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<document id="D617BDCCE457C5EDB57F2E595D41AE41" ID-CLB-Dataset="276483" ID-DOI="10.1111/j.1096-3642.2009.00552.x" ID-GBIF-Dataset="3be82545-1797-46a0-95fc-3e0fc44c9af0" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440497297" checkinUser="valdenar" docAuthor="Zhang, Kuiyan, Yang, Ding, Ren, Dong &amp; Shih, Chungkun" docDate="2010" docId="038296595831FF89FE809384FF3554C2" docLanguage="en" docName="j.1096-3642.2009.00552.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Strenorhagio Zhang &amp; Yang &amp; Ren &amp; Shih 2010, GEN. NOV." docType="treatment" docVersion="5" lastPageNumber="566" masterDocId="FFBBEE215832FF8DFFBB9529FFCA5766" masterDocTitle="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)" masterLastPageNumber="572" masterPageNumber="563" pageNumber="566" updateTime="1699284570084" updateUser="ExternalLinkService">
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<mods:title id="4AD4827023611B2302656943C06164F3">An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)</mods:title>
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<mods:namePart id="C66C6CC2395B7303DEC95A7CA07C05CA">Zhang, Kuiyan</mods:namePart>
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<mods:namePart id="F9AC0F86CBA1E3C9EA988AF373297AE9">Yang, Ding</mods:namePart>
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<mods:namePart id="F45EECF3B89A9426C738B78C191E2C44">Ren, Dong</mods:namePart>
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<mods:namePart id="CEE0D6C5D3E6F0BF35BE585DE88CB657">Shih, Chungkun</mods:namePart>
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<treatment id="038296595831FF89FE809384FF3554C2" ID-GBIF-Taxon="217238219" LSID="urn:lsid:plazi:treatment:038296595831FF89FE809384FF3554C2" httpUri="http://treatment.plazi.org/id/038296595831FF89FE809384FF3554C2" lastPageId="4" lastPageNumber="566" pageId="3" pageNumber="566">
<subSubSection id="C33174C45831FF8EFE809384FDBE51D9" box="[315,628,1706,1727]" pageId="3" pageNumber="566" type="nomenclature">
<paragraph id="8B94274F5831FF8EFE809384FDBE51D9" blockId="3.[315,628,1706,1727]" box="[315,628,1706,1727]" pageId="3" pageNumber="566">
<heading id="D0DC90235831FF8EFE809384FDBE51D9" box="[315,628,1706,1727]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="566" reason="4">
<taxonomicName id="4C2B5CCC5831FF8EFE809384FE3B51D8" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[315,497,1709,1726]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">STRENORHAGIO</taxonomicName>
<emphasis id="B95FFB5D5831FF8EFE409384FDBE51D9" bold="true" box="[507,628,1706,1727]" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A26C46265831FF8EFE409384FDBE51D9" box="[507,628,1706,1727]" pageId="3" pageNumber="566" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33174C45831FF89FF1F93F9FF3554C2" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566" type="etymology">
<paragraph id="8B94274F5831FF8EFF1F93F9FF1A5045" blockId="3.[164,779,1744,1827]" pageId="3" pageNumber="566">
<emphasis id="B95FFB5D5831FF8EFF1F93F9FEE85183" box="[164,290,1744,1765]" italics="true" pageId="3" pageNumber="566">Etymology:</emphasis>
The generic name is derived from the Greek S
<emphasis id="B95FFB5D5831FF8EFEBF93C6FEF85062" box="[260,306,1775,1796]" italics="true" pageId="3" pageNumber="566">tren</emphasis>
(stout) and the genus
<taxonomicName id="4C2B5CCC5831FF8EFD8293C6FD405062" authorityName="Fabricius" authorityYear="1775" box="[569,650,1775,1796]" class="Insecta" family="Rhagionidae" genus="Rhagio" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B95FFB5D5831FF8EFD8293C6FD405062" box="[569,650,1775,1796]" italics="true" pageId="3" pageNumber="566">Rhagio</emphasis>
</taxonomicName>
(rhagionid fly).
