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<document id="D6EA33939209697D3BA0708B4D498441" ID-CLB-Dataset="276483" ID-DOI="10.1111/j.1096-3642.2009.00552.x" ID-GBIF-Dataset="3be82545-1797-46a0-95fc-3e0fc44c9af0" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440497297" checkinUser="valdenar" docAuthor="Zhang, Kuiyan, Yang, Ding, Ren, Dong &amp; Shih, Chungkun" docDate="2010" docId="038296595834FF8AFC8F93FCFAF15267" docLanguage="en" docName="j.1096-3642.2009.00552.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Strenorhagio conjugovenius Zhang &amp; Yang &amp; Ren &amp; Shih 2010" docType="treatment" docVersion="6" lastPageNumber="570" masterDocId="FFBBEE215832FF8DFFBB9529FFCA5766" masterDocTitle="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)" masterLastPageNumber="572" masterPageNumber="563" pageNumber="569" updateTime="1699284570084" updateUser="ExternalLinkService">
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<mods:title id="6A36A0955B43367DE023F5215662A2E8">An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)</mods:title>
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<mods:namePart id="FE05841DD9B5D47E46BD1A5C300AE5B3">Zhang, Kuiyan</mods:namePart>
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<mods:namePart id="6EB6BCF9ECC8ECF821F128C95B63DFD2">Yang, Ding</mods:namePart>
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<mods:namePart id="F4CC0118586685D7222E22AA2E4E87C9">Ren, Dong</mods:namePart>
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<mods:namePart id="B16A8FB7F95A49986183870217F6E843">Shih, Chungkun</mods:namePart>
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<treatment id="038296595834FF8AFC8F93FCFAF15267" ID-DOI="http://doi.org/10.5281/zenodo.5491008" ID-GBIF-Taxon="217238223" ID-Zenodo-Dep="5491008" LSID="urn:lsid:plazi:treatment:038296595834FF8AFC8F93FCFAF15267" httpUri="http://treatment.plazi.org/id/038296595834FF8AFC8F93FCFAF15267" lastPageId="7" lastPageNumber="570" pageId="6" pageNumber="569">
<subSubSection id="C33174C45834FF8BFC8F93FCFAC0506C" pageId="6" pageNumber="569" type="nomenclature">
<paragraph id="8B94274F5834FF8BFC8F93FCFAC0506C" blockId="6.[820,1407,1749,1802]" pageId="6" pageNumber="569">
<heading id="D0DC90235834FF8BFC8F93FCFAC0506C" centered="true" fontSize="9" level="2" pageId="6" pageNumber="569" reason="4">
<taxonomicName id="4C2B5CCC5834FF8BFC8F93FCFB77518C" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[820,1213,1749,1770]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="570" phylum="Arthropoda" rank="species" species="conjugovenius" status="sp. nov.">
<emphasis id="B95FFB5D5834FF8BFC8F93FCFB77518C" bold="true" box="[820,1213,1749,1770]" italics="true" pageId="6" pageNumber="569">STRENORHAGIO CONJUGOVENIUS</emphasis>
</taxonomicName>
ZHANG, YANG &amp; REN
<emphasis id="B95FFB5D5834FF8BFC5B93D1FB64506C" bold="true" box="[992,1198,1781,1802]" pageId="6" pageNumber="569">GEN. ET SP. NOV.</emphasis>
(
<figureCitation id="13103BCA5834FF8BFB7A93D0FAC8506C" box="[1217,1282,1781,1802]" captionStart="Figure 7" captionStartId="7.[164,243,690,709]" captionTargetBox="[164,1443,199,659]" captionTargetId="figure-594@7.[743,1444,199,660]" captionTargetPageId="7" captionText="Figure 7. Strenorhagio conjugovenius sp. nov. Holotype, body with wings in dorsal view. A, photograph. B, camera lucida drawing." figureDoi="http://doi.org/10.5281/zenodo.5438136" httpUri="https://zenodo.org/record/5438136/files/figure.png" pageId="6" pageNumber="569">FIG. 7</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33174C45834FF8AFC9D9235FAF15267" lastPageId="7" lastPageNumber="570" pageId="6" pageNumber="569" type="etymology">
<paragraph id="8B94274F5834FF8BFC9D9235FB105009" blockId="6.[806,1421,1820,1903]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFC9D9235FC6E5057" box="[806,932,1820,1841]" italics="true" pageId="6" pageNumber="569">Etymology:</emphasis>
The specific name refers to the Latin
<emphasis id="B95FFB5D5834FF8BFC909212FC485036" box="[811,898,1851,1872]" italics="true" pageId="6" pageNumber="569">conjugo</emphasis>
(conjoint) and ve
<emphasis id="B95FFB5D5834FF8BFBDB9212FB5B5036" box="[1120,1169,1851,1872]" italics="true" pageId="6" pageNumber="569">nius</emphasis>
(vein), because vein R
<subScript id="17AF250A5834FF8BFC83924AFC9A5009" attach="left" box="[824,848,1891,1903]" fontSize="5" pageId="6" pageNumber="569">2+3</subScript>
and crossvein rm are coalesced.
