, A., Zabala, I. &amp; Ibanez, M. (1990) Estudio preliminar de las comunidades de Pagurus alatus Fabricius, 1775 y Parapagurus pilosimanus S. l. Smith, 1879 (Crustacea Decapoda) y Epizoanthus paguriphilus Verrill, 1883 (Anthozoa Zoantarida) de la fosa de Cap Breton (Golfo de Vizcaya). Lurralde, 13, 193 - 206." type="journal article" year="1990">
Urzelai 
<emphasis id="B95EA8718614FFBB1091D56A9F5EFAAB" box="[750,798,1330,1355]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 1990
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB110CD56D9807FAAB" author="Serrano" box="[883,1095,1330,1355]" pageId="23" pageNumber="174" refString="Serrano, A., Sanchez, F. &amp; Garcia-Castrillo, G. (2006) Epibenthic communities of trawlable grounds of the Cantabrian Sea. Scientia Marina, 70 S 1, 149 - 159." type="journal article" year="2006">
Serrano 
<emphasis id="B95EA8718614FFBB11B0D56A9FBFFAAB" box="[975,1023,1330,1355]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2006
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB162CD56D995AFAAB" author="Cartes" box="[1107,1306,1330,1355]" pageId="23" pageNumber="174" refString="Cartes, J. E., Serrano, A., Velasco, F., Parra, S. &amp; Sanchez, F. (2007) Community structure and dynamics of deep-water decapod assemblages from Le Danois Bank (Cantabrian Sea, NE Atlantic): Influence of environmental variables and food availability. Progress in Oceanography, 75, 797 - 816. http: // dx. doi. org / 10.1016 / j. pocean. 2007.09.003" type="journal article" year="2007">
Cartes 
<emphasis id="B95EA8718614FFBB16DED56A9891FAAB" box="[1185,1233,1330,1355]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2007
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB175AD56C9CB6FA8E" author="Sanchez" pageId="23" pageNumber="174" refString="Sanchez, F., Serrano, A., Parra, A., Ballesteros, M. &amp; Cartes, J. E. (2008) Habitat characteristics as determinant of the structure and spatial distribution of epibenthic and demersal communities of Le Danois Bank (Cantabrian Sea, N Spain). Journal of Marine Systems, 72, 64 - 86. http: // dx. doi. org / 10.1016 / j. jmarsys. 2007.04.008" type="journal article" year="2008">
Sánchez 
<emphasis id="B95EA8718614FFBB17F7D56A9CECFA8E" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2008
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB137BD5089D9FFA8E" author="Serrano" box="[260,479,1365,1390]" pageId="23" pageNumber="174" refString="Serrano, A., Sanchez, F., Punzon, A., Velasco, F. &amp; Olaso, I. (2011) Deep sea megafaunal assemblages off the northern Iberian slope related to environmental factors. Scientia Marina, 75 (3), 425 - 437. http: // dx. doi. org / 10.3989 / scimar. 2011.75 n 3425" type="journal article" year="2011">
Serrano 
<emphasis id="B95EA8718614FFBB131CD5089DD5FA8E" box="[355,405,1365,1390]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2011
</bibRefCitation>
), off 
<collectingCountry id="F33D34F38614FFBB1061D50B9E3BFA8E" box="[542,635,1365,1390]" name="Portugal" pageId="23" pageNumber="174">Portugal</collectingCountry>
(
<bibRefCitation id="EFBB09928614FFBB10F4D50B9F38FA8E" author="Monteiro" box="[651,888,1365,1390]" pageId="23" pageNumber="174" refString="Monteiro, P., Araujo, A., Erzini, K. &amp; Castro, M. (2001) Discards of the Algarve (southern Portugal) crustacean trawl fishery. Hydrobiologia, 449, 267 - 277. http: // dx. doi. org / 10.1023 / A: 1017575429808" type="journal article" year="2001">
Monteiro 
<emphasis id="B95EA8718614FFBB1082D5089F70FA8E" box="[765,816,1365,1390]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2001
</bibRefCitation>
) and the Mediterranean Sea (
<bibRefCitation id="EFBB09928614FFBB16B7D50B99D7FA8E" author="Kocak" box="[1224,1431,1365,1390]" pageId="23" pageNumber="174" refString="Kocak, C., Katagan, T. &amp; Kocatas, A. (2001) Anomurans of the Aegean coasts of Turkey and reported species from Turkish seas. Turkish Journal of Zoology, 25, 305 - 311." type="journal article" year="2001">
Koçak 
<emphasis id="B95EA8718614FFBB1764D508990DFA8E" box="[1307,1357,1365,1390]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2001
