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<mods:title id="801CEC7C565A26538970E63E7775CBE6">Redescription of Ve r t i g o (Vertigo) nitidula (Mousson, 1876) (Gastropoda: Vertiginidae) from the Caucasus region</mods:title>
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<emphasis id="B9C1EADAF817FFFB8584B725FF24F9A4" bold="true" box="[151,237,1615,1641]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
(
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<emphasis id="B9C1EADAF817FFFB8476B725FE0DF9A4" bold="true" box="[357,452,1615,1641]" italics="true" pageId="0" pageNumber="75">nitidula</emphasis>
(
<bibRefCitation id="EF244B39F817FFFB84C0B724FD59F9A5" author="Mousson" box="[467,656,1614,1640]" pageId="0" pageNumber="75" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
)
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<emphasis id="B9C1EADAF817FFFB8584B7FDFF05F961" box="[151,204,1687,1708]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
(
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<emphasis id="B9C1EADAF817FFFB85C8B7FDFEEDF961" box="[219,292,1687,1708]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
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)
<emphasis id="B9C1EADAF817FFFB8420B7FDFE47F961" box="[307,398,1687,1708]" italics="true" pageId="0" pageNumber="75">pygmaea</emphasis>
<taxonomicName id="4CB54D4BF817FFFB8487B7FDFD2EF966" authority="Mousson, 1876: 143" authorityName="Mousson" authorityPageNumber="143" authorityYear="1876" box="[404,743,1686,1708]" kingdom="Animalia" pageId="0" pageNumber="75" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF817FFFB84D3B7FCFDC6F966" box="[448,527,1686,1707]" italics="true" pageId="0" pageNumber="75">nitidula</emphasis>
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: 143
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.
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(
<emphasis id="B9C1EADAF817FFFB85C8B7DFFEEDF907" box="[219,292,1717,1738]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
)
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: 390
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, 407, pl. 10 fig. 6;
<treatmentCitation id="0A1410D9F817FFFB8602B7DFFC1EF907" author="Boettger" box="[785,983,1717,1738]" page="184" pageId="0" pageNumber="75" year="1883">
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: 184
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.
<emphasis id="B9C1EADAF817FFFB8584B7BEFF05F924" box="[151,204,1748,1769]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
<taxonomicName id="4CB54D4BF817FFFB85C0B7B9FE49F924" ID-CoL="7TLMG" box="[211,384,1747,1769]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF817FFFB85C8B7BEFEEDF924" box="[219,292,1748,1769]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
)
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var.
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: 407
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, pl. 10 fig. 7;
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: 141
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.
<emphasis id="B9C1EADAF817FFFB8584B799FF05F8C5" box="[151,204,1779,1800]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
<taxonomicName id="4CB54D4BF817FFFB85C0B798FE49F8C5" ID-CoL="7TLMG" box="[211,384,1778,1800]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF817FFFB85C8B799FEEDF8C5" box="[219,292,1779,1800]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF817FFFB8420B799FE49F8C5" box="[307,384,1779,1800]" italics="true" pageId="0" pageNumber="75">sieversi</emphasis>
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var.
<emphasis id="B9C1EADAF817FFFB84A1B798FDD4F8CA" box="[434,541,1778,1799]" italics="true" pageId="0" pageNumber="75">punctulum</emphasis>
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[
<emphasis id="B9C1EADAF817FFFB873FB799FD81F8C5" box="[556,584,1779,1800]" italics="true" pageId="0" pageNumber="75">sic</emphasis>
]:
<treatmentCitation id="0A1410D9F817FFFB874DB798FCEDF8CA" author="Boettger" box="[606,804,1778,1799]" page="407" pageId="0" pageNumber="75" year="1879">
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: 407
</treatmentCitation>
.
<emphasis id="B9C1EADAF817FFFB8584B67BFF05F8EB" box="[151,204,1809,1830]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
<taxonomicName id="4CB54D4BF817FFFB85C0B67BFE51F8EB" ID-CoL="5B9C8" box="[211,408,1809,1830]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="substriata" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF817FFFB85C8B67BFEEDF8EB" box="[219,292,1809,1830]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF817FFFB8420B67BFE51F8EB" box="[307,408,1809,1830]" italics="true" pageId="0" pageNumber="75">substriata</emphasis>
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var.
<emphasis id="B9C1EADAF817FFFB84D9B67BFE32F8EB" box="[458,507,1809,1830]" italics="true" pageId="0" pageNumber="75">mitis</emphasis>
<treatmentCitation id="0A1410D9F817FFFB8711B67BFD04F8EB" author="Boettger" box="[514,717,1809,1830]" page="140" pageId="0" pageNumber="75" year="1880">
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: 140
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, pl. 4 fig. 7.
<emphasis id="B9C1EADAF817FFFB8584B65AFF05F888" box="[151,204,1840,1861]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
<taxonomicName id="4CB54D4BF817FFFB85C0B645FE49F888" ID-CoL="7TLMG" box="[211,384,1839,1861]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF817FFFB85C8B65AFEEDF888" box="[219,292,1840,1861]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF817FFFB8420B65AFE49F888" box="[307,384,1840,1861]" italics="true" pageId="0" pageNumber="75">sieversi</emphasis>
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var.
<emphasis id="B9C1EADAF817FFFB84A1B645FDE5F889" box="[434,556,1839,1860]" italics="true" pageId="0" pageNumber="75">subalpestris</emphasis>
<treatmentCitation id="0A1410D9F817FFFB8720B65AFD37F889" author="Boettger" box="[563,766,1839,1861]" page="141" pageId="0" pageNumber="75" year="1880">
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: 141
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.
<taxonomicName id="4CB54D4BF817FFFB8584B625FE4CF8A9" ID-CoL="7TLMG" box="[151,389,1870,1892]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi" subGenus="Alaea">
<emphasis id="B9C1EADAF817FFFB8584B625FF29F8A9" box="[151,224,1871,1892]" italics="true" pageId="0" pageNumber="75">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF817FFFB85FCB624FEE3F8AE" box="[239,298,1870,1891]" italics="true" pageId="0" pageNumber="75">Alaea</emphasis>
)
<emphasis id="B9C1EADAF817FFFB842BB625FE4CF8A9" box="[312,389,1871,1892]" italics="true" pageId="0" pageNumber="75">sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF817FFFB849FB624FD27F8AE" authority="Boettger 1886a: 309" authorityName="Boettger" authorityPageNumber="309" authorityYear="1886" box="[396,750,1870,1892]" kingdom="Animalia" pageId="0" pageNumber="75" rank="variety" variety="punctum">
var.
<emphasis id="B9C1EADAF817FFFB84ABB625FDC6F8A9" box="[440,527,1871,1892]" italics="true" pageId="0" pageNumber="75">punctum</emphasis>
:
<treatmentCitation id="0A1410D9F817FFFB870FB624FD27F8AE" author="Boettger" box="[540,750,1870,1892]" page="309" pageId="0" pageNumber="75" year="1886">
<bibRefCitation id="EF244B39F817FFFB870FB624FD75F8A9" author="Boettger" box="[540,700,1870,1892]" pageId="0" pageNumber="75" refString="Boettger, O. (1886 a) Die Binnenmollusken des Talysch-Gebietes. Nach neuesten Materialien bearbeitet. In: Radde, G. (Ed.), Die Fauna und Flora des sudwestlichen Caspi-Gebietes. Wissenschaftliche Beitrage zu den Reisen an der persischrussischen Grenze. Brockhaus, Leipzig, pp. 255 - 350, pls. 2 - 3. http: // dx. doi. org / 10.5962 / bhl. title. 36963" type="book chapter" year="1886" yearSuffix="a">Boettger 1886a</bibRefCitation>
: 309
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F817FFFB8584B607FD68F84E" blockId="0.[151,1074,1686,2014]" box="[151,673,1901,1923]" pageId="0" pageNumber="75">
<treatmentCitationGroup id="ABA511E6F817FFFB8584B607FD68F84E" box="[151,673,1901,1923]" pageId="0" pageNumber="75">
<taxonomicName id="4CB54D4BF817FFFB8584B607FEFAF84F" ID-CoL="7TLMG" box="[151,307,1901,1922]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF817FFFB8584B607FEFAF84F" box="[151,307,1901,1922]" italics="true" pageId="0" pageNumber="75">Vertigo sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF817FFFB8429B607FD54F84E" authority="Boettger 1886b: 151" authorityName="Boettger" authorityPageNumber="151" authorityYear="1886" box="[314,669,1901,1923]" kingdom="Animalia" pageId="0" pageNumber="75" rank="variety" variety="punctum">
var.
<emphasis id="B9C1EADAF817FFFB8475B607FE74F84F" box="[358,445,1901,1922]" italics="true" pageId="0" pageNumber="75">punctum</emphasis>
:
<treatmentCitation id="0A1410D9F817FFFB84D9B607FD54F84E" author="Boettger" box="[458,669,1901,1923]" page="151" pageId="0" pageNumber="75" year="1886">
<bibRefCitation id="EF244B39F817FFFB84D9B607FDA3F84E" author="Boettger" box="[458,618,1901,1923]" pageId="0" pageNumber="75" refString="Boettger, O. (1886 b) Neuntes Verzeichniss (IX) von Mollusken der Kaukasuslander nach Sendungen des Hrn. Hans Leder, z. Z. in Helenendorf bei Elisabetpol (Transkaukasien). Jahrbucher der deutschen malakozoologischen Gesellschaft, 13, 121 - 156, pl. 3." type="journal article" year="1886" yearSuffix="b">Boettger 1886b</bibRefCitation>
: 151
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F817FFFB8584B6E6FE29F80D" blockId="0.[151,1074,1686,2014]" pageId="0" pageNumber="75">
<treatmentCitationGroup id="ABA511E6F817FFFB8584B6E6FE29F80D" pageId="0" pageNumber="75">
<taxonomicName id="4CB54D4BF817FFFB8584B6E6FEFAF86C" ID-CoL="7TLMG" box="[151,307,1932,1953]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF817FFFB8584B6E6FEFAF86C" box="[151,307,1932,1953]" italics="true" pageId="0" pageNumber="75">Vertigo sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF817FFFB8429B6E6FD6EF86C" authority="Boettger 1889: 25" authorityName="Boettger" authorityPageNumber="25" authorityYear="1889" box="[314,679,1931,1953]" kingdom="Animalia" pageId="0" pageNumber="75" rank="variety" variety="subalpestris">
var.
<emphasis id="B9C1EADAF817FFFB8476B6E1FE16F86D" box="[357,479,1931,1952]" italics="true" pageId="0" pageNumber="75">subalpestris</emphasis>
:
<treatmentCitation id="0A1410D9F817FFFB84FEB6E6FD6EF86C" author="Boettger" box="[493,679,1931,1953]" page="25" pageId="0" pageNumber="75" year="1889">
<bibRefCitation id="EF244B39F817FFFB84FEB6E6FD4BF86D" author="Boettger" box="[493,642,1931,1953]" pageId="0" pageNumber="75" refString="Boettger, O. (1889) Zehntes Verzeichnis (XII) von Mollusken der Kaukasuslander. Berichte der senckenbergischen naturforschenden Gesellschaft, 1888 / 1889, 3 - 37, pl. 1." type="journal article" year="1889">Boettger 1889</bibRefCitation>
: 25
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A1410D9F817FFFB87A2B6E1FCA1F86C" author="Pilsbry" box="[689,872,1931,1953]" page="196" pageId="0" pageNumber="75" year="1919">
<bibRefCitation id="EF244B39F817FFFB87A2B6E1FCFCF86D" author="Pilsbry" box="[689,821,1931,1953]" pageId="0" pageNumber="75" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry 1919</bibRefCitation>
: 196
</treatmentCitation>
.