</paragraph>
<paragraph id="8B94274F5831FF8EFF1F9270FB965364" blockId="3.[164,670,1881,1902]" lastBlockId="3.[826,1441,943,1026]" pageId="3" pageNumber="566">
<emphasis id="B95FFB5D5831FF8EFF1F9270FDF75008" box="[164,573,1881,1902]" italics="true" pageId="3" pageNumber="566">
<typeStatus id="549099ED5831FF8EFF1F9270FF125008" box="[164,216,1881,1902]" pageId="3" pageNumber="566">Type</typeStatus>
species:
<taxonomicName id="4C2B5CCC5831FF8EFE849270FDF75008" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[319,573,1881,1902]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="deviatus" status="sp. nov.">Strenorhagio deviatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A26C46265831FF8EFDFC9270FD545008" box="[583,670,1881,1902]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95FFB5D5831FF8EFC819686FAEE54A2" box="[826,1316,943,965]" italics="true" pageId="3" pageNumber="566">
Included species:
<taxonomicName id="4C2B5CCC5831FF8EFBA09686FAEE54A2" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1051,1316,943,964]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="deviatus" status="sp. nov.">Strenorhagio deviatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A26C46265831FF8EFA839699FA5754A3" box="[1336,1437,944,965]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C2B5CCC5831FF8EFC8196E7FC085485" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[826,962,974,995]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="grimaldi" status="sp. nov.">
<emphasis id="B95FFB5D5831FF8EFC8196E7FC085485" box="[826,962,974,995]" italics="true" pageId="3" pageNumber="566">S. grimaldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFC7596E7FBE15485" box="[974,1067,974,995]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C2B5CCC5831FF8EFB8796E7FB375482" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1084,1277,974,996]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5831FF8EFB8796E7FB375482" box="[1084,1277,974,996]" italics="true" pageId="3" pageNumber="566">S. asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFAB296E7FAAC5485" box="[1289,1382,974,995]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C2B5CCC5831FF8EFC8196C4FC305364" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[826,1018,1005,1026]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="conjugovenius" status="sp. nov.">
<emphasis id="B95FFB5D5831FF8EFC8196C4FC305364" box="[826,1018,1005,1026]" italics="true" pageId="3" pageNumber="566">S. conjugovenius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFBBE96C4FB965364" box="[1029,1116,1005,1026]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B94274F5831FF8EFC819118FAF3527A" blockId="3.[826,1441,1072,1308]" pageId="3" pageNumber="566">
<emphasis id="B95FFB5D5831FF8EFC819118FC7B5320" box="[826,945,1073,1094]" italics="true" pageId="3" pageNumber="566">Diagnosis:</emphasis>
Body stout. Vein R
<subScript id="17AF250A5831FF8EFB7C9113FB155320" attach="left" box="[1223,1247,1082,1094]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
distinctly and strongly bent upwards at base; crossvein rm at basal third of cell d, proximal to or coalesced with beginning of vein R
<subScript id="17AF250A5831FF8EFB9B91BFFBF253C4" attach="left" box="[1056,1080,1174,1186]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
, beginning of vein R
<subScript id="17AF250A5831FF8EFA9491BFFAFD53C4" attach="left" box="[1327,1335,1174,1186]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
distal to apex of cell d; all branches of vein M long, veins M
<subScript id="17AF250A5831FF8EFA22919CFA6B53A7" attach="left" box="[1433,1441,1205,1217]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
and M
<subScript id="17AF250A5831FF8EFC3791FAFC5E53B9" attach="left" box="[908,916,1235,1247]" fontSize="5" pageId="3" pageNumber="566">2</subScript>
bifurcating distal to crossvein mm; vein CuP present; cell r
<subScript id="17AF250A5831FF8EFBA391DBFBEA5398" attach="left" box="[1048,1056,1266,1278]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
wider than in
<taxonomicName id="4C2B5CCC5831FF8EFB6E91C1FAAD539B" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1237,1383,1256,1277]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5831FF8EFB6E91C1FAAD539B" box="[1237,1383,1256,1277]" italics="true" pageId="3" pageNumber="566">Uranorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFAC991C1FCA0527A" pageId="3" pageNumber="566" rank="genus">gen. nov.</taxonomicNameLabel>
; cell d slender, far from wing margin.
</paragraph>
<paragraph id="8B94274F5831FF8EFC819063FA8F5219" blockId="3.[826,1441,1354,1407]" pageId="3" pageNumber="566">
<emphasis id="B95FFB5D5831FF8EFC819063FC065239" box="[826,972,1354,1375]" italics="true" pageId="3" pageNumber="566">Distribution:</emphasis>
The genus is known only from
<collectingCountry id="F33C67DF5831FF8EFAEE9063FA575206" box="[1365,1437,1354,1376]" name="China" pageId="3" pageNumber="566">China</collectingCountry>
, and is currently represented by four species.