</paragraph>
<caption id="DF5477C75835FF8AFF1F979BFE805585" ID-DOI="http://doi.org/10.5281/zenodo.5438136" ID-Zenodo-Dep="5438136" httpUri="https://zenodo.org/record/5438136/files/figure.png" pageId="7" pageNumber="570" startId="7.[164,243,690,709]" targetBox="[164,1443,199,659]" targetPageId="7">
<paragraph id="8B94274F5835FF8AFF1F979BFE805585" blockId="7.[164,1441,690,739]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFF1F979BFD0955A0" bold="true" box="[164,707,690,710]" pageId="7" pageNumber="570">
Figure 7.
<taxonomicName id="4C2B5CCC5835FF8AFEA2979BFDA855A3" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[281,610,690,709]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="conjugovenius" status="sp. nov.">
<emphasis id="B95FFB5D5835FF8AFEA2979BFDA855A3" bold="true" box="[281,610,690,709]" italics="true" pageId="7" pageNumber="570">Strenorhagio conjugovenius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265835FF8AFDD1979AFD0955A0" box="[618,707,691,710]" pageId="7" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, body with wings in dorsal view. A, photograph. B, camera lucida drawing.
</paragraph>
</caption>
<paragraph id="8B94274F5835FF8AFF1F9633FEA354CF" blockId="7.[164,779,793,938]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFF1F9633FED15449" box="[164,283,794,815]" italics="true" pageId="7" pageNumber="570">Diagnosis:</emphasis>
Hind femur swollen at apical part. Crossvein rm at basal third of cell d, coalesced with vein R
<subScript id="17AF250A5835FF8AFF0D9648FF04540B" attach="left" box="[182,206,865,877]" fontSize="5" pageId="7" pageNumber="570">2+3</subScript>
at vein Rs (with vein Rs
<subScript id="17AF250A5835FF8AFDB89648FDC1540B" attach="left" box="[515,523,865,877]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
having disappeared); mouth of cell sc wide open, about three times as wide as that of cell r
<subScript id="17AF250A5835FF8AFEE196B7FEA854CC" attach="left" box="[346,354,926,938]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
.
</paragraph>
<paragraph id="8B94274F5835FF8AFF1F96FFFD97536D" blockId="7.[164,778,982,1035]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFF1F96FFFEC4548D" box="[164,270,982,1003]" italics="true" pageId="7" pageNumber="570">
<typeStatus id="549099ED5835FF8AFF1F96FFFEC2548D" box="[164,264,982,1003]" pageId="7" pageNumber="570" type="holotype">Holotype</typeStatus>
:
</emphasis>
CNU-DIB-NN2007022, an almost complete adult body with wings in dorsal view.
</paragraph>
<paragraph id="8B94274F5835FF8AFF1F9111FE5953CF" blockId="7.[164,779,1080,1193]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFF1F9111FE00532B" box="[164,458,1080,1101]" italics="true" pageId="7" pageNumber="570">Type locality and horizon:</emphasis>
Daohugou Village, Shantou Township, Ningcheng County,
<collectingRegion id="49EFE9AD5835FF8AFDBB917EFD79530A" box="[512,691,1110,1132]" country="China" name="Nei Mongol" pageId="7" pageNumber="570">Inner Mongolia</collectingRegion>
,
<collectingCountry id="F33C67DF5835FF8AFD05917FFCCC530A" box="[702,774,1110,1132]" name="China" pageId="7" pageNumber="570">China</collectingCountry>
; Jiulongshan Formation, the Middle Jurassic (AalenianBajocian).