<emphasis id="B9C1EADAF817FFFB8584B6C1FEE9F872" box="[151,288,1962,1984]" italics="true" pageId="0" pageNumber="75">
Pupa
<taxonomicName id="4CB54D4BF817FFFB85C0B6C0FEE9F872" ID-CoL="7TLMG" authority="Rosen 1907: 205" authorityName="Rosen" authorityPageNumber="205" authorityYear="1907" box="[211,288,1962,1983]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
:
<treatmentCitation id="0A1410D9F817FFFB843EB6C0FE15F80D" author="Rosen" box="[301,476,1962,1984]" page="205" pageId="0" pageNumber="75" year="1907">
<bibRefCitation id="EF244B39F817FFFB843EB6C0FE60F872" author="Rosen" box="[301,425,1962,1983]" pageId="0" pageNumber="75" refString="Rosen, O. von (1907) Zum Polymorphismus von Helix vulgaris Rossm. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft, 39, 198 - 206." type="journal article" year="1907">Rosen 1907</bibRefCitation>
: 205
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F817FFFB8584B6A3FDF8F813" blockId="0.[151,1074,1686,2014]" box="[151,561,1993,2014]" pageId="0" pageNumber="75">
<treatmentCitationGroup id="ABA511E6F817FFFB8584B6A3FDF8F813" box="[151,561,1993,2014]" pageId="0" pageNumber="75">
<emphasis id="B9C1EADAF817FFFB8584B6A3FF05F813" box="[151,204,1993,2014]" italics="true" pageId="0" pageNumber="75">Pupa</emphasis>
<taxonomicName id="4CB54D4BF817FFFB85C0B6A3FDE4F813" ID-CoL="7TLMG" authority="Rosen 1911: 121" authorityName="Rosen" authorityPageNumber="121" authorityYear="1911" box="[211,557,1993,2014]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="75" phylum="Mollusca" rank="species" species="sieversi" subGenus="Alaea">
(
<emphasis id="B9C1EADAF817FFFB85C8B6A3FEDFF813" box="[219,278,1993,2014]" italics="true" pageId="0" pageNumber="75">Alaea</emphasis>
)
<emphasis id="B9C1EADAF817FFFB8436B6A3FEBBF813" box="[293,370,1993,2014]" italics="true" pageId="0" pageNumber="75">sieversi</emphasis>
:
<treatmentCitation id="0A1410D9F817FFFB846CB6A3FDE4F813" author="Rosen" box="[383,557,1993,2014]" page="121" pageId="0" pageNumber="75" year="1911">
<bibRefCitation id="EF244B39F817FFFB846CB6A3FE33F813" author="Rosen" box="[383,506,1993,2014]" pageId="0" pageNumber="75" refString="Rosen, O. von (1911) Die Mollusken Ciskaukasiens und speciell des Kuban-Gebietes. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. - Petersbourg, 16, 86 - 142, pls. 2 - 3." type="journal article" year="1911">Rosen 1911</bibRefCitation>
: 121
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B1F3FD30FF63" blockId="1.[151,1436,152,603]" box="[151,761,152,174]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B1F3FD30FF63" box="[151,761,152,174]" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B1F3FE85FF63" ID-CoL="5B9C8" box="[151,332,152,174]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA8584B1F3FE85FF63" box="[151,332,152,174]" italics="true" pageId="1" pageNumber="76">Vertigo substriata</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF816FFFA8441B1F3FDBBFF63" authority="Pilsbry 1919: 173" authorityName="Pilsbry" authorityPageNumber="173" authorityYear="1919" box="[338,626,152,174]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF816FFFA846EB1F3FE67FF63" box="[381,430,153,174]" italics="true" pageId="1" pageNumber="76">mitis</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA84AFB1F2FDBBFF63" author="Pilsbry" box="[444,626,152,174]" page="173" pageId="1" pageNumber="76" year="1919">
<bibRefCitation id="EF244B39F816FFFA84AFB1F2FD88FF63" author="Pilsbry" box="[444,577,152,174]" pageId="1" pageNumber="76" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry 1919</bibRefCitation>
: 173
</treatmentCitation>
</taxonomicName>
, pl. 17 fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B1D2FCA9FF26" blockId="1.[151,1436,152,603]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B1D2FCA9FF26" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B1D2FE3EFF01" ID-CoL="7TLMG" authority="Pilsbry 1919: 195" authorityName="Pilsbry" authorityPageNumber="195" authorityYear="1919" box="[151,503,182,205]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF816FFFA8584B1D2FEFAFF00" box="[151,307,184,205]" italics="true" pageId="1" pageNumber="76">Vertigo sieversi</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA8452B1DCFE3EFF01" author="Pilsbry" box="[321,503,182,204]" page="195" pageId="1" pageNumber="76" year="1919">
<bibRefCitation id="EF244B39F816FFFA8452B1DCFE0CFF01" author="Pilsbry" box="[321,453,182,204]" pageId="1" pageNumber="76" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry 1919</bibRefCitation>
: 195
</treatmentCitation>
</taxonomicName>
, pl. 17 fig. 1112;
<treatmentCitation id="0A1410D9F816FFFA87A5B1DCFC3CFF01" author="Sysoev" box="[694,1013,182,204]" page="241" pageId="1" pageNumber="76" year="2005">
<bibRefCitation id="EF244B39F816FFFA87A5B1DCFC0BFF01" author="Sysoev" box="[694,962,182,204]" pageId="1" pageNumber="76" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2005) Neformalnaya gruppa Stylommatophora. In: Kantor, Y. I. &amp; Sysoev, A. V. (Eds.), Katalog Mollyuskov Rossii i sopredelnykh stran. KMK Scientific Press, Moscow, pp. 228 - 308." type="book chapter" year="2005">Sysoev &amp; Schileyko 2005</bibRefCitation>
: 241
</treatmentCitation>
;
<bibRefCitation id="EF244B39F816FFFA86ECB1DCFB16FF01" author="Pokryszko" box="[1023,1247,182,205]" pageId="1" pageNumber="76" refString="Pokryszko, B. M., Cameron, R. A. D., Mumladze, L. &amp; Tarknishvili, D. (2011) Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society, 102, 239 - 250. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2010.01575. x" type="journal article" year="2011">
Pokryszko
<emphasis id="B9C1EADAF816FFFA8163B1D2FB6DFF01" box="[1136,1188,182,205]" italics="true" pageId="1" pageNumber="76">et al.</emphasis>
2011
</bibRefCitation>
: Appendix S2, S3;
<treatmentCitation id="0A1410D9F816FFFA85D4B1BFFE3EFF26" author="Sysoev" box="[199,503,213,235]" page="51" pageId="1" pageNumber="76" year="2009">
<bibRefCitation id="EF244B39F816FFFA85D4B1BFFE1BFF26" author="Sysoev" box="[199,466,213,235]" pageId="1" pageNumber="76" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2009) Land snails and slugs of Russia and adjacent countries. Pensoft, Sofia and Moscow, 312 pp., 142 pls." type="book" year="2009">Sysoev &amp; Schileyko 2009</bibRefCitation>
: 51
</treatmentCitation>
, fig. 20M; Mumladze 2000: 14, 16.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B19FFC93FEC7" blockId="1.[151,1436,152,603]" box="[151,858,244,266]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B19FFC93FEC7" box="[151,858,244,266]" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B19FFEFAFEC7" ID-CoL="7TLMG" box="[151,307,245,266]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF816FFFA8584B19FFEFAFEC7" box="[151,307,245,266]" italics="true" pageId="1" pageNumber="76">Vertigo sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF816FFFA8429B19FFE19FEC7" box="[314,464,244,266]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="punctulum">
var.
<emphasis id="B9C1EADAF816FFFA8476B19EFE19FEC7" box="[357,464,244,266]" italics="true" pageId="1" pageNumber="76">punctulum</emphasis>
</taxonomicName>
[
<emphasis id="B9C1EADAF816FFFA84CCB19FFE32FEC7" box="[479,507,245,266]" italics="true" pageId="1" pageNumber="76">sic</emphasis>
]:
<treatmentCitation id="0A1410D9F816FFFA8702B19EFD0EFEC7" author="Pilsbry" box="[529,711,244,266]" page="196" pageId="1" pageNumber="76" year="1919">
<bibRefCitation id="EF244B39F816FFFA8702B19EFD5CFEC7" author="Pilsbry" box="[529,661,244,266]" pageId="1" pageNumber="76" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry 1919</bibRefCitation>
: 196
</treatmentCitation>
, pl. 17 fig. 13.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B079FC52FEE5" blockId="1.[151,1436,152,603]" box="[151,923,274,296]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B079FC52FEE5" box="[151,923,274,296]" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B079FE65FEE5" ID-CoL="5B9C8" box="[151,428,274,296]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata" subGenus="Vertigo">
<emphasis id="B9C1EADAF816FFFA8584B079FF29FEE5" box="[151,224,275,296]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF816FFFA85FCB079FEF1FEE5" box="[239,312,275,296]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF816FFFA8455B078FE65FEE5" box="[326,428,274,296]" italics="true" pageId="1" pageNumber="76">substriata</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF816FFFA84A1B079FC5EFEE5" authority="Likharev &amp; Rammelmeier 1952: 135" authorityName="Likharev &amp; Rammelmeier" authorityPageNumber="135" authorityYear="1952" box="[434,919,274,296]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF816FFFA84CDB079FDC6FEE5" box="[478,527,275,296]" italics="true" pageId="1" pageNumber="76">mitis</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA870FB078FC5EFEE5" author="Likharev" box="[540,919,274,296]" page="135" pageId="1" pageNumber="76" year="1952">
<bibRefCitation id="EF244B39F816FFFA870FB078FCADFEE5" author="Likharev" box="[540,868,274,296]" pageId="1" pageNumber="76" refString="Likharev, I. M. &amp; Rammelmeier, E. S. (1952) Nazemnye mollyuski fauny SSSR. In: Opred. po faune SSSR. 43. Akademiya Nauk SSSR, Moskva, Leningrad, 512 pp. [English translation 1962 in Israel Program for Scientific Translations, Jerusalem]" type="book" year="1952">Likharev &amp; Rammelmeier 1952</bibRefCitation>
: 135
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B058FC7FFE8A" blockId="1.[151,1436,152,603]" box="[151,950,305,327]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B058FC7FFE8A" box="[151,950,305,327]" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B058FE65FE8A" ID-CoL="5B9C8" box="[151,428,305,327]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata" subGenus="Vertigo">
<emphasis id="B9C1EADAF816FFFA8584B058FF29FE8A" box="[151,224,306,327]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF816FFFA85FCB058FEF1FE8A" box="[239,312,306,327]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF816FFFA8455B05BFE65FE8A" box="[326,428,305,327]" italics="true" pageId="1" pageNumber="76">substriata</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF816FFFA84A1B058FC7BFE8A" ID-CoL="7TLMG" authority="Likharev &amp; Rammelmeier 1952: 135" authorityName="Likharev &amp; Rammelmeier" authorityPageNumber="135" authorityYear="1952" box="[434,946,305,327]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="variety" species="substriata" variety="sieversi">
var.
<emphasis id="B9C1EADAF816FFFA84CDB058FDE2FE8A" box="[478,555,306,327]" italics="true" pageId="1" pageNumber="76">sieversi</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA872BB05BFC7BFE8A" author="Likharev" box="[568,946,305,327]" page="135" pageId="1" pageNumber="76" year="1952">
<bibRefCitation id="EF244B39F816FFFA872BB05BFCB6FE8A" author="Likharev" box="[568,895,305,327]" pageId="1" pageNumber="76" refString="Likharev, I. M. &amp; Rammelmeier, E. S. (1952) Nazemnye mollyuski fauny SSSR. In: Opred. po faune SSSR. 43. Akademiya Nauk SSSR, Moskva, Leningrad, 512 pp. [English translation 1962 in Israel Program for Scientific Translations, Jerusalem]" type="book" year="1952">Likharev &amp; Rammelmeier 1952</bibRefCitation>
: 135
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B03BFE95FE0F" blockId="1.[151,1436,152,603]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B03BFE95FE0F" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B03BFD81FEAB" ID-CoL="5B9C8" authority="Lezhawa 1962: 328" authorityName="Lezhawa" authorityPageNumber="328" authorityYear="1962" box="[151,584,336,358]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA8584B03BFE9CFEAB" box="[151,341,336,358]" italics="true" pageId="1" pageNumber="76">Vertigo substriata</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA847FB03AFD81FEAB" author="Lezhawa" box="[364,584,336,358]" page="328" pageId="1" pageNumber="76" year="1962">
<bibRefCitation id="EF244B39F816FFFA847FB03AFDC7FEAB" author="Lezhawa" box="[364,526,336,358]" pageId="1" pageNumber="76" refString="Lezhawa, G. (1962) K izucheniyu nazemnoy malakofauny gornoy Tusheti. Soobshcheniya Akademii Nauk Gruzinskoy SSR, 29, 327 - 332." type="journal article" year="1962">Lezhawa 1962</bibRefCitation>
: 328
</treatmentCitation>
</taxonomicName>
[misidentified];
<treatmentCitation id="0A1410D9F816FFFA8616B03AFC0FFEAB" author="Lezhawa" box="[773,966,336,358]" page="9" pageId="1" pageNumber="76" year="1966">
<bibRefCitation id="EF244B39F816FFFA8616B03AFC6EFEAB" author="Lezhawa" box="[773,935,336,358]" pageId="1" pageNumber="76" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
: 9
</treatmentCitation>
[misidentified];
<treatmentCitation id="0A1410D9F816FFFA8197B03AFA52FEAB" author="Aliev" box="[1156,1435,336,358]" page="34" pageId="1" pageNumber="76" year="1972">
<bibRefCitation id="EF244B39F816FFFA8197B03AFAA6FEAB" author="Aliev" box="[1156,1391,336,358]" pageId="1" pageNumber="76" refString="Aliev, S. Yu. &amp; Aliev, G. G. (1972) Nazemnye mollyuski Malogo Kavkaza territorii Azerbaydzhanskoy SSR, ikh rol v rasprostranenii gelmintozov selskokhozyaystvennykh zhivotnykh. Uchenye zapiski Azerbaydzhanskogo gosudarstvennogo universiteta. Seriya biologicheskikh nauk, 3, 33 - 36." type="journal article" year="1972">Aliev &amp; Aliev 1972</bibRefCitation>
: 34
</treatmentCitation>
[misidentified];
<treatmentCitation id="0A1410D9F816FFFA8461B004FD89FE49" author="Javelidze" box="[370,576,366,388]" page="29" pageId="1" pageNumber="76" year="1972">
<bibRefCitation id="EF244B39F816FFFA8461B004FDDAFE49" author="Javelidze" box="[370,531,366,388]" pageId="1" pageNumber="76" refString="Javelidze, G. (1972) Sakartvelos khmeletis moluskebis sarkveve. Tbilisis Universitetis, Tbilisi, 142 pp." type="book" year="1972">Javelidze 1972</bibRefCitation>
: 29
</treatmentCitation>
, fig. 21 [misidentified];
<treatmentCitation id="0A1410D9F816FFFA8641B004FBD2FE49" author="Lezhawa" box="[850,1051,366,388]" page="24" pageId="1" pageNumber="76" year="1973">
<bibRefCitation id="EF244B39F816FFFA8641B004FC38FE49" author="Lezhawa" box="[850,1009,366,388]" pageId="1" pageNumber="76" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">Lezhawa 1973</bibRefCitation>
: 24
</treatmentCitation>
[misidentified];
<treatmentCitation id="0A1410D9F816FFFA81C0B004FA55FE49" author="Lezhawa" box="[1235,1436,366,388]" page="86" pageId="1" pageNumber="76" year="2000">
<bibRefCitation id="EF244B39F816FFFA81C0B004FABBFE49" author="Lezhawa" box="[1235,1394,366,388]" pageId="1" pageNumber="76" refString="Lezhawa, G. (2000) The land shelled molluscs of Georgia. Proceedings of the Institute of Zoology, Tbilisi, 20, 85 - 95." type="journal article" year="2000">Lezhawa 2000</bibRefCitation>
: 86
</treatmentCitation>
[misidentified];
<treatmentCitation id="0A1410D9F816FFFA847DB0E7FD7FFE6E" author="Sysoev" box="[366,694,397,419]" page="241" pageId="1" pageNumber="76" year="2005">
<bibRefCitation id="EF244B39F816FFFA847DB0E7FD4AFE6E" author="Sysoev" box="[366,643,397,419]" pageId="1" pageNumber="76" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2005) Neformalnaya gruppa Stylommatophora. In: Kantor, Y. I. &amp; Sysoev, A. V. (Eds.), Katalog Mollyuskov Rossii i sopredelnykh stran. KMK Scientific Press, Moscow, pp. 228 - 308." type="book chapter" year="2005">Sysoev &amp; Schileyko 2005</bibRefCitation>