</paragraph>
<paragraph id="8B94274F5831FF89FC819085FF3554C2" blockId="3.[826,1441,1452,1903]" lastBlockId="4.[144,759,849,932]" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566">
<emphasis id="B95FFB5D5831FF8EFC819085FC6C52A7" box="[826,934,1452,1473]" italics="true" pageId="3" pageNumber="566">Remarks:</emphasis>
We treat
<taxonomicName id="4C2B5CCC5831FF8EFB899085FB0C52A7" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1074,1222,1452,1473]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5831FF8EFB899085FB0C52A7" box="[1074,1222,1452,1473]" italics="true" pageId="3" pageNumber="566">Strenorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFB6D9084FA8952A4" box="[1238,1347,1453,1474]" pageId="3" pageNumber="566" rank="genus">gen. nov.</taxonomicNameLabel>
as the second genus of family
<taxonomicName id="4C2B5CCC5831FF8EFBEF90E2FAEE5287" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1108,1316,1483,1505]" class="Insecta" family="Uranorhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="family" status="fam. nov.">Uranorhagionidae</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFA9490E2FA575286" box="[1327,1437,1483,1505]" pageId="3" pageNumber="566" rank="family">fam. nov.</taxonomicNameLabel>
, because it has the following characters: body stout; hind tibia with a spur; veins R
<subScript id="17AF250A5831FF8EFB2E933BFB575178" attach="left" box="[1173,1181,1554,1566]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
and Rs
<subScript id="17AF250A5831FF8EFB48933BFB315178" attach="left" box="[1267,1275,1554,1566]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
long; vein R
<subScript id="17AF250A5831FF8EFA32933BFA6B5178" attach="left" box="[1417,1441,1554,1566]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
strongly bent upwards at base; veins M
<subScript id="17AF250A5831FF8EFA949318FAFD515B" attach="left" box="[1327,1335,1585,1597]" fontSize="5" pageId="3" pageNumber="566">1</subScript>
and M
<subScript id="17AF250A5831FF8EFA229318FA6B515B" attach="left" box="[1433,1441,1585,1597]" fontSize="5" pageId="3" pageNumber="566">2</subScript>
bifurcating distal to crossvein mm; five posterior cells present and wide open, etc. However, we can easily separate it from
<taxonomicName id="4C2B5CCC5831FF8EFBF793AAFB1451FE" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1100,1246,1667,1688]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5831FF8EFBF793AAFB1451FE" box="[1100,1246,1667,1688]" italics="true" pageId="3" pageNumber="566">Uranorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFB5393AAFA8451FE" box="[1256,1358,1667,1688]" pageId="3" pageNumber="566" rank="genus">gen. nov.</taxonomicNameLabel>
by the following features: vein R
<subScript id="17AF250A5831FF8EFBDB9382FBA251D1" attach="left" box="[1120,1128,1707,1719]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
beginning distal to apex of cell d; crossvein rm at basal third of cell d, proximal to or coalesced with beginning of vein R
<subScript id="17AF250A5831FF8EFAE993C1FAA05192" attach="left" box="[1362,1386,1768,1780]" fontSize="5" pageId="3" pageNumber="566">2+3</subScript>
; all branches of vein M long; cell r
<subScript id="17AF250A5831FF8EFB16922EFB7F5075" attach="left" box="[1197,1205,1799,1811]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
wider than that in
<taxonomicName id="4C2B5CCC5831FF8EFC819235FC065057" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[826,972,1820,1841]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5831FF8EFC819235FC065057" box="[826,972,1820,1841]" italics="true" pageId="3" pageNumber="566">Uranorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFC6C9235FB8B5057" box="[983,1089,1820,1841]" pageId="3" pageNumber="566" rank="genus">gen. nov.</taxonomicNameLabel>
; cell d slender, far from wing margin. In
<taxonomicName id="4C2B5CCC5831FF8EFC709213FB975029" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[971,1117,1850,1871]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FFB5D5831FF8EFC709213FB975029" box="[971,1117,1850,1871]" italics="true" pageId="3" pageNumber="566">Uranorhagio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265831FF8EFBD69212FB165036" box="[1133,1244,1851,1872]" pageId="3" pageNumber="566" rank="genus">gen. nov.</taxonomicNameLabel>
, vein R
<subScript id="17AF250A5831FF8EFAFB926DFA825036" attach="left" box="[1344,1352,1860,1872]" fontSize="5" pageId="3" pageNumber="566">4</subScript>
begins proximal to the apex of cell d, crossvein rm is located at the basal 3/7 of cell d, distal to the beginning of vein R
<subScript id="17AF250A5836FF89FF659653FF3C54E0" attach="left" box="[222,246,890,902]" fontSize="5" pageId="4" pageNumber="567">2+3</subScript>
; at least vein M
<subScript id="17AF250A5836FF89FE7F9653FE0654E0" attach="left" box="[452,460,890,902]" fontSize="5" pageId="4" pageNumber="567">3</subScript>
is short; cell r
<subScript id="17AF250A5836FF89FD3F9653FD4654E0" attach="left" box="[644,652,890,902]" fontSize="5" pageId="4" pageNumber="567">4</subScript>
is much narrower.
</paragraph>
</subSubSection>
</treatment>
</document>