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB12E8D5299D21FA70" author="Abello" box="[151,353,1399,1424]" pageId="23" pageNumber="174" refString="Abello, P., Carbonell, A. &amp; Torres, P. (2002) Biogeography of epibenthic crustaceans on the shelf and upper slope off the Iberian Peninsula Mediterranean coasts: implications for the establishment of natural management areas. Scientia Marina, 66 (2), 183 - 198." type="journal article" year="2002">
Abelló 
<emphasis id="B95EA8718614FFBB1296D5279D59FA70" box="[233,281,1399,1424]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1312D5299E67FA70" author="Biagi" box="[365,551,1399,1424]" pageId="23" pageNumber="174" refString="Biagi, F., Sartor, P., Ardizzone, G. D., Belcari, P., Belluscio, A. &amp; Serena, F. (2002) Analysis of demersal assemblages off the Tuscany and Latium coasts (north-western Mediterranean). Scientia Marina, 66 (2), 233 - 242." type="journal article" year="2002">
Biagi 
<emphasis id="B95EA8718614FFBB13D0D5279D9FFA70" box="[431,479,1399,1424]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2002
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; 
<bibRefCitation id="EFBB09928614FFBB104CD5299F0CFA70" author="Pipitone" box="[563,844,1399,1424]" pageId="23" pageNumber="174" refString="Pipitone, C. &amp; Arculeo, M. (2003) The marine Crustacea Decapoda of Sicily (central Mediterranean Sea): a checklist with remarks on their distribution. Italian Journal of Zoology, 70, 69 - 78. http: // dx. doi. org / 10.1080 / 11250000309356498" type="journal article" year="2003">Pipitone &amp; Arculeo 2003</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1127D529986DFA70" author="Colloca" box="[856,1069,1399,1424]" pageId="23" pageNumber="174" refString="Colloca, F., Carpentieri, P., Balestri, E. &amp; Ardizzone, G. D. (2004) A critical habitat for Mediterranean fish resources: shelfbreak areas with Leptometra phalangium (Echinodermata: Crinoidea). Marine Biology, 145 (6), 1129 - 1142. http: // dx. doi. org / 10.1007 / s 00227 - 004 - 1405 - 8" type="journal article" year="2004">
Colloca 
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; 
<bibRefCitation id="EFBB09928614FFBB1647D5269963FA70" author="Company" box="[1080,1315,1399,1424]" pageId="23" pageNumber="174" refString="Company, J. B., Maiorano, P., Tselepides, A., Politou, C. - Y., Plaity, W., Rotllant, G. &amp; Sarda, F. (2004). Deep-sea decapod crustaceans in the western and central Mediterranean Sea: preliminary aspects of species distribution, biomass and population structure. Scientia Marina, 68 (3), 73 - 86." type="journal article" year="2004">
Company 
<emphasis id="B95EA8718614FFBB16D4D527989AFA70" box="[1195,1242,1399,1424]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1750D5269CB5FA53" author="Ungaro" pageId="23" pageNumber="174" refString="Ungaro, N., Marano, C. A., Ceriola, L. &amp; Martino, M. (2005) Distribution of demersal crustaceans in the southern Adriatic Sea. Acta Adriatica, 46 (1), 27 - 40." type="journal article" year="2005">
Ungaro 
<emphasis id="B95EA8718614FFBB17F7D5279CECFA53" italics="true" pageId="23" pageNumber="174">et al</emphasis>
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</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB137ED5C59DFEFA53" author="Mura" box="[257,446,1434,1459]" pageId="23" pageNumber="174" refString="Mura, M., Orru, F. &amp; Cau, A. (2006) Reproduction Strategy of the Deep-sea Hermit Crabs Pagurus alatus and Pagurus excavatus of the Central-Western Mediterranean Sea. Hydrobiologia, 557 (1), 51 - 57. http: // dx. doi. org / 10.1007 / s 10750 - 005 - 1307 - x" type="journal article" year="2006">
Mura 
<emphasis id="B95EA8718614FFBB133AD5C29D35FA53" box="[325,373,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
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</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB13B4D5C59E3FFA53" author="Ates" box="[459,639,1434,1459]" pageId="23" pageNumber="174" refString="Ates, A. S., Katagan, T. &amp; Kocatas, A. (2006) Bathymetric distribution of decapod crustaceans on the continental shelf along the Aegean coasts of Turkey. Crustaceana, 79 (2), 129 - 141. http: // dx. doi. org / 10.1163 / 156854006776952928" type="journal article" year="2006">