: 241
</treatmentCitation>
[partly misidentified];
<treatmentCitation id="0A1410D9F816FFFA86BFB0E7FB20FE6E" author="Sysoev" box="[940,1257,397,419]" page="51" pageId="1" pageNumber="76" year="2009">
<bibRefCitation id="EF244B39F816FFFA86BFB0E7FB08FE6E" author="Sysoev" box="[940,1217,397,419]" pageId="1" pageNumber="76" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2009) Land snails and slugs of Russia and adjacent countries. Pensoft, Sofia and Moscow, 312 pp., 142 pls." type="book" year="2009">Sysoev &amp; Schileyko 2009</bibRefCitation>
: 51
</treatmentCitation>
, fig. 20N [partly misidentified].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B0A1FE3EFE32" blockId="1.[151,1436,152,603]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B0A1FE3EFE32" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B0A1FD57FE2D" ID-CoL="7TLMG" authority="Akramowski 1968: 26" authorityName="Akramowski" authorityPageNumber="26" authorityYear="1968" box="[151,670,458,480]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="subSpecies" species="substriata" subSpecies="sieversi">
<emphasis id="B9C1EADAF816FFFA8584B0A1FE6DFE2D" box="[151,420,458,480]" italics="true" pageId="1" pageNumber="76">Vertigo substriata sieversi</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA84A7B0A0FD57FE2D" author="Akramowski" box="[436,670,458,480]" page="26" pageId="1" pageNumber="76" year="1968">
<bibRefCitation id="EF244B39F816FFFA84A7B0A0FDBCFE2D" author="Akramowski" box="[436,629,458,480]" pageId="1" pageNumber="76" refString="Akramowski, N. N. (1968) Nekotorye dopolneniya k faune mollyuskov Armenii. Vklyuchaya novosti dlya fauny Zakavkazya i SSSR. In: Mollyuski i ikh pol v yekosistemakh. Avtoreferaty dokladov, Nauka, Leningrad, 150 pp. [pp. 26 - 27]" type="book chapter" year="1968">Akramowski 1968</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A1410D9F816FFFA87B8B0A0FC41FE2D" author="Akramowski" box="[683,904,458,480]" page="6" pageId="1" pageNumber="76" year="1971">
<bibRefCitation id="EF244B39F816FFFA87B8B0A0FCA4FE2D" author="Akramowski" box="[683,877,458,480]" pageId="1" pageNumber="76" refString="Akramowski, N. N. (1971) Kratkiy katalog sovremennoy fauny mollyuskov Sovetskoy Armenii. Biologicheskiy zhurnal Armenii, 24, 3 - 12." type="journal article" year="1971">Akramowski 1971</bibRefCitation>
: 6
</treatmentCitation>
;
<treatmentCitation id="0A1410D9F816FFFA8686B0A0FB5CFE2D" author="Asadov" box="[917,1173,458,480]" page="101" pageId="1" pageNumber="76" year="1975">
<bibRefCitation id="EF244B39F816FFFA8686B0A0FBA9FE2D" author="Asadov" box="[917,1120,458,480]" pageId="1" pageNumber="76" refString="Asadov, S. M., Akramowski, N. N. &amp; Dzhabbarov, D. G. (1975) K izucheniyu fauny nazemnykh mollyuskov Malogo Kavkaza v predelakh Azerbaydzhanskoy SSR. Izvestiya Akademii Nauk Azerbaydzhanskoy SSR, Seriya biologicheskick nauk, 1975, 100 - 106." type="journal article" year="1975">
Asadov
<emphasis id="B9C1EADAF816FFFA86FFB0A1FBE8FE2D" box="[1004,1057,458,480]" italics="true" pageId="1" pageNumber="76">et al.</emphasis>
1975
</bibRefCitation>
: 101
</treatmentCitation>
;
<treatmentCitation id="0A1410D9F816FFFA81B1B0A0FA51FE2D" author="Akramowski" box="[1186,1432,458,480]" page="136" pageId="1" pageNumber="76" year="1976">
<bibRefCitation id="EF244B39F816FFFA81B1B0A0FAAAFE2D" author="Akramowski" box="[1186,1379,458,480]" pageId="1" pageNumber="76" refString="Akramowski, N. N. (1976) Fauna Armyanskoy SSR. Mollyuski (Mollusca). Akademiya nauk Armyanskoy SSR, Institut Zoologii, Yerevan, 268 pp., 16 pls." type="book" year="1976">Akramowski 1976</bibRefCitation>
: 136
</treatmentCitation>
, text-fig. 56, 57B, pl. 5 fig. 48.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B363FEACFDF1" blockId="1.[151,1436,152,603]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B363FEACFDF1" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B363FD7AFDD3" ID-CoL="5B9C8" authority="Schileyko 1984: 201" authorityName="Schileyko" authorityPageNumber="201" authorityYear="1984" box="[151,691,520,542]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata" subGenus="Vertigo">
<emphasis id="B9C1EADAF816FFFA8584B363FF29FDD3" box="[151,224,521,542]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF816FFFA85E5B363FEF6FDD3" box="[246,319,521,542]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF816FFFA8445B362FE72FDD3" box="[342,443,520,542]" italics="true" pageId="1" pageNumber="76">substriata</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA84C3B362FD7AFDD3" author="Schileyko" box="[464,691,520,542]" page="201" pageId="1" pageNumber="76" year="1984">
<bibRefCitation id="EF244B39F816FFFA84C3B362FDB0FDD3" author="Schileyko" box="[464,633,520,542]" pageId="1" pageNumber="76" refString="Schileyko, A. A. (1984) Nazemnye mollyuski podotryada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In: Fauna SSSR. Mollyuski. III (3). Nauka, Leningrad, pp. 1 - 399." type="journal article" year="1984">Schileyko 1984</bibRefCitation>
: 201
</treatmentCitation>
</taxonomicName>
, fig. 118-119 [partly misidentified];
<treatmentCitation id="0A1410D9F816FFFA8159B362FADCFDD3" author="Kijashko" box="[1098,1301,520,542]" page="11" pageId="1" pageNumber="76" year="2000">
<bibRefCitation id="EF244B39F816FFFA8159B362FB23FDD3" author="Kijashko" box="[1098,1258,520,542]" pageId="1" pageNumber="76" refString="Kijashko, P. V. (2000) Nazemnye mollyuski Lagonakskogo nagorya (Severo-Zapadnyi Kavkaz). Rossiyskaya Akademiya Nauk, Zoologicheskiy Institut, Sankt-Peterburg, 24 pp." type="book" year="2000">Kijashko 2000</bibRefCitation>
: 11
</treatmentCitation>
, 12, 14, 21 [misidentified].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B32CFAABFD96" blockId="1.[151,1436,152,603]" box="[151,1378,581,603]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="ABA511E6F816FFFA8584B32CFAABFD96" box="[151,1378,581,603]" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA8584B32CFDBCFD96" ID-CoL="7TLMG" authority="Schileyko 1984: 204" authorityName="Schileyko" authorityPageNumber="204" authorityYear="1984" box="[151,629,581,603]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
<emphasis id="B9C1EADAF816FFFA8584B32CFF29FD96" box="[151,224,582,603]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF816FFFA85FCB32CFEF1FD96" box="[239,312,582,603]" italics="true" pageId="1" pageNumber="76">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF816FFFA8455B32CFE5AFD96" box="[326,403,582,603]" italics="true" pageId="1" pageNumber="76">sieversi</emphasis>
:
<treatmentCitation id="0A1410D9F816FFFA84B2B32FFDBCFD96" author="Schileyko" box="[417,629,581,603]" page="204" pageId="1" pageNumber="76" year="1984">
<bibRefCitation id="EF244B39F816FFFA84B2B32FFD8AFD96" author="Schileyko" box="[417,579,581,603]" pageId="1" pageNumber="76" refString="Schileyko, A. A. (1984) Nazemnye mollyuski podotryada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In: Fauna SSSR. Mollyuski. III (3). Nauka, Leningrad, pp. 1 - 399." type="journal article" year="1984">Schileyko 1984</bibRefCitation>
: 204
</treatmentCitation>
</taxonomicName>
, fig. 123;
<treatmentCitation id="0A1410D9F816FFFA87C8B32CFC4EFD96" author="Egorov" box="[731,903,581,603]" page="35" pageId="1" pageNumber="76" year="2008">
<bibRefCitation id="EF244B39F816FFFA87C8B32CFCABFD96" author="Egorov" box="[731,866,581,603]" pageId="1" pageNumber="76" refString="Egorov, R. V. (2008) Treasure of Russian Shells, Supplement 5. Illustrated catalogue of the recent terrestrial molluscs of Russia and adjacent regions. Colus-Doveri, Moscow, 179 pp." type="book" year="2008">Egorov 2008</bibRefCitation>
: 35
</treatmentCitation>
, pl. 5 fig. 32;
<treatmentCitation id="0A1410D9F816FFFA8106B32FFB0CFD96" author="Kijashko" box="[1045,1221,581,603]" page="9" pageId="1" pageNumber="76" year="2000">
<bibRefCitation id="EF244B39F816FFFA8106B32FFB67FD96" author="Kijashko" box="[1045,1198,581,603]" pageId="1" pageNumber="76" refString="Kijashko, P. V. (2000) Nazemnye mollyuski Lagonakskogo nagorya (Severo-Zapadnyi Kavkaz). Rossiyskaya Akademiya Nauk, Zoologicheskiy Institut, Sankt-Peterburg, 24 pp." type="book" year="2000">Kijashko 2000</bibRefCitation>
: 9
</treatmentCitation>
, 12, 14, 18, 21.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B0A36C8F816FFFA8584B3ECFE61FC72" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
<emphasis id="B9C1EADAF816FFFA8584B3ECFEB8FD52" bold="true" box="[151,369,646,671]" pageId="1" pageNumber="76">
Shell (
<figureCitation id="138E2A4DF816FFFA85F0B3ECFEEEFD52" box="[227,295,646,671]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
A
<collectingCountry id="F3A27658F816FFFA845FB3ECFEAAFD52" box="[332,355,646,671]" name="Iceland" pageId="1" pageNumber="76">O</collectingCountry>
).
</emphasis>
Shell elongate-ovate or rarely ovate with a rounded apex; perforate; upper whorls irregularly rib-striated; the striation becomes weaker on the body whorl; corneous or greenish-brown; subtranslucent; 4.5
<emphasis id="B9C1EADAF816FFFA8093B3C0FA47FD0E" bold="true" box="[1408,1422,682,707]" pageId="1" pageNumber="76"></emphasis>
5 moderately convex whorls; body whorl slightly ascending towards the aperture, with a shallow impression behind the weak cervical crest above the lower palatal fold; aperture rounded triangular; insertions of peristome connected by a weak callus; peristome slightly expanded, not reflected, only slightly thickened; palatal margin straight with only a shallow indentation; occasionally with a white callus at the inside corresponding to the cervical crest; parietal lamella higher than angular lamella, which is sometimes completely missing; columellar lamella strong; lower palatal fold stronger than upper palatal fold; basal fold more or less strong, missing in some specimens or even whole populations.
</paragraph>
</subSubSection>
<subSubSection id="C3AF6543F816FFF985D4B2A0FA4BFE45" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76" type="description">
<paragraph id="8B0A36C8F816FFFA85D4B2A0FBF9FBCB" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
<emphasis id="B9C1EADAF816FFFA85D4B2A0FE15FC2E" bold="true" box="[199,476,970,995]" pageId="1" pageNumber="76">
Measurements (
<figureCitation id="138E2A4DF816FFFA8495B2A0FE07FC2E" box="[390,462,970,995]" captionStart="FIGURE 2" captionStartId="3.[151,250,1258,1280]" captionTargetId="figure@3.[340,1248,191,1236]" captionTargetPageId="3" captionText="FIGURE 2. Variation in shell size and shape of Vertigo (Vertigo) nitidula (Mousson)." figureDoi="http://doi.org/10.5281/zenodo.8387550" httpUri="https://zenodo.org/record/8387550/files/figure.png" pageId="1" pageNumber="76">Fig. 2</figureCitation>
).
</emphasis>
<specimenCount id="9DB3FD41F816FFFA84F7B2A0FDB7FC2E" box="[484,638,970,995]" pageId="1" pageNumber="76" type="generic">57 specimens</specimenCount>
from 13 populations (n=1
<emphasis id="B9C1EADAF816FFFA86BAB2A0FC7EFC2E" bold="true" box="[937,951,970,995]" pageId="1" pageNumber="76"></emphasis>
18 per population): shell height: 1.502.20 mm, mean 1.85 + 0.18 mm; shell diameter: 0.901.15 mm, mean 1.03 + 0.06 mm.
</paragraph>
<paragraph id="8B0A36C8F816FFFA85D4B578FA9FFBBF" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
<emphasis id="B9C1EADAF816FFFA85D4B578FE77FBE6" bold="true" box="[199,446,1042,1067]" pageId="1" pageNumber="76">Species delimitation.</emphasis>
<taxonomicName id="4CB54D4BF816FFFA84D5B57EFDF3FBE6" box="[454,570,1042,1067]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA84D5B57EFDF3FBE6" box="[454,570,1042,1067]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4CB54D4BF816FFFA87C1B57EFBD0FBE6" authority="Jeffreys, 1833" authorityName="Jeffreys" authorityYear="1833" box="[722,1049,1042,1067]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA87C1B57EFC97FBE6" box="[722,862,1042,1067]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
(
<bibRefCitation id="EF244B39F816FFFA867DB578FBD8FBE6" author="Jeffreys" box="[878,1041,1042,1067]" pageId="1" pageNumber="76" refString="Jeffreys, J. G. (1833) A Supplement to the ' Synopsis of the Testaceous Pneumobranchous Mollusca of Great Britain'. Transactions of the Linnean Society of London, 16 (3), 505 - 523." type="journal article" year="1833">Jeffreys, 1833</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="138E2A4DF816FFFA8139B578FBA1FBE6" box="[1066,1128,1042,1067]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
PV), but differs from this species in the finer radial sculpture on the upper whorls, which are ribbed in
<taxonomicName id="4CB54D4BF816FFFA86E0B55DFBB7FB82" box="[1011,1150,1078,1103]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA86E0B55DFBB7FB82" box="[1011,1150,1078,1103]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
, and the only moderately convex whorls. Moreover, most specimens of
<taxonomicName id="4CB54D4BF816FFFA878FB536FCC7FBBE" box="[668,782,1114,1139]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA878FB536FCC7FBBE" box="[668,782,1114,1139]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
are more elongated than specimens of
<taxonomicName id="4CB54D4BF816FFFA81D7B536FA86FBBE" box="[1220,1359,1114,1139]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA81D7B536FA86FBBE" box="[1220,1359,1114,1139]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B0A36C8F816FFFA85D4B514FB47FAA3" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
The shell variability within
<taxonomicName id="4CB54D4BF816FFFA8717B515FDB0FB5A" box="[516,633,1150,1175]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8717B515FDB0FB5A" box="[516,633,1150,1175]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
is high. The varieties
<emphasis id="B9C1EADAF816FFFA8669B515FBDBFB5B" box="[890,1042,1151,1174]" italics="true" pageId="1" pageNumber="76">
Pupa
<taxonomicName id="4CB54D4BF816FFFA86AEB515FBDBFB5B" box="[957,1042,1151,1174]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF816FFFA810FB515FA94FB5A" authority="Boettger, 1879" authorityName="Boettger" authorityYear="1879" box="[1052,1373,1151,1175]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="punctum">
var.
<emphasis id="B9C1EADAF816FFFA815EB515FB67FB5B" box="[1101,1198,1151,1174]" italics="true" pageId="1" pageNumber="76">punctum</emphasis>
<bibRefCitation id="EF244B39F816FFFA81A4B515FA94FB5A" author="Boettger" box="[1207,1373,1151,1175]" pageId="1" pageNumber="76" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger, 1879</bibRefCitation>
</taxonomicName>
(
<figureCitation id="138E2A4DF816FFFA807DB514FF68FB76" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
C) and
<emphasis id="B9C1EADAF816FFFA85EAB5CEFE58FB76" box="[249,401,1188,1211]" italics="true" pageId="1" pageNumber="76">
Pupa
<taxonomicName id="4CB54D4BF816FFFA842FB5CEFE58FB76" box="[316,401,1188,1211]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF816FFFA8489B5C9FCC9FB76" authority="Boettger, 1880" authorityName="Boettger" authorityYear="1880" box="[410,768,1186,1211]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="subalpestris">
var.
