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<document id="D92E6ABABABEC2C52B6D3A96300C5155" ID-CLB-Dataset="269966" ID-DOI="10.5281/zenodo.5337424" ID-GBIF-Dataset="7c26f212-ab38-4e92-b461-cdb7dc073af5" ID-ISSN="0253-116X" ID-Zenodo-Dep="5337424" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630178635768" checkinUser="carolina" docAuthor="Puthz, V." docDate="2012" docId="038313247C48FFCBFF01FC843C8EE0F9" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1431-1475.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Stenus pertricosus Puthz 2012, nov.sp." docType="treatment" docVersion="8" lastPageNumber="1451" masterDocId="FFBA6B5C7C5BFFDFFFB3FFE33D00E112" masterDocTitle="On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)" masterLastPageNumber="1475" masterPageNumber="1431" pageNumber="1450" updateTime="1699176368675" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="54B42B113A898840B797DE2392DDAA44">On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)</mods:title>
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<mods:namePart id="BBEEC6BCB7D1B34821800ED3218A1FF5">Puthz, V.</mods:namePart>
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<mods:title id="E1881DB192E28E8104F470E158836470">Linzer biologische Beiträge</mods:title>
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<mods:date id="C3C4BF3866C894E397CE602B9D697EF5">2012</mods:date>
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<mods:number id="E8E41FC338E852A1E3E3F27A78182D6E">2012-12-28</mods:number>
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<treatment id="038313247C48FFCBFF01FC843C8EE0F9" ID-DOI="http://doi.org/10.5281/zenodo.8392900" ID-GBIF-Taxon="213626804" ID-Zenodo-Dep="8392900" LSID="urn:lsid:plazi:treatment:038313247C48FFCBFF01FC843C8EE0F9" httpUri="http://treatment.plazi.org/id/038313247C48FFCBFF01FC843C8EE0F9" lastPageId="20" lastPageNumber="1451" pageId="19" pageNumber="1450">
<subSubSection id="C330F1B97C48FFCCFF01FC843F4EE26F" box="[178,590,871,893]" pageId="19" pageNumber="1450" type="nomenclature">
<paragraph id="8B95A2327C48FFCCFF01FC843F4EE26F" blockId="19.[178,590,871,893]" box="[178,590,871,893]" pageId="19" pageNumber="1450">
<heading id="D0DD155E7C48FFCCFF01FC843F4EE26F" box="[178,590,871,893]" fontSize="8" level="2" pageId="19" pageNumber="1450" reason="2">
<emphasis id="B95E7E207C48FFCCFF01FC843F4EE26F" bold="true" box="[178,590,871,893]" pageId="19" pageNumber="1450">
<taxonomicName id="4C2AD9B17C48FFCCFF01FC843C72E26E" ID-CoL="52B55" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[178,370,871,892]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1450" phylum="Arthropoda" rank="species" species="pertricosus" status="sp. nov.">
<emphasis id="B95E7E207C48FFCCFF01FC843C72E26E" bold="true" box="[178,370,871,892]" italics="true" pageId="19" pageNumber="1450">Stenus pertricosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C48FFCCFECBFC843CC4E26F" box="[376,452,871,893]" pageId="19" pageNumber="1450" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1311BEB77C48FFCCFE62FC843F21E26F" box="[465,545,871,893]" captionStart="Figs 42-54" captionStartId="41.[178,218,1315,1335]" captionTargetBox="[178,1085,225,1295]" captionTargetId="figure-158@41.[178,1085,219,1295]" captionTargetPageId="41" captionText="Figs 42-54: Ventral aspect of aedeagus (42-45), paraglossa (46, 47), metasternum (48), spermatheca (49-54) of Stenus perpunctus nov.sp. (42, T 69, PT), S. perscitus nov.sp. (43, T28, PT), S. pertricosus nov.sp. (44, T 61, PT), S. perfectus nov.sp. (45, T 119, PT), S. perpauperculus nov.sp. (46, 52, PTT), S. pilicornis FAUVEL (47, 48), S. mysterialis PUTHZ (49), S. perpusillus nov.sp. (50), S. perpropinquus nov.sp. (51, T 52, PT), S. perpauper nov.sp. (53,T 60, PT), S. perpastus (54, T 91, PT). Scale bar = 0.1mm (42=43-45)." figureDoi="http://doi.org/10.5281/zenodo.5337432" httpUri="https://zenodo.org/record/5337432/files/figure.png" pageId="19" pageNumber="1450">Figs 44</figureCitation>