Ates 
<emphasis id="B95EA8718614FFBB107AD5C29E76FA53" box="[517,566,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2006
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB10F4D5C49F19FA53" author="Fanelli" box="[651,857,1434,1459]" pageId="23" pageNumber="174" refString="Fanelli, E. F., Colloca, F. &amp; Ardizzone, G. D. (2007) Decapod crustacean assemblages off the West coast of central Italy (western Mediterranean). Scientia Marina, 71 (1), 19 - 28." type="journal article" year="2007">
Fanelli 
<emphasis id="B95EA8718614FFBB10A0D5C29F50FA53" box="[735,784,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2007
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB111AD5C5986DFA53" author="Cartes" box="[869,1069,1434,1459]" pageId="23" pageNumber="174" refString="Cartes, J. E., Maynou, F., Fanelli, E., Papiol, V. &amp; Lloris, D. (2009) Long-term changes in the composition and diversity of deep-slope megabenthos and trophic webs off Catalonia (western Mediterranean): are trends related to climatic oscillations? Progress in Oceanography, 82, 32 - 46. http: // dx. doi. org / 10.1016 / j. pocean. 2009.03.003" type="journal article" year="2009">
Cartes 
<emphasis id="B95EA8718614FFBB11CCD5C29FA4FA53" box="[947,996,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1646D5C4994AFA53" author="Follesa" box="[1081,1290,1434,1459]" pageId="23" pageNumber="174" refString="Follesa, M. C., Porcu, C., Gastoni, A., Mulas, A., Sabatini, A. &amp; Cau, A. (2009) Community structure of bathyal decapod crustaceans off South-Eastern Sardinian deep-waters (Central-Western Mediterranean). Marine Ecology, 30 (s 1), 188 - 199. http: // dx. doi. org / 10.1111 / j. 1439 - 0485.2009.00323. x" type="journal article" year="2009">
Follesa 
<emphasis id="B95EA8718614FFBB16EFD5C29881FA53" box="[1168,1217,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFBB09928614FFBB1769D5C49C92FA36" author="Papiol" pageId="23" pageNumber="174" refString="Papiol, V., Cartes, J. E., Fanelli, E. &amp; Maynou, F. (2012) Influence of environmental variables on the spatio-temporal dynamics of bentho-pelagic assemblages in the middle slope of the Baleric Basin (NW Mediterranean). Deep-Sea Research I, 61, 84 - 99. http: // dx. doi. org / 10.1016 / j. dsr. 2011.11.008" type="journal article" year="2012">
Papiol 
<emphasis id="B95EA8718614FFBB171BD5C299D5FA53" box="[1380,1429,1434,1459]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33027E88614FFB412B8D5819F4BFEDA" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="discussion">
<paragraph id="8B9574638614FFBB12B8D58198ECFA18" blockId="23.[151,1437,533,2014]" box="[199,1196,1503,1528]" pageId="23" pageNumber="174">
<emphasis id="B95EA8718614FFBB12B8D5819D72FA18" bold="true" box="[199,306,1503,1528]" pageId="23" pageNumber="174">Remarks</emphasis>
. Our specimens coincide with the descriptions and figures given by 
<bibRefCitation id="EFBB09928614FFBB1661D58198E8FA18" author="Ingle" box="[1054,1192,1503,1528]" pageId="23" pageNumber="174" refString="Ingle, R. W. (1993) Hermit crabs of the Northeastern Atlantic Ocean and the Mediterranean Sea: an illustrated key. Chapman &amp; Hall, London, 495 pp." type="book" year="1993">Ingle (1993)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9574638614FFBB12E8D65C9D00F927" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
The taxonomic status of 
<taxonomicName id="4C2A0FE08614FFBB13BAD65A9E62F9FB" box="[453,546,1538,1563]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB13BAD65A9E62F9FB" box="[453,546,1538,1563]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2A0FE08614FFBB1047D65A9EFEF9FB" box="[568,702,1540,1563]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB1047D65A9EFEF9FB" box="[568,702,1540,1563]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0FE08614FFBB117CD65A9FC7F9FB" box="[771,903,1538,1563]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB117CD65A9FC7F9FB" box="[771,903,1538,1563]" italics="true" pageId="23" pageNumber="174">P. variabilis</emphasis>