<emphasis id="B9C1EADAF816FFFA84D8B5C8FD9BFB76" box="[459,594,1186,1211]" italics="true" pageId="1" pageNumber="76">subalpestris</emphasis>
<bibRefCitation id="EF244B39F816FFFA8749B5C9FCC9FB76" author="Boettger" box="[602,768,1187,1211]" pageId="1" pageNumber="76" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger, 1880</bibRefCitation>
</taxonomicName>
(
<figureCitation id="138E2A4DF816FFFA8603B5C8FC86FB76" box="[784,847,1186,1211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
D) were based on differences in shell shape and the development of the angular lamella. However, these as well as other variations of shell characters are not characteristic for specific geographic regions. Shell size and shape vary continuously (
<figureCitation id="138E2A4DF816FFFA8181B580FB15FACE" box="[1170,1244,1258,1283]" captionStart="FIGURE 2" captionStartId="3.[151,250,1258,1280]" captionTargetId="figure@3.[340,1248,191,1236]" captionTargetPageId="3" captionText="FIGURE 2. Variation in shell size and shape of Vertigo (Vertigo) nitidula (Mousson)." figureDoi="http://doi.org/10.5281/zenodo.8387550" httpUri="https://zenodo.org/record/8387550/files/figure.png" pageId="1" pageNumber="76">Fig. 2</figureCitation>
). Different shell shapes and the presence or absence of angular lamella and basal fold are found in various combinations and different forms can co-occur within a single population (see
<figureCitation id="138E2A4DF816FFFA8676B458FC6FFA86" box="[869,934,1330,1355]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
HJ and K
<emphasis id="B9C1EADAF816FFFA813FB458FBF3FA86" bold="true" box="[1068,1082,1330,1355]" pageId="1" pageNumber="76"></emphasis>
<collectingCountry id="F3A27658F816FFFA8129B459FB98FA86" box="[1082,1105,1331,1355]" name="Iceland" pageId="1" pageNumber="76">O</collectingCountry>
). Thus, we cannot split this complex into smaller entities and place the named varieties in the synonymy of
<taxonomicName id="4CB54D4BF816FFFA8107B43DFB4EFAA2" box="[1044,1159,1366,1391]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8107B43DFB4EFAA2" box="[1044,1159,1366,1391]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B0A36C8F816FFFA85D4B411FC54F9A7" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
<bibRefCitation id="EF244B39F816FFFA85D4B411FEB2FA5E" author="Boettger" box="[199,379,1403,1427]" pageId="1" pageNumber="76" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger (1880)</bibRefCitation>
described a short
<emphasis id="B9C1EADAF816FFFA8743B416FD57FA5E" box="[592,670,1404,1427]" italics="true" pageId="1" pageNumber="76">Vert igo</emphasis>
form from Abastumani and Kazbek as
<emphasis id="B9C1EADAF816FFFA8178B416FB30FA5E" box="[1131,1273,1402,1427]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA819AB410FB30FA5E" box="[1161,1273,1402,1427]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">substriata</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF816FFFA8011B411FAA3FA5E" box="[1282,1386,1403,1427]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF816FFFA8027B416FAA3FA5E" box="[1332,1386,1404,1427]" italics="true" pageId="1" pageNumber="76">mitis</emphasis>
</taxonomicName>
. He stated that it differs from
<emphasis id="B9C1EADAF816FFFA84AAB4F5FDE3FA7B" box="[441,554,1439,1462]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA84C6B4F5FDE3FA7B" box="[469,554,1439,1462]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
(=
<taxonomicName id="4CB54D4BF816FFFA8747B4F5FD01FA7A" box="[596,712,1438,1463]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8747B4F5FD01FA7A" box="[596,712,1438,1463]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
) in the more strongly sculptured, less turrited shell. However, a re-examination of the
<typeStatus id="540E886AF816FFFA8489B4A9FE35FA16" box="[410,508,1475,1499]" pageId="1" pageNumber="76" type="syntype">syntypes</typeStatus>
(
<figureCitation id="138E2A4DF816FFFA871CB4A8FD98FA16" box="[527,593,1474,1499]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
E) has shown that the upper whorls of this form are not ribbed as in
<taxonomicName id="4CB54D4BF816FFFA8094B4AEFECEFA32" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA8094B4AEFECEFA32" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
, but coincide in sculpture with those of
<taxonomicName id="4CB54D4BF816FFFA87D7B48DFCFFFA32" box="[708,822,1510,1535]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA87D7B48DFCFFFA32" box="[708,822,1510,1535]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
. The three available
<typeStatus id="540E886AF816FFFA810CB48DFB48FA32" box="[1055,1153,1511,1535]" pageId="1" pageNumber="76" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF816FFFA81B7B48DFAE6FA32" box="[1188,1327,1510,1535]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA81ACB48CFAE6FA32" box="[1215,1327,1510,1535]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">substriata</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF816FFFA8025B48DFA52FA33" box="[1334,1435,1511,1534]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF816FFFA8076B48DFA52FA33" box="[1381,1435,1511,1534]" italics="true" pageId="1" pageNumber="76">mitis</emphasis>
</taxonomicName>
from Abastumani differ from the three specimens from the same locality determined as
<emphasis id="B9C1EADAF816FFFA8184B766FAC2F9EE" box="[1175,1291,1548,1571]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA81A5B766FAC2F9EE" box="[1206,1291,1548,1571]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
by
<bibRefCitation id="EF244B39F816FFFA802FB761FF28F98A" author="Boettger" pageId="1" pageNumber="76" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger (1880)</bibRefCitation>
in the smaller shell height and the reduction of the angular lamella, but not in shell sculpture. Thus, we place
<emphasis id="B9C1EADAF816FFFA8584B73EFEEBF9A6" box="[151,290,1618,1643]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA85A1B738FEEBF9A6" box="[178,290,1618,1643]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">substriata</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF816FFFA843AB739FDF5F9A6" authority="Boettger, 1880" authorityName="Boettger" authorityYear="1880" box="[297,572,1619,1643]" kingdom="Animalia" pageId="1" pageNumber="76" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF816FFFA8449B73EFE59F9A6" box="[346,400,1620,1643]" italics="true" pageId="1" pageNumber="76">mitis</emphasis>
<bibRefCitation id="EF244B39F816FFFA8484B739FDF5F9A6" author="Boettger" box="[407,572,1619,1643]" pageId="1" pageNumber="76" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">
Boettger,
<quantity id="4C4D9B2DF816FFFA8717B739FDF5F9A6" box="[516,572,1619,1643]" metricMagnitude="1" metricUnit="m" metricValue="4.7752" pageId="1" pageNumber="76" unit="in" value="1880.0">1880</quantity>
</bibRefCitation>
</taxonomicName>
in the synonymy of
<taxonomicName id="4CB54D4BF816FFFA8630B73EFC5FF9A6" box="[803,918,1618,1643]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8630B73EFC5FF9A6" box="[803,918,1618,1643]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B0A36C8F816FFFA85D4B71CFF26F8D2" blockId="1.[151,1437,646,2003]" pageId="1" pageNumber="76">
<bibRefCitation id="EF244B39F816FFFA85D4B71CFE43F942" author="Schileyko" box="[199,394,1654,1679]" pageId="1" pageNumber="76" refString="Schileyko, A. A. (1984) Nazemnye mollyuski podotryada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In: Fauna SSSR. Mollyuski. III (3). Nauka, Leningrad, pp. 1 - 399." type="journal article" year="1984">Schileyko (1984)</bibRefCitation>
considered the lack of the basal fold as the decisive diagnostic feature for distinguishing
<taxonomicName id="4CB54D4BF816FFFA8094B71DFF27F97E" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8094B71DFF27F97E" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
from
<taxonomicName id="4CB54D4BF816FFFA8427B7F6FE09F97E" box="[308,448,1690,1715]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA8427B7F6FE09F97E" box="[308,448,1690,1715]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
. However, the basal fold is present in the
<typeStatus id="540E886AF816FFFA868EB7F1FC36F97E" box="[925,1023,1691,1715]" pageId="1" pageNumber="76" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4CB54D4BF816FFFA8136B7F6FB51F97E" box="[1061,1176,1690,1715]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8136B7F6FB51F97E" box="[1061,1176,1690,1715]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
(
<figureCitation id="138E2A4DF816FFFA81BBB7F0FB2EF97E" box="[1192,1255,1690,1715]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
A) as well as in the
<typeStatus id="540E886AF816FFFA85D1B7D5FEEDF91A" box="[194,292,1727,1751]" pageId="1" pageNumber="76" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF816FFFA845EB7D5FE09F91B" box="[333,448,1727,1750]" italics="true" pageId="1" pageNumber="76">
P.
<taxonomicName id="4CB54D4BF816FFFA8478B7D5FE09F91B" box="[363,448,1727,1750]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
(
<figureCitation id="138E2A4DF816FFFA84C1B7D4FDDAF91A" box="[466,531,1726,1751]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
B).
<collectingCountry id="F3A27658F816FFFA8750B7D5FDABF91A" box="[579,610,1727,1751]" name="American Samoa" pageId="1" pageNumber="76">As</collectingCountry>
already noted by
<bibRefCitation id="EF244B39F816FFFA8620B7D5FC20F91A" author="Boettger" box="[819,1001,1727,1751]" pageId="1" pageNumber="76" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger (1880)</bibRefCitation>
, a basal fold may be missing in some populations of
<taxonomicName id="4CB54D4BF816FFFA842EB78EFE66F936" box="[317,431,1762,1787]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA842EB78EFE66F936" box="[317,431,1762,1787]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
. The same is true for
<taxonomicName id="4CB54D4BF816FFFA87B3B78EFCE2F936" box="[672,811,1762,1787]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA87B3B78EFCE2F936" box="[672,811,1762,1787]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
. Thus, this character cannot be used to distinguish these species.
</paragraph>
<paragraph id="8B0A36C8F816FFF985D4B646FB6DFF18" blockId="1.[151,1437,646,2003]" lastBlockId="2.[151,1437,151,752]" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
<taxonomicName id="4CB54D4BF816FFFA85D4B646FE50F88E" box="[199,409,1834,1859]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA85D4B646FE50F88E" box="[199,409,1834,1859]" italics="true" pageId="1" pageNumber="76">Vertigo substriata</emphasis>
</taxonomicName>
has been reported repeatedly from the Caucasus region. However, most vouchers determined as
<taxonomicName id="4CB54D4BF816FFFA842FB625FE00F8AA" box="[316,457,1870,1895]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA842FB625FE00F8AA" box="[316,457,1870,1895]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
turned out to belong to
<taxonomicName id="4CB54D4BF816FFFA87C9B625FC87F8AA" box="[730,846,1870,1895]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA87C9B625FC87F8AA" box="[730,846,1870,1895]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
. Obviously, they were misidentified as
<taxonomicName id="4CB54D4BF816FFFA801CB625FA55F8AA" box="[1295,1436,1870,1895]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA801CB625FA55F8AA" box="[1295,1436,1870,1895]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
because of the unclear delimitation of
<taxonomicName id="4CB54D4BF816FFFA8754B61EFD72F846" box="[583,699,1906,1931]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF816FFFA8754B61EFD72F846" box="[583,699,1906,1931]" italics="true" pageId="1" pageNumber="76">V. nitidula</emphasis>
</taxonomicName>
and
<taxonomicName id="4CB54D4BF816FFFA87E1B61EFCB7F846" box="[754,894,1906,1931]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="76" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF816FFFA87E1B61EFCB7F846" box="[754,894,1906,1931]" italics="true" pageId="1" pageNumber="76">V. substriata</emphasis>
</taxonomicName>
. Only three samples from the Chegem valley in
<collectingCountry id="F3A27658F816FFFA8584B6FCFF2BF862" box="[151,226,1942,1967]" name="Russia" pageId="1" pageNumber="76">Russia</collectingCountry>
(ZIN 96/2,
<figureCitation id="138E2A4DF816FFFA847AB6FCFE61F862" box="[361,424,1942,1967]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
S), from Kobi in
<collectingCountry id="F3A27658F816FFFA877DB6FCFD02F862" box="[622,715,1942,1967]" name="Georgia" pageId="1" pageNumber="76">Georgia</collectingCountry>
,
<geoCoordinate id="EE81500FF816FFFA87C5B6FDFCFBF862" box="[726,818,1943,1967]" direction="north" orientation="latitude" pageId="1" pageNumber="76" precision="925" value="42.55">42°33'N</geoCoordinate>
<geoCoordinate id="EE81500FF816FFFA8629B6FDFC5FF862" box="[826,918,1943,1967]" direction="east" orientation="longitude" pageId="1" pageNumber="76" precision="925" value="44.5">44°30'E</geoCoordinate>
, (ZIN 95/30,
<figureCitation id="138E2A4DF816FFFA8123B6FCFBA7F862" box="[1072,1134,1942,1967]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="1" pageNumber="76">Fig. 1</figureCitation>
T
<emphasis id="B9C1EADAF816FFFA8190B6FCFB58F862" bold="true" box="[1155,1169,1942,1967]" pageId="1" pageNumber="76"></emphasis>
V) and from Arakani in Dagestan,
<geoCoordinate id="EE81500FF816FFFA8402B6D1FEA7F81E" box="[273,366,1979,2003]" direction="north" orientation="latitude" pageId="1" pageNumber="76" precision="925" value="42.616665">42°37'N</geoCoordinate>
<geoCoordinate id="EE81500FF816FFFA846AB6D1FE1BF81E" box="[377,466,1979,2003]" direction="east" orientation="longitude" pageId="1" pageNumber="76" precision="925" value="47.0">47°00'E</geoCoordinate>
(ZMMU Lc-7656/1), all on the northern slope of the Greater Caucasus, belong to
<taxonomicName id="4CB54D4BF815FFF98584B1F3FE96FF7D" box="[151,351,151,176]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF815FFF98584B1F3FE96FF7D" box="[151,351,151,176]" italics="true" pageId="2" pageNumber="77">Vertigo substriata</emphasis>
</taxonomicName>
. Near Kobi
<taxonomicName id="4CB54D4BF815FFF984FAB1F3FD95FF7D" box="[489,604,151,176]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF815FFF984FAB1F3FD95FF7D" box="[489,604,151,176]" italics="true" pageId="2" pageNumber="77">V. nitidula</emphasis>
</taxonomicName>
and
<taxonomicName id="4CB54D4BF815FFF98780B1F3FCD7FF7D" box="[659,798,151,176]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF815FFF98780B1F3FCD7FF7D" box="[659,798,151,176]" italics="true" pageId="2" pageNumber="77">V. substriata</emphasis>
</taxonomicName>
occur in sympatry, but do not show transitions (compare
<figureCitation id="138E2A4DF815FFF98584B1D6FF1DFF19" box="[151,212,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
F
<emphasis id="B9C1EADAF815FFF985FBB1D6FF3FFF18" bold="true" box="[232,246,188,213]" pageId="2" pageNumber="77"></emphasis>
G versus 1T
<emphasis id="B9C1EADAF815FFF9846DB1D6FE45FF18" bold="true" box="[382,396,188,213]" pageId="2" pageNumber="77"></emphasis>
V). This confirms the classification of
<taxonomicName id="4CB54D4BF815FFF9862EB1D7FC79FF18" box="[829,944,188,213]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF815FFF9862EB1D7FC79FF18" box="[829,944,188,213]" italics="true" pageId="2" pageNumber="77">V. nitidula</emphasis>
</taxonomicName>
as a separate species.