,
<figureCitation id="1311BEB77C48FFCCFD9FFC843F46E26F" box="[556,582,871,893]" captionStart="Figs 70-77" captionStartId="43.[178,218,1128,1148]" captionTargetBox="[178,1085,219,1108]" captionTargetId="figure-109@43.[178,1085,219,1108]" captionTargetPageId="43" captionText="Figs 70-77: Spermathecae of PTT (70-75), elytra (76), semilateral aspect of abdominal segments 4- 7 (77) of Stenus permolestus nov.sp. (70, T 116), S. pertricosus nov.sp. (71, T 61), S. permodestus nov.sp. (72, T 64), S. perscitus nov.sp. (73, T 28), S. perfectus nov.sp. (74, T 19), S. pertenuis nov.sp. (75, T 161), S. nefas PUTHZ (76, 77, HT)." figureDoi="http://doi.org/10.5281/zenodo.5337436" httpUri="https://zenodo.org/record/5337436/files/figure.png" pageId="19" pageNumber="1450">71</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C48FFCCFF01FC743CCCE529" pageId="19" pageNumber="1450" type="materials_examined">
<paragraph id="8B95A2327C48FFCCFF01FC743CCCE529" blockId="19.[178,1087,919,1083]" pageId="19" pageNumber="1450">
<materialsCitation id="3B42A86F7C48FFCCFF01FC743F2BE2B9" ID-GBIF-Occurrence="4417845317" box="[178,555,919,939]" collectingDate="1976-08-22" collectionCode="Shibata" collectorName="Y. Shibata" country="Taiwan" elevation="2200" location="Tsuifeng" pageId="19" pageNumber="1450" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
Type material:
<typeStatus id="54911C907C48FFCCFE1DFC743CFFE2B9" box="[430,511,919,939]" pageId="19" pageNumber="1450" type="holotype">Holotype</typeStatus>
()
</materialsCitation>
<materialsCitation id="3B42A86F7C48FFCCFD86FC743F6FE2D1" ID-GBIF-Occurrence="4417845436" collectingDate="1976-08-22" collectionCode="Shibata, cN, cT, cP" collectorName="Y. Shibata" country="Taiwan" elevation="2200" location="Tsuifeng" pageId="19" pageNumber="1450" specimenCount="10" stateProvince="Nantou" typeStatus="paratype">
and 5, 5 -
<typeStatus id="54911C907C48FFCCFD60FC743E2BE2B9" box="[723,811,919,939]" pageId="19" pageNumber="1450" type="paratype">paratypes</typeStatus>
:
<collectingCountry id="F33DE2A27C48FFCCFC8BFC743E7DE2B9" box="[824,893,919,939]" name="Taiwan" pageId="19" pageNumber="1450">Taiwan</collectingCountry>
:
<collectingRegion id="49EE6CD07C48FFCCFC39FC74390AE2B9" box="[906,1034,919,939]" country="Taiwan" name="Nantou" pageId="19" pageNumber="1450">Nantou Hsien</collectingRegion>
: near
<location id="8EF5F4E97C48FFCCFF6CFC4C3C2FE2D1" LSID="urn:lsid:plazi:treatment:038313247C48FFCBFF01FC843C8EE0F9:8EF5F4E97C48FFCCFF6CFC4C3C2FE2D1" box="[223,303,943,963]" country="Taiwan" name="Tsuifeng" pageId="19" pageNumber="1450" stateProvince="Nantou">Tsuifeng</location>
,
<quantity id="4CD20FD77C48FFCCFE8EFC4C3C7CE2D1" box="[317,380,943,963]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="1450" unit="m" value="2200.0">
<elevation id="000745017C48FFCCFE8EFC4C3C7CE2D1" box="[317,380,943,963]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="1450" unit="m" value="2200.0">2200m</elevation>
</quantity>
,
<date id="FF9484F27C48FFCCFE39FC4C3CFDE2D1" box="[394,509,943,963]" pageId="19" pageNumber="1450" value="1976-08-22">
<collectingDate id="EFD07D1A7C48FFCCFE39FC4C3CFDE2D1" box="[394,509,943,963]" pageId="19" pageNumber="1450" value="1976-08-22">22.VIII.1976</collectingDate>
</date>
,
<collectorName id="26DFC7E47C48FFCCFDB8FC4C3F6FE2D1" box="[523,623,943,963]" pageId="19" pageNumber="1450">Y. Shibata</collectorName>
</materialsCitation>
.