</taxonomicName>
has been controversial for many years. The differences between 
<taxonomicName id="4C2A0FE08614FFBB13F6D6789E4FF9DD" box="[393,527,1574,1597]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB13F6D6789E4FF9DD" box="[393,527,1574,1597]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0FE08614FFBB102BD6789E97F9DE" box="[596,727,1573,1598]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB102BD6789E97F9DE" box="[596,727,1573,1598]" italics="true" pageId="23" pageNumber="174">P. variabilis</emphasis>
</taxonomicName>
were established by 
<bibRefCitation id="EFBB09928614FFBB11A2D67899D8F9DE" author="Milne-Edwards" box="[989,1432,1573,1598]" pageId="23" pageNumber="174" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1900) Crustaces decapodes. Premiere partie. Brachyures et Anomoures. In: Milne- Edwards, A. (Ed.), Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882, 1883. Masson, Paris, pp. 1 - 396, 32 pls." type="journal article" year="1900">A. Milne-Edwards &amp; Bouvier (1900)</bibRefCitation>
. Following that opinion, 
<bibRefCitation id="EFBB09928614FFBB13D3D6199E05F980" author="Selbie" box="[428,581,1607,1632]" pageId="23" pageNumber="174" refString="Selbie, C. M. (1921) The Decapoda Reptantia of the coasts of Ireland. Part II: Paguridea. Fisheries Ireland Scientific Investigations, 1, 1 - 68." type="journal article" year="1921">Selbie (1921)</bibRefCitation>
indicated that the most reliable features to separate both species were the number of unpaired pleopods in males (three in 
<taxonomicName id="4C2A0FE08614FFBB10E1D6329F61F963" box="[670,801,1642,1667]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB10E1D6329F61F963" box="[670,801,1642,1667]" italics="true" pageId="23" pageNumber="174">P. variabilis</emphasis>
</taxonomicName>
and four in 
<taxonomicName id="4C2A0FE08614FFBB11DBD632986BF963" box="[932,1067,1644,1667]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB11DBD632986BF963" box="[932,1067,1644,1667]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
) and the length of the dorso-outer process of the antennal peduncular segment 2 that reaches, or passes, the base of the terminal joint in 
<taxonomicName id="4C2A0FE08614FFBB1694D6D09909F946" box="[1259,1353,1677,1702]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB1694D6D09909F946" box="[1259,1353,1677,1702]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
but not in 
<taxonomicName id="4C2A0FE08614FFBB12CCD6EF9D79F928" box="[179,313,1713,1736]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB12CCD6EF9D79F928" box="[179,313,1713,1736]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9574638614FFBB12B8D68D9921F896" blockId="23.[151,1437,533,2014]" pageId="23" pageNumber="174">
<bibRefCitation id="EFBB09928614FFBB12B8D68D9DD9F90B" author="Forest" box="[199,409,1746,1771]" pageId="23" pageNumber="174" refString="Forest, J. (1955) Crustaces Decapodes, Pagurides. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de L'Atlantique Sud (1948 - 1949). Resultats Scientifiques. Institut Royal des Sciences Naturelles de Belge, 3 (4), 23 - 147." type="journal article" year="1955">Forest (1955: 109)</bibRefCitation>
also followed this differentiation, considering 
<taxonomicName id="4C2A0FE08614FFBB11D2D68A9874F90B" box="[941,1076,1748,1771]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB11D2D68A9874F90B" box="[941,1076,1748,1771]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
to be identical to 
<taxonomicName id="4C2A0FE08614FFBB1775D68A9927F90B" box="[1290,1383,1746,1771]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB1775D68A9927F90B" box="[1290,1383,1746,1771]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
(the latter name having priority) and different from 
<taxonomicName id="4C2A0FE08614FFBB10DCD6A89F66F8EE" box="[675,806,1781,1806]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB10DCD6A89F66F8EE" box="[675,806,1781,1806]" italics="true" pageId="23" pageNumber="174">P. variabilis</emphasis>
</taxonomicName>
. This author also indicated the presence of two 