</paragraph>
<paragraph id="8B0A36C8F815FFF985D4B18AFA4BFE45" blockId="2.[151,1437,151,752]" pageId="2" pageNumber="77">
We re-examined samples of
<taxonomicName id="4CB54D4BF815FFF98709B18BFD62FF35" box="[538,683,223,248]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF815FFF98709B18BFD62FF35" box="[538,683,223,248]" italics="true" pageId="2" pageNumber="77">V. substriata</emphasis>
</taxonomicName>
from the Ural (
<figureCitation id="138E2A4DF815FFF9867CB1B5FC7DFF35" box="[879,948,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
Q; voucher to
<bibRefCitation id="EF244B39F815FFF9817BB1B5FA82FF35" author="Horsak" box="[1128,1355,223,248]" pageId="2" pageNumber="77" refString="Horsak, M., Chytry, M., Danihelka, J., Koci, M., Kubesova, S., Lososova, Z., Otypkova, Z. &amp; Tichy, L. (2010) Snail faunas in the Southern Ural forests and their relations to vegetation: an analogue of the Early Holocene assemblages of Central Europe? Journal of Molluscan Studies, 76, 1 - 10. http: // dx. doi. org / 10.1093 / mollus / eyp 039" type="journal article" year="2010">
Horsák
<emphasis id="B9C1EADAF815FFF981D6B18BFACAFF35" box="[1221,1283,223,248]" italics="true" pageId="2" pageNumber="77">et al.</emphasis>
2010
</bibRefCitation>
), Altai mountains (
<figureCitation id="138E2A4DF815FFF98408B06EFE93FED1" box="[283,346,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
R; voucher to
<bibRefCitation id="EF244B39F815FFF984EFB06EFCEEFED1" author="Pokryszko" box="[508,807,260,285]" pageId="2" pageNumber="77" refString="Pokryszko, B. M. &amp; Horsak, M. (2007) Pupilloidea (Pupillidae, Vertiginidae, Valloniidae, Gastrocoptinae) of the Altay-a travel in space and time. In: Jordaens, K., Van Houtte, N., Van Goethem, J. &amp; Backeljau, T. (Eds.), World Congress of Malacology, Antwerp, Belgium, 15 - 20 July 2007, Abstracts. Unitas Malacologica, Antwerp, pp. 170." type="journal article" year="2007">Pokryszko &amp; Horsák 2007</bibRefCitation>
) and the Crimean Nature Reserve (IZK; partly vouchers to
<bibRefCitation id="EF244B39F815FFF985AAB04DFDE6FE8D" author="Balashov" box="[185,559,295,320]" pageId="2" pageNumber="77" refString="Balashov, I. A. &amp; Baidashnikov, A. A. (2012) The first findings of a slug Boettgerilla pallens (Stylommatophora, Boettgerillidae) in Crimea. Ruthenica, 22, 111 - 114." type="journal article" year="2012">Balashov &amp; Baidashnikov 2012</bibRefCitation>
) and confirmed their specific identity with Central and Western European specimens (
<figureCitation id="138E2A4DF815FFF9840EB026FE97FEA9" box="[285,350,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
P). Consequently,
<taxonomicName id="4CB54D4BF815FFF98722B027FD6CFEA8" box="[561,677,332,357]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF815FFF98722B027FD6CFEA8" box="[561,677,332,357]" italics="true" pageId="2" pageNumber="77">V. nitidula</emphasis>
</taxonomicName>
can be ranked as a Caucasian endemic, while
<taxonomicName id="4CB54D4BF815FFF981AEB027FA83FEA8" box="[1213,1354,332,357]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF815FFF981AEB027FA83FEA8" box="[1213,1354,332,357]" italics="true" pageId="2" pageNumber="77">V. substriata</emphasis>
</taxonomicName>
ranges from Western Europe to the Ural and the north side of the Great Caucasus with isolated occurrences in the Altai.
</paragraph>
</subSubSection>
<subSubSection id="C3AF6543F815FFFE85D4B0FEFBF9FC90" lastPageId="5" lastPageNumber="80" pageId="2" pageNumber="77" type="materials_examined">
<paragraph id="8B0A36C8F815FFF985D4B0FEFB96FD3D" blockId="2.[151,1437,151,752]" pageId="2" pageNumber="77">
<emphasis id="B9C1EADAF815FFF985D4B0FEFEBEFE60" bold="true" box="[199,375,404,429]" pageId="2" pageNumber="77">Nomenclature.</emphasis>
<bibRefCitation id="EF244B39F815FFF98491B0FFFDF6FE60" author="Mousson" box="[386,575,404,429]" pageId="2" pageNumber="77" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson (1876)</bibRefCitation>
was the first who described this species as
<emphasis id="B9C1EADAF815FFF9815EB0FFFB41FE61" box="[1101,1160,405,428]" italics="true" pageId="2" pageNumber="77">Pupa</emphasis>
(
<taxonomicName id="4CB54D4BF815FFF9818FB0FFFB24FE61" box="[1180,1261,405,428]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="genus">
<emphasis id="B9C1EADAF815FFF9818FB0FFFB24FE61" box="[1180,1261,405,428]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
</taxonomicName>
)
<emphasis id="B9C1EADAF815FFF98011B0FFFAAEFE61" box="[1282,1383,405,428]" italics="true" pageId="2" pageNumber="77">pygmaea</emphasis>
<taxonomicName id="4CB54D4BF815FFF98061B0FFFF27FE1D" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF815FFF98584B0DDFF27FE1D" box="[151,238,439,464]" italics="true" pageId="2" pageNumber="77">nitidula</emphasis>
</taxonomicName>
based on material collected in Tabatsquri by G. Sievers. However, his description does not well characterize the species. It does not even well fit the
<typeStatus id="540E886AF815FFF987EAB0B7FC92FE38" box="[761,859,477,501]" pageId="2" pageNumber="77" type="syntype">syntypes</typeStatus>
(
<figureCitation id="138E2A4DF815FFF9867FB0B6FC67FE39" box="[876,942,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
A; the size of this
<typeStatus id="540E886AF815FFF98194B0B7FB17FE38" box="[1159,1246,477,501]" pageId="2" pageNumber="77" type="syntype">syntype</typeStatus>
is 1.9 mm x 1.0 mm; the other
<typeStatus id="540E886AF815FFF98428B36AFE5BFDD5" box="[315,402,512,536]" pageId="2" pageNumber="77" type="syntype">syntype</typeStatus>
is damaged). Nevertheless, there is no doubt about the authenticity of the
<typeStatus id="540E886AF815FFF981C0B36AFAF1FDD5" box="[1235,1336,512,536]" pageId="2" pageNumber="77" type="syntype">syntypes</typeStatus>
, because there is only one sample of
<emphasis id="B9C1EADAF815FFF984C1B34FFDB3FDF1" box="[466,634,549,572]" italics="true" pageId="2" pageNumber="77">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF815FFF98791B34FFCC2FDF0" box="[642,779,548,573]" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF815FFF987A7B34EFCC2FDF0" box="[692,779,548,573]" italics="true" pageId="2" pageNumber="77">nitidula</emphasis>
</taxonomicName>
in Moussons collection. There are no specimens of other
<taxonomicName id="4CB54D4BF815FFF98584B323FF21FDAD" box="[151,232,585,608]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="genus">
<emphasis id="B9C1EADAF815FFF98584B323FF21FDAD" box="[151,232,585,608]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
</taxonomicName>
species from Tabatsquri in Moussons or in any other examined collection. The authenticity of the
<typeStatus id="540E886AF815FFF98072B322FA55FDAD" box="[1377,1436,584,608]" pageId="2" pageNumber="77">types</typeStatus>
gets further support from the fact that
<bibRefCitation id="EF244B39F815FFF9875AB307FD35FD49" author="Boettger" box="[585,764,620,645]" pageId="2" pageNumber="77" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger (1879)</bibRefCitation>
got conspecific specimens from the same locality from the same collector. The discrepancies between the
<typeStatus id="540E886AF815FFF987C7B3FAFCCDFD65" box="[724,772,656,680]" pageId="2" pageNumber="77">type</typeStatus>
specimens and the description might be explained by Moussons insufficient optical equipment or his impaired vision as a result of his age. Disagreements between Moussons descriptions and his vouchers were also found in several other microsnails.
</paragraph>
<caption id="DFCA6640F815FFF98584B7B9FF0FF80D" httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="2" pageNumber="77" targetBox="[151,1436,806,1726]" targetPageId="2">
<paragraph id="8B0A36C8F815FFF98584B7B9FF0FF80D" blockId="2.[151,1437,1747,1984]" pageId="2" pageNumber="77">
<emphasis id="B9C1EADAF815FFF98584B7B9FEDCF924" bold="true" box="[151,277,1747,1769]" pageId="2" pageNumber="77">FIGURE 1.</emphasis>
<taxonomicName id="4CB54D4BF815FFF9840FB7BEFEACF924" box="[284,357,1748,1769]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="genus">
<emphasis id="B9C1EADAF815FFF9840FB7BEFEACF924" box="[284,357,1748,1769]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
</taxonomicName>
spp., shells.
<emphasis id="B9C1EADAF815FFF984FDB7B9FDEEF924" bold="true" box="[494,551,1747,1769]" pageId="2" pageNumber="77">AO.</emphasis>
<taxonomicName id="4CB54D4BF815FFF9873CB7BEFC6EF924" authority="Mousson" authorityName="Mousson" box="[559,935,1747,1769]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="nitidula" subGenus="Vertigo">
<emphasis id="B9C1EADAF815FFF9873CB7BEFDB1F924" box="[559,632,1748,1769]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF815FFF9879AB7BEFD1BF924" box="[649,722,1748,1769]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF815FFF987F0B7B9FCFBF924" box="[739,818,1747,1769]" italics="true" pageId="2" pageNumber="77">nitidula</emphasis>
(Mousson)
</taxonomicName>
.
<emphasis id="B9C1EADAF815FFF986A6B7B9FC07F924" bold="true" box="[949,974,1747,1769]" pageId="2" pageNumber="77">A.</emphasis>
Georgia: Tabatsquri (syntype of
<emphasis id="B9C1EADAF815FFF98035B7BEFA54F924" box="[1318,1437,1748,1769]" italics="true" pageId="2" pageNumber="77">P. pygmaea</emphasis>
<taxonomicName id="4CB54D4BF815FFF98584B799FEDDF8C5" box="[151,276,1778,1800]" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF815FFF985D6B798FEDDF8C5" box="[197,276,1778,1800]" italics="true" pageId="2" pageNumber="77">nitidula</emphasis>
</taxonomicName>
ZMZ 514689).
<emphasis id="B9C1EADAF815FFF984ADB798FE1CF8C5" bold="true" box="[446,469,1778,1800]" pageId="2" pageNumber="77">B.</emphasis>
Georgia, Tabatsquri (syntype of
<emphasis id="B9C1EADAF815FFF98623B799FC51F8C5" box="[816,920,1779,1800]" italics="true" pageId="2" pageNumber="77">
P.
<taxonomicName id="4CB54D4BF815FFF98658B799FC51F8C5" box="[843,920,1779,1800]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
SMF 2129).
<emphasis id="B9C1EADAF815FFF98134B798FB89F8C5" bold="true" box="[1063,1088,1778,1800]" pageId="2" pageNumber="77">C.</emphasis>
Georgia, Manglisi (syntype of
<emphasis id="B9C1EADAF815FFF9809AB799FF2DF8EA" italics="true" pageId="2" pageNumber="77">
P.
<taxonomicName id="4CB54D4BF815FFF98584B678FF2DF8EA" box="[151,228,1810,1831]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF815FFF985FCB678FEBFF8EA" box="[239,374,1810,1831]" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="punctum">
var.
<emphasis id="B9C1EADAF815FFF9840CB678FEBFF8EA" box="[287,374,1810,1831]" italics="true" pageId="2" pageNumber="77">punctum</emphasis>
</taxonomicName>
SMF 2125).
<emphasis id="B9C1EADAF815FFF9871FB67BFDECF8EA" bold="true" box="[524,549,1809,1831]" pageId="2" pageNumber="77">D.</emphasis>
Georgia, Kazbek (syntype of
<emphasis id="B9C1EADAF815FFF9867FB678FC1EF8EA" box="[876,983,1810,1831]" italics="true" pageId="2" pageNumber="77">
P.
<taxonomicName id="4CB54D4BF815FFF98699B678FC1EF8EA" box="[906,983,1810,1831]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF815FFF986F1B678FB44F8EA" box="[994,1165,1809,1831]" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="subalpestris">
var.
<emphasis id="B9C1EADAF815FFF98100B67BFB44F8EA" box="[1043,1165,1809,1831]" italics="true" pageId="2" pageNumber="77">subalpestris</emphasis>
</taxonomicName>
SMF 2122).
<emphasis id="B9C1EADAF815FFF98031B67BFAF3F8EA" bold="true" box="[1314,1338,1809,1831]" pageId="2" pageNumber="77">E.</emphasis>
Georgia, Abastumani (syntype of
<emphasis id="B9C1EADAF815FFF98487B65BFDDDF888" box="[404,532,1839,1862]" italics="true" pageId="2" pageNumber="77">
P.
<taxonomicName id="4CB54D4BF815FFF984BDB645FDDDF888" box="[430,532,1839,1861]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata">substriata</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF815FFF98708B65AFDB1F88B" box="[539,632,1840,1862]" kingdom="Animalia" pageId="2" pageNumber="77" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF815FFF98754B65BFDB1F88B" box="[583,632,1841,1862]" italics="true" pageId="2" pageNumber="77">mitis</emphasis>
</taxonomicName>
SMF 2282).
<emphasis id="B9C1EADAF815FFF98610B645FCFFF888" bold="true" box="[771,822,1839,1861]" pageId="2" pageNumber="77">FG.</emphasis>
Georgia, Kobi (ZIN 27).
<emphasis id="B9C1EADAF815FFF98153B645FBBDF888" bold="true" box="[1088,1140,1839,1861]" pageId="2" pageNumber="77">HJ.</emphasis>
Russia, lower Adylsu valley (ZMH 80612).
<emphasis id="B9C1EADAF815FFF98425B624FEB8F8A9" bold="true" box="[310,369,1870,1892]" pageId="2" pageNumber="77">KO.</emphasis>
Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614).
<emphasis id="B9C1EADAF815FFF986ADB624FC26F8A9" bold="true" box="[958,1007,1870,1892]" pageId="2" pageNumber="77">PV.</emphasis>
<taxonomicName id="4CB54D4BF815FFF986EBB625FAB4F8A9" authority="Jeffreys" authorityName="Jeffreys" box="[1016,1405,1870,1892]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="77" phylum="Mollusca" rank="species" species="substriata" subGenus="Vertigo">
<emphasis id="B9C1EADAF815FFF986EBB625FB88F8A9" box="[1016,1089,1871,1892]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF815FFF98141B625FB52F8A9" box="[1106,1179,1871,1892]" italics="true" pageId="2" pageNumber="77">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF815FFF981BFB624FADBF8A9" box="[1196,1298,1870,1892]" italics="true" pageId="2" pageNumber="77">substriata</emphasis>
(Jeffreys)
</taxonomicName>
.