</paragraph>
<paragraph id="8B94274F5835FF8AFF1F91FEFDCD526C" blockId="7.[164,779,1238,1903]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFF1F91FEFEE6538A" box="[164,300,1239,1260]" italics="true" pageId="7" pageNumber="570">Description:</emphasis>
Body length, 14.0 mm; wing length,
<quantity id="4CD38AAA5835FF8AFF1F91DCFEC3526C" box="[164,265,1269,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="7" pageNumber="570" unit="mm" value="10.5">10.5 mm</quantity>
; wing width,
<quantity id="4CD38AAA5835FF8AFE1191DCFDC8526C" box="[426,514,1269,1290]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="570" unit="mm" value="3.7">3.7 mm</quantity>
.
</paragraph>
<paragraph id="8B94274F5835FF8AFF07903AFD815201" blockId="7.[164,779,1238,1903]" pageId="7" pageNumber="570">Body stout. Head round, slightly narrower than thorax. Eyes large, bare; ommatidia clearly visible, upper facets not distinctly enlarged.</paragraph>
<paragraph id="8B94274F5835FF8AFF079059FE5152A4" blockId="7.[164,779,1238,1903]" pageId="7" pageNumber="570">Legs pubescent. Hind femur swollen at apical part, tibia slender. First tarsomere distinctly longer than following tarsomeres.</paragraph>
<paragraph id="8B94274F5835FF8AFF0790E2FAF454CF" blockId="7.[164,779,1238,1903]" lastBlockId="7.[826,1441,793,999]" pageId="7" pageNumber="570">
Wing hyaline; veins thick and strong. Vein Sc ending slightly beyond middle of wing. Veins R
<subScript id="17AF250A5835FF8AFD7590DDFD1C5166" attach="left" box="[718,726,1524,1536]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
and Rs
<subScript id="17AF250A5835FF8AFF7A933BFF035178" attach="left" box="[193,201,1554,1566]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
long; vein R
<subScript id="17AF250A5835FF8AFEC6933BFE5C5178" attach="left" box="[381,406,1554,1566]" fontSize="5" pageId="7" pageNumber="570">2+3</subScript>
distinctly and strongly bent upwards at base; vein R
<subScript id="17AF250A5835FF8AFE039318FE0A515B" attach="left" box="[440,448,1585,1597]" fontSize="5" pageId="7" pageNumber="570">5</subScript>
ending slightly beyond wing apex. Crossvein rm at basal third of cell d, and coalesced with vein R
<subScript id="17AF250A5835FF8AFE109347FE0E511C" attach="left" box="[427,452,1646,1658]" fontSize="5" pageId="7" pageNumber="570">2+3</subScript>
at vein Rs (with vein Rs
<subScript id="17AF250A5835FF8AFCB99347FCC0511C" attach="left" box="[770,778,1646,1658]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
disappearing); vein Rs
<subScript id="17AF250A5835FF8AFE0893A4FE7151FF" attach="left" box="[435,443,1677,1689]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
shorter than vein Rs
<subScript id="17AF250A5835FF8AFD7693A4FD1F51FF" attach="right" box="[717,725,1677,1689]" fontSize="5" pageId="7" pageNumber="570">3</subScript>
. All branches of vein M long, veins M
<subScript id="17AF250A5835FF8AFD999382FDE051D1" attach="left" box="[546,554,1707,1719]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
and M
<subScript id="17AF250A5835FF8AFDC69382FD4F51D1" attach="left" box="[637,645,1707,1719]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
bifurcating distal of crossvein mm, basal trunk of veins M
<subScript id="17AF250A5835FF8AFD7493E3FD1D51B0" attach="left" box="[719,727,1738,1750]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
and M
<subScript id="17AF250A5835FF8AFF0093C1FF095192" attach="left" box="[187,195,1768,1780]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
slightly shorter than crossvein mm. Vein CuA
<subScript id="17AF250A5835FF8AFCB993C1FCC05192" attach="left" box="[770,778,1768,1780]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
arising from cell bm; vein CuP present. Mouth of cell sc wide open, about three times as wide as that of cell r
<subScript id="17AF250A5835FF8AFF14926DFF7D5036" attach="left" box="[175,183,1860,1872]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
; relative length of mouths of cells r