<taxonomicName id="4C2A0FE08614FFBB1740D6A899DCF8ED" box="[1343,1436,1782,1805]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA8718614FFBB1740D6A899DCF8ED" box="[1343,1436,1782,1805]" italics="true" pageId="23" pageNumber="174">Pagurus</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4C2A0FE08614FFBB12BFD7479D78F8D0" box="[192,312,1817,1840]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA8718614FFBB12BFD7479D78F8D0" box="[192,312,1817,1840]" italics="true" pageId="23" pageNumber="174">Eupagurus</emphasis>
</taxonomicName>
) groups in West Africa, the first group including 
<taxonomicName id="4C2A0FE08614FFBB1113D7479F89F8D0" box="[876,969,1815,1840]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB1113D7479F89F8D0" box="[876,969,1815,1840]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
, with four unpaired pleopods in males (a feature of the current concept of 
<taxonomicName id="4C2A0FE08614FFBB106CD7629EDAF8B3" box="[531,666,1852,1875]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB106CD7629EDAF8B3" box="[531,666,1852,1875]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
), and the second group, including 
<taxonomicName id="4C2A0FE08614FFBB1653D76298EFF8B3" box="[1068,1199,1850,1875]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA8718614FFBB1653D76298EFF8B3" box="[1068,1199,1850,1875]" italics="true" pageId="23" pageNumber="174">P. variabilis</emphasis>
</taxonomicName>
, with three unpaired pleopods in males (a feature of the current concept of 
<taxonomicName id="4C2A0FE08614FFBB109BD7009F02F896" box="[740,834,1885,1910]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB109BD7009F02F896" box="[740,834,1885,1910]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
). Forest opinion was shared by 
<bibRefCitation id="EFBB09928614FFBB16E0D703991CF896" author="Zariquiey" box="[1183,1372,1885,1910]" pageId="23" pageNumber="174" refString="Zariquiey Alvarez, R. (1968) Crustaceos Decapodos Ibericos. Investigacion Pesquera, 32, 1 - 510." type="journal article" year="1968">Zariquiey (1968)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9574638614FFB412B8D7219E47FF32" blockId="23.[151,1437,533,2014]" lastBlockId="24.[151,1437,151,315]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
Based on 
<bibRefCitation id="EFBB09928614FFBB134BD7219D8BF878" author="Selbie" box="[308,459,1919,1944]" pageId="23" pageNumber="174" refString="Selbie, C. M. (1921) The Decapoda Reptantia of the coasts of Ireland. Part II: Paguridea. Fisheries Ireland Scientific Investigations, 1, 1 - 68." type="journal article" year="1921">Selbie (1921)</bibRefCitation>
, 
<bibRefCitation id="EFBB09928614FFBB13A8D7219E20F878" author="Ingle" box="[471,608,1919,1944]" pageId="23" pageNumber="174" refString="Ingle, R. W. (1985) Northeastern Atlantic and Mediterranean hermit crabs (Crustacea: Anomura: Paguroidea: Paguridae). I. The genus Pagurus Fabricius, 1775. Journal of Natural History, 19, 745 - 769. http: // dx. doi. org / 10.1080 / 00222938500770461" type="journal article" year="1985">Ingle (1985)</bibRefCitation>
indicated that there were no consistent records of 
<taxonomicName id="4C2A0FE08614FFBB16F2D7DF9953F878" box="[1165,1299,1921,1944]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB16F2D7DF9953F878" box="[1165,1299,1921,1944]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
north of the southern part of the Bay of Biscay. Taking into account that 
<taxonomicName id="4C2A0FE08614FFBB115BD7FA9FC2F85B" box="[804,898,1954,1979]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB115BD7FA9FC2F85B" box="[804,898,1954,1979]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
was described after material collected in 
<collectingCountry id="F33D34F38614FFBB173AD7FC99D8F85B" box="[1349,1432,1954,1979]" name="Iceland" pageId="23" pageNumber="174">Iceland</collectingCountry>
, Ingle concluded that 
<taxonomicName id="4C2A0FE08614FFBB13F4D7989DA8F83E" box="[395,488,1989,2014]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA8718614FFBB13F4D7989DA8F83E" box="[395,488,1989,2014]" italics="true" pageId="23" pageNumber="174">P. alatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0FE08614FFBB1059D7989EEDF83D" box="[550,685,1990,2013]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB1059D7989EEDF83D" box="[550,685,1990,2013]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