<emphasis id="B9C1EADAF815FFF9809BB624FA53F8A9" bold="true" box="[1416,1434,1870,1892]" pageId="2" pageNumber="77">P.</emphasis>
Germany, Nübel (ZMH 60278);
<emphasis id="B9C1EADAF815FFF984F3B607FE33F84E" bold="true" box="[480,506,1901,1923]" pageId="2" pageNumber="77">Q.</emphasis>
Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098).
<emphasis id="B9C1EADAF815FFF9816CB607FB51F84E" bold="true" box="[1151,1176,1901,1923]" pageId="2" pageNumber="77">R.</emphasis>
Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096).
<emphasis id="B9C1EADAF815FFF98743B6E1FDADF86C" bold="true" box="[592,612,1931,1953]" pageId="2" pageNumber="77">S.</emphasis>
Russia, Chegem valley (ZIN 96).
<emphasis id="B9C1EADAF815FFF986D5B6E1FC30F86C" bold="true" box="[966,1017,1931,1953]" pageId="2" pageNumber="77">TV.</emphasis>
Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DFCA6640F814FFF88584B580FC33FACD" ID-DOI="http://doi.org/10.5281/zenodo.8387550" ID-Zenodo-Dep="8387550" box="[151,1018,1258,1280]" httpUri="https://zenodo.org/record/8387550/files/figure.png" pageId="3" pageNumber="78" startId="3.[151,250,1258,1280]" targetBox="[345,1239,199,1229]" targetPageId="3">
<paragraph id="8B0A36C8F814FFF88584B580FC33FACD" blockId="3.[151,1018,1258,1280]" box="[151,1018,1258,1280]" pageId="3" pageNumber="78">
<emphasis id="B9C1EADAF814FFF88584B580FEDBFACD" bold="true" box="[151,274,1258,1280]" pageId="3" pageNumber="78">FIGURE 2.</emphasis>
Variation in shell size and shape of
<taxonomicName id="4CB54D4BF814FFF88793B581FC3FFACD" authority="Mousson" authorityName="Mousson" box="[640,1014,1258,1280]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="nitidula" subGenus="Vertigo">
<emphasis id="B9C1EADAF814FFF88793B581FD00FACD" box="[640,713,1259,1280]" italics="true" pageId="3" pageNumber="78">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF814FFF887CBB581FCE8FACD" box="[728,801,1259,1280]" italics="true" pageId="3" pageNumber="78">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF814FFF8863CB580FCB7FACD" box="[815,894,1258,1280]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
(Mousson)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B0A36C8F814FFF885D4B44DFB03F9F1" blockId="3.[151,1437,1319,2029]" pageId="3" pageNumber="78">
Although
<bibRefCitation id="EF244B39F814FFF8842FB442FE3AFA8D" author="Boettger" box="[316,499,1320,1344]" pageId="3" pageNumber="78" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger (1879)</bibRefCitation>
supposed that the
<taxonomicName id="4CB54D4BF814FFF887C6B443FCEAFA8D" box="[725,803,1321,1344]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="genus">
<emphasis id="B9C1EADAF814FFF887C6B443FCEAFA8D" box="[725,803,1321,1344]" italics="true" pageId="3" pageNumber="78">Vertigo</emphasis>
</taxonomicName>
species from the
<typeStatus id="540E886AF814FFF886EEB442FBE4FA8D" box="[1021,1069,1320,1344]" pageId="3" pageNumber="78">type</typeStatus>
locality of
<emphasis id="B9C1EADAF814FFF881AFB443FAAEFA8D" box="[1212,1383,1321,1344]" italics="true" pageId="3" pageNumber="78">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF814FFF88060B442FF27FAA8" kingdom="Animalia" pageId="3" pageNumber="78" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF814FFF88584B426FF27FAA8" box="[151,238,1356,1381]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
</taxonomicName>
that he also got from G. Sievers is identical with the taxon described by
<bibRefCitation id="EF244B39F814FFF88125B427FB3BFAA8" author="Mousson" box="[1078,1266,1356,1381]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson (1876)</bibRefCitation>
, he renamed it
<emphasis id="B9C1EADAF814FFF88584B41BFF1BFA45" box="[151,210,1393,1416]" italics="true" pageId="3" pageNumber="78">Pupa</emphasis>
(
<emphasis id="B9C1EADAF814FFF885F0B41BFEF8FA45" box="[227,305,1393,1416]" italics="true" pageId="3" pageNumber="78">Vert igo</emphasis>
)
<taxonomicName id="4CB54D4BF814FFF88457B41BFE50FA45" box="[324,409,1393,1416]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF814FFF88457B41BFE50FA45" box="[324,409,1393,1416]" italics="true" pageId="3" pageNumber="78">sieversi</emphasis>
</taxonomicName>
. This name was used for the species by most of the later authors.
<bibRefCitation id="EF244B39F814FFF88190B405FAEDFA45" author="Pilsbry" box="[1155,1316,1391,1416]" pageId="3" pageNumber="78" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry (1919)</bibRefCitation>
noted that he was not sure whether the name
<taxonomicName id="4CB54D4BF814FFF88735B4FEFDB4FA60" box="[550,637,1428,1453]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF814FFF88735B4FEFDB4FA60" box="[550,637,1428,1453]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
</taxonomicName>
should be rejected. He mentioned
<taxonomicName id="4CB54D4BF814FFF8811EB4FEFA51FA60" authority="Lamarck, 1816" authorityName="Lamarck" authorityYear="1816" box="[1037,1432,1428,1453]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF814FFF8811EB4FEFB28FA60" box="[1037,1249,1428,1453]" italics="true" pageId="3" pageNumber="78">Tornatella nitidula</emphasis>
<bibRefCitation id="EF244B39F814FFF881F9B4FEFA51FA60" author="Lamarck" box="[1258,1432,1428,1453]" pageId="3" pageNumber="78" refString="Lamarck, J. B. P. A. (1816) Mollusques et polypes divers. In: Tableau encyclopedique et methodique des trois regnes de la nature. Vers, coquilles, mollusques et polypiers. Vol. 3. (23). Agasse, Paris, 16 pp., pls. 391 - 488." type="book chapter" year="1816">Lamarck, 1816</bibRefCitation>
</taxonomicName>
, which is today included in
<emphasis id="B9C1EADAF814FFF884CEB4D3FDD1FA1D" box="[477,536,1465,1488]" italics="true" pageId="3" pageNumber="78">Pupa</emphasis>
<bibRefCitation id="EF244B39F814FFF88737B4DDFD0CFA1D" author="Roding" box="[548,709,1463,1488]" pageId="3" pageNumber="78" refString="Roding, P. F. (1798) Museum Boltenianum sive catalogus cimeliorum e tribus regnis naturae quae olim collegerat Joa. Fried. Bolten. Pars secunda continens Conchylia sive Testacea univalvia, bivalvia &amp; multivalvia. J. C. Trappii, Hamburg, viii + 199 pp. http: // dx. doi. org / 10.5962 / bhl. title. 16250" type="book" year="1798">Röding, 1798</bibRefCitation>
. However,
<emphasis id="B9C1EADAF814FFF8865FB4D3FC30FA1D" box="[844,1017,1465,1488]" italics="true" pageId="3" pageNumber="78">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF814FFF88117B4D2FA84FA1D" authority="Mousson, 1876" authorityName="Mousson" authorityYear="1876" box="[1028,1357,1463,1488]" kingdom="Animalia" pageId="3" pageNumber="78" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF814FFF8812AB4DDFB59FA1D" box="[1081,1168,1463,1488]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
<bibRefCitation id="EF244B39F814FFF8818FB4D2FA84FA1D" author="Mousson" box="[1180,1357,1463,1488]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
</taxonomicName>
is not homonymous with
<emphasis id="B9C1EADAF814FFF8847DB4B7FE60FA39" box="[366,425,1501,1524]" italics="true" pageId="3" pageNumber="78">Pupa</emphasis>
<taxonomicName id="4CB54D4BF814FFF884BCB4B6FD02FA38" authority="(Lamarck, 1816)" authorityName="Lamarck" authorityYear="1816" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[431,715,1500,1525]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF814FFF884BCB4B6FDCFFA38" box="[431,518,1500,1525]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
(
<bibRefCitation id="EF244B39F814FFF88707B4B6FD0AFA38" author="Lamarck" box="[532,707,1500,1525]" pageId="3" pageNumber="78" refString="Lamarck, J. B. P. A. (1816) Mollusques et polypes divers. In: Tableau encyclopedique et methodique des trois regnes de la nature. Vers, coquilles, mollusques et polypiers. Vol. 3. (23). Agasse, Paris, 16 pp., pls. 391 - 488." type="book chapter" year="1816">Lamarck, 1816</bibRefCitation>
)
</taxonomicName>
, because it has been established in the homonymous genus
<emphasis id="B9C1EADAF814FFF88072B4B7FA54FA39" box="[1377,1437,1501,1524]" italics="true" pageId="3" pageNumber="78">Pupa</emphasis>
<bibRefCitation id="EF244B39F814FFF88584B495FEAEF9D5" author="Draparnaud" box="[151,359,1535,1560]" pageId="3" pageNumber="78" refString="Draparnaud, J. (1801) Tableau des mollusques terrestres et fluviatiles de la France. Renaud, Montpellier; Bossange, Masson &amp; Besson, Paris, 116 pp. http: // dx. doi. org / 10.5962 / bhl. title. 40947" type="book" year="1801">Draparnaud, 1801</bibRefCitation>
and homonymy between identical species-group names in combination with homonymous generic names established for different nominal genera is to be disregarded (Art. 57.8.1.
<collectionCode id="EDA4AE0DF814FFF88164B74EFB77F9F1" box="[1143,1214,1572,1596]" pageId="3" pageNumber="78">ICZN</collectionCode>
).
</paragraph>
<paragraph id="8B0A36C8F814FFF885D4B723FBBAF901" blockId="3.[151,1437,1319,2029]" pageId="3" pageNumber="78">
<emphasis id="B9C1EADAF814FFF885D4B723FEA4F9AD" box="[199,365,1609,1632]" italics="true" pageId="3" pageNumber="78">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF814FFF88467B722FD64F9AD" authority="Mousson, 1876" authorityName="Mousson" authorityYear="1876" box="[372,685,1607,1632]" kingdom="Animalia" pageId="3" pageNumber="78" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF814FFF884B7B72DFE32F9AD" box="[420,507,1607,1632]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
<bibRefCitation id="EF244B39F814FFF88712B722FD64F9AD" author="Mousson" box="[513,685,1607,1632]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
</taxonomicName>
has been considered a synonym of
<taxonomicName id="4CB54D4BF814FFF8812BB723FAB0F9AD" authority="Jeffreys, 1833" authorityName="Jeffreys" authorityYear="1833" box="[1080,1401,1607,1632]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF814FFF8812BB723FB0AF9AD" box="[1080,1219,1607,1632]" italics="true" pageId="3" pageNumber="78">V. substriata</emphasis>
(
<bibRefCitation id="EF244B39F814FFF881C2B72DFAB8F9AD" author="Jeffreys" box="[1233,1393,1607,1632]" pageId="3" pageNumber="78" refString="Jeffreys, J. G. (1833) A Supplement to the ' Synopsis of the Testaceous Pneumobranchous Mollusca of Great Britain'. Transactions of the Linnean Society of London, 16 (3), 505 - 523." type="journal article" year="1833">Jeffreys, 1833</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EF244B39F814FFF88584B706FE94F949" author="Schileyko" box="[151,349,1644,1669]" pageId="3" pageNumber="78" refString="Schileyko, A. A. (1984) Nazemnye mollyuski podotryada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In: Fauna SSSR. Mollyuski. III (3). Nauka, Leningrad, pp. 1 - 399." type="journal article" year="1984">Schileyko (1984)</bibRefCitation>
,
<bibRefCitation id="EF244B39F814FFF8847FB706FD67F948" author="Sysoev" box="[364,686,1644,1669]" pageId="3" pageNumber="78" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2005) Neformalnaya gruppa Stylommatophora. In: Kantor, Y. I. &amp; Sysoev, A. V. (Eds.), Katalog Mollyuskov Rossii i sopredelnykh stran. KMK Scientific Press, Moscow, pp. 228 - 308." type="book chapter" year="2005">Sysoev &amp; Schileyko (2005)</bibRefCitation>
,
<bibRefCitation id="EF244B39F814FFF887AEB707FCADF949" author="Egorov" box="[701,868,1644,1669]" pageId="3" pageNumber="78" refString="Egorov, R. V. (2008) Treasure of Russian Shells, Supplement 5. Illustrated catalogue of the recent terrestrial molluscs of Russia and adjacent regions. Colus-Doveri, Moscow, 179 pp." type="book" year="2008">Egorov (2008)</bibRefCitation>
and
<bibRefCitation id="EF244B39F814FFF886B3B706FB2BF949" author="Sysoev" box="[928,1250,1644,1669]" pageId="3" pageNumber="78" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2009) Land snails and slugs of Russia and adjacent countries. Pensoft, Sofia and Moscow, 312 pp., 142 pls." type="book" year="2009">Sysoev &amp; Schileyko (2009)</bibRefCitation>
. However, a reexamination of the
<typeStatus id="540E886AF814FFF88466B7FAFE1EF965" box="[373,471,1680,1704]" pageId="3" pageNumber="78" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF814FFF884ECB7FBFD61F965" box="[511,680,1681,1704]" italics="true" pageId="3" pageNumber="78">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF814FFF887A3B7FAFC39F965" authority="Mousson, 1876" authorityName="Mousson" authorityYear="1876" box="[688,1008,1679,1704]" kingdom="Animalia" pageId="3" pageNumber="78" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF814FFF887F1B7E5FCF0F965" box="[738,825,1679,1704]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
<bibRefCitation id="EF244B39F814FFF88651B7FAFC39F965" author="Mousson" box="[834,1008,1679,1704]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
</taxonomicName>
from Tabatsquri (
<figureCitation id="138E2A4DF814FFF881ADB7E5FB36F965" box="[1214,1279,1679,1704]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1769]" captionTargetBox="[151,1436,806,1726]" captionTargetId="figure@2.[151,1436,806,1726]" captionTargetPageId="2" captionText="FIGURE 1. Vertigo spp., shells. A O. Vertigo (Vertigo) nitidula (Mousson). A. Georgia: Tabatsquri (syntype of P. pygmaea var. nitidula ZMZ 514689). B. Georgia, Tabatsquri (syntype of P. sieversi SMF 2129). C. Georgia, Manglisi (syntype of P. sieversi var. punctum SMF 2125). D. Georgia, Kazbek (syntype of P. sieversi var. subalpestris SMF 2122). E. Georgia, Abastumani (syntype of P. substriata var. mitis SMF 2282). F G. Georgia, Kobi (ZIN 27). H J. Russia, lower Adylsu valley (ZMH 80612). K O. Armenia, Tsaghkadzor, Mount Teghenis (ZMH 80614). P V. Vertigo (Vertigo) substriata (Jeffreys). P. Germany, Nübel (ZMH 60278); Q. Russia, Zilairskoe plateau, Novoaleksandrovka (ZMH 96098). R. Russia, Altai Republic, 2 km W of Zaisanskaya Elan (ZMH 96096). S. Russia, Chegem valley (ZIN 96). T V. Georgia, Kobi (ZIN 95). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/287228/files/figure.png" pageId="3" pageNumber="78">Fig. 1</figureCitation>