<subScript id="17AF250A5835FF8AFDD4926DFDBD5036" attach="left" box="[623,631,1860,1872]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
, r
<subScript id="17AF250A5835FF8AFD2E926DFD675036" attach="left" box="[661,685,1860,1872]" fontSize="5" pageId="7" pageNumber="570">2+3</subScript>
, and r
<subScript id="17AF250A5835FF8AFCB9926DFCC05036" attach="left" box="[770,778,1860,1872]" fontSize="5" pageId="7" pageNumber="570">4</subScript>
about 4: 5: 4. Cell bm about as wide as cell br; cell d slender, far from wing margin. Five posterior cells present and wide open; mouths of cell m
<subScript id="17AF250A5835FF8AFA8E966BFAF75428" attach="left" box="[1333,1341,834,846]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
slightly narrower than that of cell m
<subScript id="17AF250A5835FF8AFB169648FB7F540B" attach="left" box="[1197,1205,865,877]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
; mouth of cell m
<subScript id="17AF250A5835FF8AFA229648FA6B540B" attach="left" box="[1433,1441,865,877]" fontSize="5" pageId="7" pageNumber="570">3</subScript>
narrow, about 2/5 as wide as that of cell cua
<subScript id="17AF250A5835FF8AFA959656FAFC54ED" attach="left" box="[1326,1334,895,907]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
. Cell cup distinctly open; anal lobe small and narrow.
</paragraph>
<paragraph id="8B94274F5835FF8AFCE8969BFA945481" blockId="7.[826,1441,793,999]" pageId="7" pageNumber="570">Abdomen cylindrical, robust, and pubescent. Eight segments visible; segment I evidently shortest.</paragraph>
<paragraph id="8B94274F5835FF8AFC81913FFAF15267" blockId="7.[826,1441,1046,1282]" pageId="7" pageNumber="570">
<emphasis id="B95FFB5D5835FF8AFC81913FFB25534D" box="[826,1263,1046,1067]" italics="true" pageId="7" pageNumber="570">
Remarks:
<taxonomicName id="4C2B5CCC5835FF8AFC0A913FFB25534D" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[945,1263,1046,1067]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="conjugovenius" status="sp. nov.">Strenorhagio conjugovenius</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A26C46265835FF8AFB40913FFA9E534D" box="[1275,1364,1046,1067]" pageId="7" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
is the fourth species in
<taxonomicName id="4C2B5CCC5835FF8AFBBF911DFB52532F" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1028,1176,1076,1097]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5835FF8AFBBF911DFB52532F" box="[1028,1176,1076,1097]" italics="true" pageId="7" pageNumber="570">Strenorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265835FF8AFB1A911CFACD532C" box="[1185,1287,1077,1098]" pageId="7" pageNumber="570" rank="genus">gen. nov.</taxonomicNameLabel>
It looks very much like
<taxonomicName id="4C2B5CCC5835FF8AFC04917AFB4B530F" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[959,1153,1107,1129]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5835FF8AFC04917AFB4B530F" box="[959,1153,1107,1129]" italics="true" pageId="7" pageNumber="570">S. asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265835FF8AFB34917AFB27530E" box="[1167,1261,1107,1128]" pageId="7" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
, but there are some unique characters to separate it from the others in the genus: the crossvein rm and vein R
<subScript id="17AF250A5835FF8AFAED91B3FAA453C0" attach="left" box="[1366,1390,1178,1190]" fontSize="5" pageId="7" pageNumber="570">2+3</subScript>
are coalesced at vein Rs (with vein Rs
<subScript id="17AF250A5835FF8AFB6A9190FB1353A3" attach="left" box="[1233,1241,1209,1221]" fontSize="5" pageId="7" pageNumber="570">2</subScript>
disappearing) in both wings; the mouth of cell sc is wide open, and about three times as wide as that of cell r
<subScript id="17AF250A5835FF8AFA9791DFFAFE5264" attach="left" box="[1324,1332,1270,1282]" fontSize="5" pageId="7" pageNumber="570">1</subScript>
.
</paragraph>
</subSubSection>
</treatment>
</document>