were two different species and reinstated 
<taxonomicName id="4C2A0FE08614FFBB16E5D7989960F83D" box="[1178,1312,1990,2013]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA8718614FFBB16E5D7989960F83D" box="[1178,1312,1990,2013]" italics="true" pageId="23" pageNumber="174">P. excavatus</emphasis>
</taxonomicName>
as a valid species. In addition, 
<bibRefCitation id="EFBB0992861BFFB413F3D0C99E5AFF50" author="Ingle" box="[396,538,151,176]" pageId="24" pageNumber="175" refString="Ingle, R. W. (1985) Northeastern Atlantic and Mediterranean hermit crabs (Crustacea: Anomura: Paguroidea: Paguridae). I. The genus Pagurus Fabricius, 1775. Journal of Natural History, 19, 745 - 769. http: // dx. doi. org / 10.1080 / 00222938500770461" type="journal article" year="1985">Ingle (1985)</bibRefCitation>
considered 
<taxonomicName id="4C2A0FE0861BFFB410D4D0C79F6FFF50" box="[683,815,151,176]" class="Malacostraca" family="Paguridae" genus="Eupagurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B95EA871861BFFB410D4D0C79F6FFF50" box="[683,815,151,176]" italics="true" pageId="24" pageNumber="175">P. variabilis</emphasis>
</taxonomicName>
, whose distribution was more boreal, to be a junior subjective synonymy of 
<taxonomicName id="4C2A0FE0861BFFB413DCD0E29E40FF33" box="[419,512,186,211]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA871861BFFB413DCD0E29E40FF33" box="[419,512,186,211]" italics="true" pageId="24" pageNumber="175">P. alatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B957463861BFFB412B8D0839F4BFEDA" blockId="24.[151,1437,151,315]" pageId="24" pageNumber="175">
<bibRefCitation id="EFBB0992861BFFB412B8D0839D17FF16" author="Ingle" box="[199,343,221,246]" pageId="24" pageNumber="175" refString="Ingle, R. W. (1993) Hermit crabs of the Northeastern Atlantic Ocean and the Mediterranean Sea: an illustrated key. Chapman &amp; Hall, London, 495 pp." type="book" year="1993">Ingle (1993)</bibRefCitation>
provided a complete synonymy for 
<taxonomicName id="4C2A0FE0861BFFB4116CD0809F31FF16" box="[787,881,221,246]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA871861BFFB4116CD0809F31FF16" box="[787,881,221,246]" italics="true" pageId="24" pageNumber="175">P. alatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2A0FE0861BFFB411CAD080987CFF15" box="[949,1084,222,245]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA871861BFFB411CAD080987CFF15" box="[949,1084,222,245]" italics="true" pageId="24" pageNumber="175">P. excavatus</emphasis>
</taxonomicName>
, which is currently accepted. Nevertheless, the report of 
<taxonomicName id="4C2A0FE0861BFFB413ABD15F9E71FEF8" box="[468,561,255,280]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="B95EA871861BFFB413ABD15F9E71FEF8" box="[468,561,255,280]" italics="true" pageId="24" pageNumber="175">P. alatus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFBB0992861BFFB41021D15E9EB8FEF8" author="Forest" box="[606,760,255,280]" pageId="24" pageNumber="175" refString="Forest, J. (1955) Crustaces Decapodes, Pagurides. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de L'Atlantique Sud (1948 - 1949). Resultats Scientifiques. Institut Royal des Sciences Naturelles de Belge, 3 (4), 23 - 147." type="journal article" year="1955">Forest (1955)</bibRefCitation>
from Loos Islands and 
<bibRefCitation id="EFBB0992861BFFB4166AD0A19894FEF8" box="[1045,1236,255,280]" pageId="24" pageNumber="175" refString="Zariquiey Alvarez, R. (1968) Crustaceos Decapodos Ibericos. Investigacion Pesquera, 32, 1 - 510." type="journal article">Zariquiey (1968)</bibRefCitation>
from the Iberian Peninsula should be definitively included in 
<taxonomicName id="4C2A0FE0861BFFB41002D17A9F44FEDB" box="[637,772,292,315]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B95EA871861BFFB41002D17A9F44FEDB" box="[637,772,292,315]" italics="true" pageId="24" pageNumber="175">P. excavatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>