A) confirmed the supposition of
<bibRefCitation id="EF244B39F814FFF88475B7DFFDD0F901" author="Boettger" box="[358,537,1716,1741]" pageId="3" pageNumber="78" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger (1879)</bibRefCitation>
that this taxon is synonymous with his
<emphasis id="B9C1EADAF814FFF886C7B7DFFBA2F901" box="[980,1131,1717,1740]" italics="true" pageId="3" pageNumber="78">
Pupa
<taxonomicName id="4CB54D4BF814FFF88105B7DFFBA2F901" box="[1046,1131,1717,1740]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="8B0A36C8F814FFF885D4B7BDFBE0F805" blockId="3.[151,1437,1319,2029]" pageId="3" pageNumber="78">
Although
<emphasis id="B9C1EADAF814FFF88428B7B3FE2FF93D" box="[315,486,1753,1776]" italics="true" pageId="3" pageNumber="78">Pupa pygmaea</emphasis>
<taxonomicName id="4CB54D4BF814FFF884E2B7B2FCFEF93D" authority="Mousson, 1876" authorityName="Mousson" authorityYear="1876" box="[497,823,1751,1776]" kingdom="Animalia" pageId="3" pageNumber="78" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF814FFF88736B7BDFDB5F93D" box="[549,636,1751,1776]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
<bibRefCitation id="EF244B39F814FFF88795B7B2FCFEF93D" author="Mousson" box="[646,823,1751,1776]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
</taxonomicName>
has not been used as a valid name after its original description, it is not a nomen oblitum in the sense of Art. 23.9
<collectionCode id="EDA4AE0DF814FFF8864BB796FC50F8D9" box="[856,921,1788,1812]" pageId="3" pageNumber="78">ICNZ</collectionCode>
because
<emphasis id="B9C1EADAF814FFF88113B797FBF2F8D9" box="[1024,1083,1789,1812]" italics="true" pageId="3" pageNumber="78">Pupa</emphasis>
<taxonomicName id="4CB54D4BF814FFF88151B797FA8DF8D9" authority="Boettger, 1879" authorityName="Boettger" authorityYear="1879" box="[1090,1348,1788,1813]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF814FFF88151B797FB5EF8D9" box="[1090,1175,1789,1812]" italics="true" pageId="3" pageNumber="78">sieversi</emphasis>
<bibRefCitation id="EF244B39F814FFF8818DB797FA8DF8D9" author="Boettger" box="[1182,1348,1788,1813]" pageId="3" pageNumber="78" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger, 1879</bibRefCitation>
</taxonomicName>
has not been used as the presumed valid name of that taxon in 25 works in the past 50 years. Actually, a survey of more than 360 works containing data about Caucasian land snails resulted in only 11 works from the past 50 years citing
<taxonomicName id="4CB54D4BF814FFF88584B603FEC0F84D" box="[151,265,1897,1920]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="sieversi">
<emphasis id="B9C1EADAF814FFF88584B603FEC0F84D" box="[151,265,1897,1920]" italics="true" pageId="3" pageNumber="78">V. sieversi</emphasis>
</taxonomicName>
as presumed valid name. Although we cannot exclude that there are a few additional papers, in which this name has been used as presumed valid name, it is very unlikely that the actual number of uses exceeds 24. Thus, the valid name of the species is
<taxonomicName id="4CB54D4BF814FFF88751B6DBFBECF805" authority="Mousson, 1876" authorityName="Mousson" authorityYear="1876" box="[578,1061,1967,1992]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="nitidula" subGenus="Vertigo">
<emphasis id="B9C1EADAF814FFF88751B6DBFD5AF805" box="[578,659,1969,1992]" italics="true" pageId="3" pageNumber="78">Vertigo</emphasis>
(
<emphasis id="B9C1EADAF814FFF887B0B6DBFD3DF805" box="[675,756,1969,1992]" italics="true" pageId="3" pageNumber="78">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF814FFF88617B6C5FC92F805" box="[772,859,1967,1992]" italics="true" pageId="3" pageNumber="78">nitidula</emphasis>
(
<bibRefCitation id="EF244B39F814FFF88679B6DAFBD5F805" author="Mousson" box="[874,1052,1967,1992]" pageId="3" pageNumber="78" refString="Mousson, A. (1876) Coquilles recueillies par M. le Dr. Sievers dans la Russie Asiatique. Journal de Conchyliologie, 24, 137 - 148, pls. 5." type="journal article" year="1876">Mousson, 1876</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B0A36C8F814FFFF85D4B6BFFC78FED0" blockId="3.[151,1437,1319,2029]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78">
<bibRefCitation id="EF244B39F814FFF885D4B6BFFEB3F821" author="Boettger" box="[199,378,2004,2029]" pageId="3" pageNumber="78" refString="Boettger, O. (1879) Kaukasische Mollusken. Gesammelt von Herrn Dr. G. Sievers in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 6, 388 - 412, pl. 10." type="journal article" year="1879">Boettger (1879)</bibRefCitation>
introduced two different spellings for a variety of
<emphasis id="B9C1EADAF814FFF886A5B6BFFB87F821" box="[950,1102,2005,2028]" italics="true" pageId="3" pageNumber="78">
Pupa
<taxonomicName id="4CB54D4BF814FFF886EAB6BFFB87F821" box="[1017,1102,2005,2028]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="78" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
:
<emphasis id="B9C1EADAF814FFF8814DB6BFFB77F821" box="[1118,1214,2005,2028]" italics="true" pageId="3" pageNumber="78">punctum</emphasis>
and
<emphasis id="B9C1EADAF814FFF881E5B6BEFAA5F820" box="[1270,1388,2004,2029]" italics="true" pageId="3" pageNumber="78">punctulum</emphasis>
. By using the spelling
<emphasis id="B9C1EADAF813FFFF8477B1F3FE56FF7D" box="[356,415,153,176]" italics="true" pageId="4" pageNumber="79">Pupa</emphasis>
<taxonomicName id="4CB54D4BF813FFFF84B5B1F2FDAFFF7D" box="[422,614,152,176]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF813FFFF84A3B1F3FE37FF7D" box="[432,510,153,176]" italics="true" pageId="4" pageNumber="79">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF813FFFF8702B1F3FDAFFF7D" box="[529,614,153,176]" italics="true" pageId="4" pageNumber="79">sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF813FFFF877DB1F2FD36FF7D" box="[622,767,152,176]" kingdom="Animalia" pageId="4" pageNumber="79" rank="variety" variety="punctum">
var.
<emphasis id="B9C1EADAF813FFFF878DB1F3FD36FF7D" box="[670,767,153,176]" italics="true" pageId="4" pageNumber="79">punctum</emphasis>
</taxonomicName>
as valid,
<bibRefCitation id="EF244B39F813FFFF8678B1F2FBD4FF7D" author="Boettger" box="[875,1053,152,176]" pageId="4" pageNumber="79" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger (1880)</bibRefCitation>
determined that this is the correct spelling (Art. 24.2.4
<collectionCode id="EDA4AE0DF813FFFF8491B1D6FE01FF19" box="[386,456,188,212]" pageId="4" pageNumber="79">ICZN</collectionCode>
).
<emphasis id="B9C1EADAF813FFFF84C8B1D7FDDFFF19" box="[475,534,189,212]" italics="true" pageId="4" pageNumber="79">Pupa</emphasis>
<taxonomicName id="4CB54D4BF813FFFF870EB1D6FD14FF19" box="[541,733,188,212]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="sieversi" subGenus="Vertigo">
(
<emphasis id="B9C1EADAF813FFFF8735B1D7FDBEFF19" box="[550,631,189,212]" italics="true" pageId="4" pageNumber="79">Vertigo</emphasis>
)
<emphasis id="B9C1EADAF813FFFF879BB1D7FD14FF19" box="[648,733,189,212]" italics="true" pageId="4" pageNumber="79">sieversi</emphasis>
</taxonomicName>
<taxonomicName id="4CB54D4BF813FFFF87F6B1D7FC45FF18" box="[741,908,188,213]" kingdom="Animalia" pageId="4" pageNumber="79" rank="variety" variety="punctulum">
var.
<emphasis id="B9C1EADAF813FFFF8605B1D6FC45FF18" box="[790,908,188,213]" italics="true" pageId="4" pageNumber="79">punctulum</emphasis>
</taxonomicName>
is therefore an incorrect and, thus, unavailable spelling. The selection of
<emphasis id="B9C1EADAF813FFFF84DDB1B5FD8DFF35" box="[462,580,223,248]" italics="true" pageId="4" pageNumber="79">punctulum</emphasis>
as the correct spelling by
<bibRefCitation id="EF244B39F813FFFF8699B1B5FBF9FF35" author="Pilsbry" box="[906,1072,223,248]" pageId="4" pageNumber="79" refString="Pilsbry, H. A. (1919) Pupillidae (Gastrocoptinae, Vertigininae). In: Pilsbry, H. A. (Ed.), Manual of Conchology, Second Series: Pulmonata. Academy of natural Sciences of Philadelphia, Philadelphia, 25 (99), 145 - 224, pls. 11 - 18." type="journal article" year="1919">Pilsbry (1919)</bibRefCitation>
is not valid as a result of the implementation of Art. 24.2.4
<collectionCode id="EDA4AE0DF813FFFF84FFB06EFDE4FED1" box="[492,557,260,284]" pageId="4" pageNumber="79">ICZN</collectionCode>
in the present edition of the Code.
</paragraph>
<paragraph id="8B0A36C8F813FFFF85D4B04DFD68FE38" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<emphasis id="B9C1EADAF813FFFF85D4B04DFEB1FE8D" bold="true" box="[199,376,295,320]" pageId="4" pageNumber="79">
<typeStatus id="540E886AF813FFFF85D4B04DFECBFE8D" box="[199,258,295,320]" pageId="4" pageNumber="79">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F3A27658F813FFFF8490B04DFE29FE8D" box="[387,480,295,320]" name="Georgia" pageId="4" pageNumber="79">Georgia</collectingCountry>
, Mtskheta-Mtianeti region: Kazbek (
<typeStatus id="540E886AF813FFFF869EB042FC38FE8D" box="[909,1009,296,320]" pageId="4" pageNumber="79" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF813FFFF810EB043FB5BFE8D" box="[1053,1170,297,320]" italics="true" pageId="4" pageNumber="79">
P.
<taxonomicName id="4CB54D4BF813FFFF812EB043FB5BFE8D" box="[1085,1170,297,320]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
<taxonomicName id="4CB54D4BF813FFFF818EB042FA93FE8D" box="[1181,1370,295,320]" kingdom="Animalia" pageId="4" pageNumber="79" rank="variety" variety="subalpestris">
var.
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2121/2, 2122/1;
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of
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P.
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</emphasis>
<taxonomicName id="4CB54D4BF813FFFF8795B027FD38FEA9" box="[646,753,333,356]" kingdom="Animalia" pageId="4" pageNumber="79" rank="variety" variety="mitis">
var.
<emphasis id="B9C1EADAF813FFFF87A8B027FD38FEA9" box="[699,753,333,356]" italics="true" pageId="4" pageNumber="79">mitis</emphasis>
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2023/3); Samtskhe-Javakheti region: Borjomi: Kura debris (
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<emphasis id="B9C1EADAF813FFFF8458B01BFE94FE45" box="[331,349,369,392]" italics="true" pageId="4" pageNumber="79">P.</emphasis>
of
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<emphasis id="B9C1EADAF813FFFF8495B01BFE12FE45" box="[390,475,369,392]" italics="true" pageId="4" pageNumber="79">sieversi</emphasis>
</taxonomicName>
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var.
<emphasis id="B9C1EADAF813FFFF8707B01BFDBCFE45" box="[532,629,369,392]" italics="true" pageId="4" pageNumber="79">punctum</emphasis>
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4595/1); Abastumani,
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(
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of
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P.
<taxonomicName id="4CB54D4BF813FFFF803FB005FA55FE45" box="[1324,1436,367,392]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">substriata</taxonomicName>
</emphasis>
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var.
<emphasis id="B9C1EADAF813FFFF85DBB0FFFF37FE61" box="[200,254,405,428]" italics="true" pageId="4" pageNumber="79">mitis</emphasis>
</taxonomicName>
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2282/1, 2289/2); Tabatsquri,
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(
<typeStatus id="540E886AF813FFFF8649B0FEFC77FE61" box="[858,958,404,428]" pageId="4" pageNumber="79" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF813FFFF86F7B0FFFBAFFE61" box="[996,1126,405,428]" italics="true" pageId="4" pageNumber="79">P. pygmaea</emphasis>
<taxonomicName id="4CB54D4BF813FFFF817EB0FFFB3CFE60" box="[1133,1269,404,429]" kingdom="Animalia" pageId="4" pageNumber="79" rank="variety" variety="nitidula">
var.
<emphasis id="B9C1EADAF813FFFF818DB0FEFB3CFE60" box="[1182,1269,404,429]" italics="true" pageId="4" pageNumber="79">nitidula</emphasis>
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ZMZ 514689/ 2;
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of
<emphasis id="B9C1EADAF813FFFF842FB0D3FE64FE1D" box="[316,429,441,464]" italics="true" pageId="4" pageNumber="79">
P.
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</emphasis>
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2029/1, 2030/10); Kvemo Kartli region: Manglisi,
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(
<typeStatus id="540E886AF813FFFF81EEB0D2FAA8FE1D" box="[1277,1377,440,464]" pageId="4" pageNumber="79" type="syntype">syntypes</typeStatus>
of
<emphasis id="B9C1EADAF813FFFF8094B0D3FF25FE39" italics="true" pageId="4" pageNumber="79">
P.
<taxonomicName id="4CB54D4BF813FFFF8584B0B7FF25FE39" box="[151,236,477,500]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="sieversi">sieversi</taxonomicName>
</emphasis>
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var.
<emphasis id="B9C1EADAF813FFFF8437B0B7FE4DFE39" box="[292,388,477,500]" italics="true" pageId="4" pageNumber="79">punctum</emphasis>
</taxonomicName>
<collectionCode id="EDA4AE0DF813FFFF8498B0B6FE0AFE39" LSID="urn:lsid:biocol.org:col:34838" box="[395,451,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="79">SMF</collectionCode>
2125/1, 245965/3).