<materialsCitation id="3B42A86F7C48FFCCFDCFFC4C3F1BE2E1" ID-GBIF-Occurrence="4417845393" collectingDate="1974-08-25" collectingDateMax="1977-08-11" collectingDateMin="1973-08-23" collectionCode="Shibata, cN, cT, cP" country="Taiwan" pageId="19" pageNumber="1450" specimenCount="40" stateProvince="Nantou" typeStatus="paratype">
<typeStatus id="54911C907C48FFCCFDCFFC4C3FD4E2D1" box="[636,724,943,963]" pageId="19" pageNumber="1450" type="paratype">Paratypes</typeStatus>
: 12, 28: same as HT, 28.VII., 22.VIII., 23.VIII.1973, 26.VII.,
<date id="FF9484F27C48FFCCFE4DFC243F73E2C9" box="[510,627,967,987]" pageId="19" pageNumber="1450" value="1974-08-25">
<collectingDate id="EFD07D1A7C48FFCCFE4DFC243F73E2C9" box="[510,627,967,987]" pageId="19" pageNumber="1450" value="1974-08-25">25.VIII.1974</collectingDate>
</date>
; 28.VII., 29. VII., 18.VIII., 20.VIII., 21.VIII.1976, 23.VII., 2.VIII., 11.VIII.1977, idem
</materialsCitation>
;
<materialsCitation id="3B42A86F7C48FFCCFD97FC3C3EFEE2E1" ID-GBIF-Occurrence="4417845401" box="[548,1022,991,1011]" collectingDate="1981-08-13" collectingDateMax="1977-08-11" collectingDateMin="1976-08-21" collectionCode="Shibata, cN, cT, cP" country="Taiwan" elevation="2150" location="Meifeng" pageId="19" pageNumber="1450" specimenCount="3" stateProvince="Nantou" typeStatus="paratype">
1, 2: near
<location id="8EF5F4E97C48FFCCFD06FC3C3E01E2E1" LSID="urn:lsid:plazi:treatment:038313247C48FFCBFF01FC843C8EE0F9:8EF5F4E97C48FFCCFD06FC3C3E01E2E1" box="[693,769,991,1011]" name="Meifeng" pageId="19" pageNumber="1450">Meifeng</location>
,
<quantity id="4CD20FD77C48FFCCFCB9FC3C3E49E2E1" box="[778,841,991,1011]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="19" pageNumber="1450" unit="m" value="2150.0">
<elevation id="000745017C48FFCCFCB9FC3C3E49E2E1" box="[778,841,991,1011]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="19" pageNumber="1450" unit="m" value="2150.0">2150m</elevation>
</quantity>
,
<date id="FF9484F27C48FFCCFCE0FC3C3EC6E2E1" box="[851,966,991,1011]" pageId="19" pageNumber="1450" value="1981-08-13">
<collectingDate id="EFD07D1A7C48FFCCFCE0FC3C3EC6E2E1" box="[851,966,991,1011]" pageId="19" pageNumber="1450" value="1981-08-13">13.VIII.1981</collectingDate>
</date>
, idem
</materialsCitation>
;
<materialsCitation id="3B42A86F7C48FFCCFBB4FC3C3FBFE519" ID-GBIF-Occurrence="4417845553" collectingDate="1991-05-03" collectionCode="Shibata, cN, cT, cP" collectorName="A. Smetana" country="Taiwan" elevation="2130" location="Meifeng" pageId="19" pageNumber="1450" specimenCount="4" stateProvince="Nantou" typeStatus="paratype">
3, 1:
<location id="8EF5F4E97C48FFCCFEBFFC143C58E519" LSID="urn:lsid:plazi:treatment:038313247C48FFCBFF01FC843C8EE0F9:8EF5F4E97C48FFCCFEBFFC143C58E519" box="[268,344,1015,1035]" country="Taiwan" name="Meifeng" pageId="19" pageNumber="1450" stateProvince="Nantou">Meifeng</location>
,
<quantity id="4CD20FD77C48FFCCFED7FC143CA4E519" box="[356,420,1015,1035]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="19" pageNumber="1450" unit="m" value="2130.0">
<elevation id="000745017C48FFCCFED7FC143CA4E519" box="[356,420,1015,1035]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="19" pageNumber="1450" unit="m" value="2130.0">2130m</elevation>
</quantity>
,
<date id="FF9484F27C48FFCCFE1CFC143F02E519" box="[431,514,1015,1035]" pageId="19" pageNumber="1450" value="1991-05-03">
<collectingDate id="EFD07D1A7C48FFCCFE1CFC143F02E519" box="[431,514,1015,1035]" pageId="19" pageNumber="1450" value="1991-05-03">3.V.1991</collectingDate>
</date>
,
<collectorName id="26DFC7E47C48FFCCFDBCFC143F78E519" box="[527,632,1015,1035]" pageId="19" pageNumber="1450">A. Smetana</collectorName>
(T 61)
</materialsCitation>
;
<materialsCitation id="3B42A86F7C48FFCCFD7FFC14393AE519" ID-GBIF-Occurrence="4417845521" box="[716,1082,1015,1035]" collectingDate="1993-07-10" collectionCode="Shibata, cN, cT, cP" collectorName="A. Smetana" country="Taiwan" elevation="2130" location="Meifeng" pageId="19" pageNumber="1450" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
1: ibidem,
<date id="FF9484F27C48FFCCFCF7FC143EAFE519" box="[836,943,1015,1035]" pageId="19" pageNumber="1450" value="1993-07-10">
<collectingDate id="EFD07D1A7C48FFCCFCF7FC143EAFE519" box="[836,943,1015,1035]" pageId="19" pageNumber="1450" value="1993-07-10">10.VII.1993</collectingDate>
</date>
, idem (T 146)
</materialsCitation>
;
<materialsCitation id="3B42A86F7C48FFCCFF6CFBEC3CC9E529" ID-GBIF-Occurrence="4417845318" collectingDate="1998-05-04" collectionCode="Shibata, cN, cT, cP" collectorName="A. Smetana" country="Taiwan" elevation="2130" location="Meifeng" pageId="19" pageNumber="1450" specimenCount="2" stateProvince="Nantou" typeStatus="paratype">
2: ibidem,
<date id="FF9484F27C48FFCCFED7FBEC3CB7E531" box="[356,439,1039,1059]" pageId="19" pageNumber="1450" value="1998-05-04">
<collectingDate id="EFD07D1A7C48FFCCFED7FBEC3CB7E531" box="[356,439,1039,1059]" pageId="19" pageNumber="1450" value="1998-05-04">4.V.1998</collectingDate>
</date>
, idem (T 199).- HT and PTT in coll.
<collectorName id="26DFC7E47C48FFCCFCB8FBEC3E4CE531" box="[779,844,1039,1059]" pageId="19" pageNumber="1450">Shibata</collectorName>
(
<collectorName id="26DFC7E47C48FFCCFCEAFBEC3E96E531" box="[857,918,1039,1059]" pageId="19" pageNumber="1450">Tokyo</collectorName>
), PTT also in coll.
<collectorName id="26DFC7E47C48FFCCFF6CFBC43C2CE529" box="[223,300,1063,1083]" pageId="19" pageNumber="1450">Smetana</collectorName>
, cN, cT and in cP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C48FFCBFF01FBA93C4EE077" lastPageId="20" lastPageNumber="1451" pageId="19" pageNumber="1450" type="description">
<paragraph id="8B95A2327C48FFCCFF01FBA93FDDE54D" blockId="19.[178,733,1097,1119]" box="[178,733,1097,1119]" pageId="19" pageNumber="1450">
Description: About as in
<taxonomicName id="4C2AD9B17C48FFCCFDBBFBA93F8DE54D" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[520,653,1098,1119]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1450" phylum="Arthropoda" rank="species" species="persubtilis" status="sp. nov.">
<emphasis id="B95E7E207C48FFCCFDBBFBA93F8DE54D" box="[520,653,1098,1119]" italics="true" pageId="19" pageNumber="1450">S. persubtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C48FFCCFD20FBA93FDDE54D" box="[659,733,1098,1119]" pageId="19" pageNumber="1450" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B95A2327C48FFCCFF01FB8C3F6CE597" blockId="19.[178,620,1135,1157]" box="[178,620,1135,1157]" pageId="19" pageNumber="1450">
Length:
<quantity id="4CD20FD77C48FFCCFEB4FB8C3C76E597" box="[263,374,1135,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="19" pageNumber="1450" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.8-3.4mm</quantity>
(forebody:
<quantity id="4CD20FD77C48FFCCFE5DFB8C3F60E597" box="[494,608,1135,1157]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="19" pageNumber="1450" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5-1.6mm</quantity>
).
</paragraph>
<paragraph id="8B95A2327C48FFCCFF01FB763DE5E5D5" blockId="19.[178,1086,1172,1223]" pageId="19" pageNumber="1450">PM of a -PT from T 61: HW: 30; DE: 16; PW: 21.5; PL: 22; EW: 27.3; EL: 24; SL: 17.5.</paragraph>
<paragraph id="8B95A2327C48FFCCFF01FB343F84E4C4" blockId="19.[178,1086,1239,1495]" pageId="19" pageNumber="1450">Male: Metasternum distinctly impressed, moderately coarsely, densely punctate, interstices smaller than half diameter of punctures. Anterior sternites simple, sternite 6 finer and denser punctate and pubescent medially than laterally, sternite 7 with a shallow posteromedian impression, which is very finely and very densely punctate and pubescent. Sternite 8 with a shallow, moderately broad apical emargination (LS: ED= 53:3). Sternite 9 with comparatively short, blackish apicolateral teeth. Aedeagus (fig. 44), median lobe broadened in anterior half, apical portion obtusely angled narrowed, apex small and prominent, expulsion hooks connected medially; parameres slightly longer than median lobe, with about 6-7 apical setae.</paragraph>
<paragraph id="8B95A2327C48FFCCFF01FA043C0EE725" blockId="19.[178,1085,1510,1591]" pageId="19" pageNumber="1450">Female: Sternite 8 at posterior margin feebly projecting, rounded apically. Valvifer with a short apicolateral tooth. Spermatheca (fig. 71), infundibulum narrow, 3x as long as broad.</paragraph>
<paragraph id="8B95A2327C4FFFCBFF01FF383C4EE077" blockId="20.[178,1086,219,358]" pageId="20" pageNumber="1451">
Median portion of frons strongly elevated, distinctly extending beyond the level of medial eye margins; pronotum and elytra with deep impressions, punctation coarser than in
<taxonomicName id="4C2AD9B17C4FFFCBFF7DFEF53C54E038" authorityName="Puthz" authorityYear="2012" box="[206,340,277,299]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1451" phylum="Arthropoda" rank="species" species="persubtilis">
<emphasis id="B95E7E207C4FFFCBFF7DFEF53C54E038" box="[206,340,277,299]" italics="true" pageId="20" pageNumber="1451">S. persubtilis</emphasis>
</taxonomicName>
, diameter of elytral punctures at least as large as apical cross section of antennal segment 2; abdomen densely punctate, interstices on tergite 7 distinctly smaller than punctures.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4FFFCBFF01FE963EA5E0BA" pageId="20" pageNumber="1451" type="discussion">
<paragraph id="8B95A2327C4FFFCBFF01FE963EA5E0BA" blockId="20.[178,1085,373,424]" pageId="20" pageNumber="1451">Comparative note: This is a relatively large species, it may be distinguished from the close relatives as quoted in the key, best by the sexual characters.</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4FFFCBFF01FE5B3C8EE0F9" pageId="20" pageNumber="1451" type="etymology">
<paragraph id="8B95A2327C4FFFCBFF01FE5B3C8EE0F9" blockId="20.[178,1085,439,491]" pageId="20" pageNumber="1451">
Etymology: Since is it difficult to identify this species I name it &quot;
<emphasis id="B95E7E207C4FFFCBFC72FE5B3933E0DF" box="[961,1075,440,461]" italics="true" pageId="20" pageNumber="1451">pertricosus</emphasis>
&quot; (Lat.= very intricate).
</paragraph>
</subSubSection>
</treatment>
</document>