</paragraph>
<paragraph id="8B0A36C8F813FFFF85D4B095FBCAFAFD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<emphasis id="B9C1EADAF813FFFF85D4B095FE73FDD5" bold="true" box="[199,442,511,536]" pageId="4" pageNumber="79">Additional material.</emphasis>
<collectingCountry id="F3A27658F813FFFF84D1B095FDC6FDD5" box="[450,527,511,536]" name="Russia" pageId="4" pageNumber="79">Russia</collectingCountry>
, Krasnodar kray: debris of Psekups river (
<bibRefCitation id="EF244B39F813FFFF86FEB36AFBB0FDD5" author="Rosen" box="[1005,1145,512,536]" pageId="4" pageNumber="79" refString="Rosen, O. von (1907) Zum Polymorphismus von Helix vulgaris Rossm. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft, 39, 198 - 206." type="journal article" year="1907">Rosen 1907</bibRefCitation>
); Lagonaki plateau, rocks behind Hotel Azish-Tau near Pereval Azishsky,
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<geoCoordinate id="EE81500FF813FFFF8624B34EFC72FDF1" box="[823,955,548,572]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="40.01361">40°00'49&quot;E</geoCoordinate>
,
<quantity id="4C4D9B2DF813FFFF86D6B34EFBD3FDF1" box="[965,1050,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="4" pageNumber="79" unit="m" value="1770.0">1770 m</quantity>
a.s.l. (
<collectionCode id="EDA4AE0DF813FFFF8171B34EFB6BFDF1" box="[1122,1186,548,572]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
<date id="FF0B1008F813FFFF81BBB34EFB34FDF0" box="[1192,1277,548,573]" pageId="4" pageNumber="79" value="10-10-01">100110</date>
/2).—Adygea: Lagonaki plateau, source of the Kurdzips River,
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<geoCoordinate id="EE81500FF813FFFF862BB32DFC5BFDAD" box="[824,914,583,608]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.983334">39°59'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF86B7B322FC1DFDAD" box="[932,980,584,608]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
37/1); Lagonaki plateau, source of the Kurdzips River,
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<geoCoordinate id="EE81500FF813FFFF84A4B306FDD8FD48" box="[439,529,620,645]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.983334">39°59'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8732B306FD98FD49" box="[545,593,620,644]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
36/15); Lagonaki plateau, source of the Kurdzips River,
<geoCoordinate id="EE81500FF813FFFF81CDB306FAF3FD49" box="[1246,1338,620,644]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="44.05">44°03'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8050B306FA55FD49" box="[1347,1436,620,644]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="40.016666">40°01'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF858DB3FAFF07FD65" box="[158,206,656,680]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
35/6, 41/9); Lagonaki plateau,
<geoCoordinate id="EE81500FF813FFFF8727B3FAFD59FD65" box="[564,656,656,680]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="44.05">44°03'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF878BB3E5FD3DFD65" box="[664,756,655,680]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.983334">39°59'E</geoCoordinate>
,
<quantity id="4C4D9B2DF813FFFF8613B3FAFC9FFD6A" box="[768,854,656,680]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="79" unit="m" value="1800.0">1800 m</quantity>
a.s.l. (
<collectionCode id="EDA4AE0DF813FFFF86B1B3FAFC1BFD65" box="[930,978,656,680]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
23/1); Lagonaki plateau, Murzikal, near the Triangulation station,
<geoCoordinate id="EE81500FF813FFFF84D4B3DEFDEDFD01" box="[455,548,692,716]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="44.033333">44°02'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8723B3DEFD45FD00" box="[560,652,692,717]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.983334">39°59'E</geoCoordinate>
,
<quantity id="4C4D9B2DF813FFFF878EB3DEFD31FD01" box="[669,760,692,716]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="4" pageNumber="79" unit="m" value="2180.0">2180 m</quantity>
a.s.l. (
<collectionCode id="EDA4AE0DF813FFFF865FB3DEFCB5FD01" box="[844,892,692,716]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
34/9); Oshten Fisht mountains (
<collectionCode id="EDA4AE0DF813FFFF8012B3DEFAF2FD01" LSID="urn:lsid:biocol.org:col:34838" box="[1281,1339,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="79">SMF</collectionCode>
2123/3, voucher to
<bibRefCitation id="EF244B39F813FFFF840BB3B2FE77FD3D" author="Boettger" box="[280,446,728,752]" pageId="4" pageNumber="79" refString="Boettger, O. (1889) Zehntes Verzeichnis (XII) von Mollusken der Kaukasuslander. Berichte der senckenbergischen naturforschenden Gesellschaft, 1888 / 1889, 3 - 37, pl. 1." type="journal article" year="1889">Boettger 1889</bibRefCitation>
); Oshten Fisht mountains, glacial moraine between Oshten and Pshekha-su mountains,
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<geoCoordinate id="EE81500FF813FFFF8411B396FE92FCD8" box="[258,347,764,789]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.883335">39°53'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8462B396FE68FCD9" box="[369,417,764,788]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
40/4); Oshten Fisht mountains, high-mountain valley between Fisht and Pshekha-su mountains,
<geoCoordinate id="EE81500FF813FFFF840EB275FEB0FCF5" box="[285,377,799,824]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="43.966667">43°58'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8490B275FE14FCF5" box="[387,477,799,824]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.9">39°54'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF84FDB24AFDD7FCF5" box="[494,542,800,824]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
39/1); Oshten Fisht mountains, SE slope of the Armyansky mountain range,
<geoCoordinate id="EE81500FF813FFFF8584B22EFF3AFC90" box="[151,243,836,861]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="43.966667">43°58'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8413B22EFE90FC90" box="[256,345,836,861]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="39.95">39°57'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF847EB22EFE54FC91" box="[365,413,836,860]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
38/1); Guzeripl
<quantity id="4C4D9B2DF813FFFF8774B22EFD7AFC90" box="[615,691,836,861]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="79" unit="km" value="10.0">10 km</quantity>
towards Abago Pasture,
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<geoCoordinate id="EE81500FF813FFFF8159B22EFB6DFC91" box="[1098,1188,836,860]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="40.2">40°12'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF81A4B22EFB2EFC91" box="[1207,1255,836,860]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
18/22); Abago Pasture, birch forest, near lodge &quot;
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&quot;,
<geoCoordinate id="EE81500FF813FFFF8793B20DFD15FC4D" box="[640,732,871,896]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="43.933334">43°56'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF87F5B202FCF6FC4D" box="[742,831,872,896]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="40.216667">40°13'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8643B202FC49FC4D" box="[848,896,872,896]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
19/48).—Karachayevo-Cherkesiya: Nizhnyaya Teberda,
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<geoCoordinate id="EE81500FF813FFFF847FB2E6FE0CFC68" box="[364,453,908,933]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="41.883335">41°53'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF84CBB2E6FDD1FC69" box="[472,536,908,932]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
80615/7); Arkhyz, near the mouth of Baritov river,
<geoCoordinate id="EE81500FF813FFFF8199B2E6FAC6FC69" box="[1162,1295,908,932]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="43.575">43°34'30”N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8009B2E6FA55FC68" box="[1306,1436,908,933]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="41.275">41°16'30”E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF858DB2DAFF3EFC05" box="[158,247,944,968]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="79">ZMMU</collectionCode>
Lc-27359/2);
<quantity id="4C4D9B2DF813FFFF8488B2DAFE2BFC05" box="[411,482,943,968]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="79" unit="km" value="21.0">21 km</quantity>
SW of Arkhyz,
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SW of the confluence of Psysh and Aman-auz rivers,
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(
<collectionCode id="EDA4AE0DF813FFFF8417B2BEFE94FC21" box="[260,349,980,1004]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="79">ZMMU</collectionCode>
Lc-27356/5, vouchers to
<bibRefCitation id="EF244B39F813FFFF8787B2BEFC02FC21" author="Sysoev" box="[660,971,980,1005]" pageId="4" pageNumber="79" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2009) Land snails and slugs of Russia and adjacent countries. Pensoft, Sofia and Moscow, 312 pp., 142 pls." type="book" year="2009">Sysoev &amp; Schileyko 2009</bibRefCitation>
, as
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<emphasis id="B9C1EADAF813FFFF86ECB2BFFB59FC20" box="[1023,1168,980,1005]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
;
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Lc-27378/3); Dombay valley in direction to Alibek glacier,
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(
<collectionCode id="EDA4AE0DF813FFFF867CB292FC66FBDD" box="[879,943,1016,1040]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
80617/3); Teberda,
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(
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2132/3, voucher to
<bibRefCitation id="EF244B39F813FFFF8467B577FDDEFBF9" author="Boettger" box="[372,535,1052,1077]" pageId="4" pageNumber="79" refString="Boettger, O. (1883) Siebentes Verzeichniss von Mollusken der Kaukasuslander nach Sendungen des Hrn. Hans Leder, z. Z. in Helenendorf bei Elisabetpol (Transkaukasien). Jahrbucher der deutschen malakozoologischen Gesellschaft, 10, 135 - 200, pl. 4 - 7." type="journal article" year="1883">Boettger 1883</bibRefCitation>
;
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54027/2, 54028/3;
<collectionCode id="EDA4AE0DF813FFFF8629B577FCA0FBF9" box="[826,873,1053,1076]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
1/1, voucher to
<bibRefCitation id="EF244B39F813FFFF8137B577FB0EFBF9" author="Boettger" box="[1060,1223,1052,1077]" pageId="4" pageNumber="79" refString="Boettger, O. (1883) Siebentes Verzeichniss von Mollusken der Kaukasuslander nach Sendungen des Hrn. Hans Leder, z. Z. in Helenendorf bei Elisabetpol (Transkaukasien). Jahrbucher der deutschen malakozoologischen Gesellschaft, 10, 135 - 200, pl. 4 - 7." type="journal article" year="1883">Boettger 1883</bibRefCitation>
;
<collectionCode id="EDA4AE0DF813FFFF81C7B577FACBFBF9" box="[1236,1282,1053,1076]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
2/2, 3/1, 4/1; ZMZ 514740/2); Teberda valley, Madniskhevi,
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(SCZENASRA/6).—Kabardino-Balkariya: Zhankhoteko,
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<geoCoordinate id="EE81500FF813FFFF848DB50EFE3EFBB1" box="[414,503,1124,1148]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="42.2">42°12'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8715B50EFDFFFBB1" box="[518,566,1124,1148]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
29/1); Shashbavat gorge below Tyrnyauz,
<geoCoordinate id="EE81500FF813FFFF8108B50EFBBEFBB0" box="[1051,1143,1124,1149]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="43.416668">43°25'N</geoCoordinate>
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(
<collectionCode id="EDA4AE0DF813FFFF81F4B50EFAEEFBB1" box="[1255,1319,1124,1148]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
80616/1); Bukhaidze,
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<geoCoordinate id="EE81500FF813FFFF8493B5EDFE10FB6D" box="[384,473,1159,1184]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="42.75">42°45'E</geoCoordinate>
(
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30/1); Itkol, forest in gorge at Mount Cheget,
<geoCoordinate id="EE81500FF813FFFF810DB5EDFBB3FB6D" box="[1054,1146,1159,1184]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="43.25">43°15'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8192B5E2FB13FB6D" box="[1153,1242,1160,1184]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="42.566666">42°34'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF81FBB5E2FAE1FB6D" box="[1256,1320,1160,1184]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
80613/2); lower Adylsu valley,
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<geoCoordinate id="EE81500FF813FFFF84E9B5C6FD9AFB09" box="[506,595,1196,1220]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="42.65">42°39'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8775B5C6FD6FFB09" box="[614,678,1196,1220]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
80612/13); Chegem gorge
<quantity id="4C4D9B2DF813FFFF86F9B5C6FBEEFB08" box="[1002,1063,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="79" unit="km" value="5.0">5 km</quantity>
S of Verkhniy Chegem village,
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<geoCoordinate id="EE81500FF813FFFF85ECB5BAFEB7FB25" box="[255,382,1232,1256]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="43.075">43°04'30''E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF8481B5BAFD0EFB25" author="Sysoev" box="[402,711,1231,1256]" pageId="4" pageNumber="79" refString="Sysoev, A. V. &amp; Schileyko, A. A. (2009) Land snails and slugs of Russia and adjacent countries. Pensoft, Sofia and Moscow, 312 pp., 142 pls." type="book" year="2009">Sysoev &amp; Schileyko 2009</bibRefCitation>
).—North Ossetia: Nizhniy Nar,
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<geoCoordinate id="EE81500FF813FFFF81A3B5BAFAC3FB25" box="[1200,1290,1232,1256]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.783333">43°47'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF800EB5BAFA84FB25" box="[1309,1357,1232,1256]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
31/1); Dzinaga,
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<geoCoordinate id="EE81500FF813FFFF8462B59EFE03FAC1" box="[369,458,1268,1292]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.7">43°42'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF84CDB59EFDFEFAC1" box="[478,567,1268,1292]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="79">ZMMU</collectionCode>
Lc-27357/5); Chmi,
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<geoCoordinate id="EE81500FF813FFFF868CB59EFC31FAC1" box="[927,1016,1268,1292]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.633335">44°38'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF811FB59EFBF5FAC1" box="[1036,1084,1268,1292]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
22/20).—Ingushetia: Armkhi,
<geoCoordinate id="EE81500FF813FFFF8584B472FF3DFAFD" box="[151,244,1304,1328]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="42.8">42°48'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF85E9B472FE9AFAFD" box="[250,339,1304,1328]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.716667">44°43'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8471B472FE5BFAFD" box="[354,402,1304,1328]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
28/2).—Chechnya: Lake Kezenoyam (
<collectionCode id="EDA4AE0DF813FFFF8659B472FCB3FAFD" box="[842,890,1304,1328]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
32/2, 33/1).
</paragraph>
<paragraph id="8B0A36C8F813FFFF85D4B456FB71F811" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<collectingCountry id="F3A27658F813FFFF85D4B456FEEDFA98" box="[199,292,1340,1365]" name="Georgia" pageId="4" pageNumber="79">Georgia</collectingCountry>
, Abkhazia: Sukhumi Military Road, near Mount Pal, Bogatskiye Skali,
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<geoCoordinate id="EE81500FF813FFFF81CDB456FAF1FA99" box="[1246,1336,1340,1364]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="41.366665">41°22'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8059B456FAB3FA99" box="[1354,1402,1340,1364]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
7/ 2).—Imereti region: Nakerala pass,
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(
<collectionCode id="EDA4AE0DF813FFFF862CB40AFC6AFAB5" box="[831,931,1376,1400]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="79">NHMW</collectionCode>
/1, voucher to
<bibRefCitation id="EF244B39F813FFFF812EB435FAF8FAB5" author="Pokryszko" box="[1085,1329,1375,1400]" pageId="4" pageNumber="79" refString="Pokryszko, B. M., Cameron, R. A. D., Mumladze, L. &amp; Tarknishvili, D. (2011) Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society, 102, 239 - 250. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2010.01575. x" type="journal article" year="2011">
Pokryszko
<emphasis id="B9C1EADAF813FFFF81A9B40BFB23FAB5" box="[1210,1258,1375,1400]" italics="true" pageId="4" pageNumber="79">et al</emphasis>
. 2011
</bibRefCitation>
).—Shida Kartli region: Tqemlovana,
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<geoCoordinate id="EE81500FF813FFFF8748B4EEFD13FA51" box="[603,730,1412,1436]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="43.716667">43°43'00''E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF87FAB4EEFB81FA50" author="Lezhawa" box="[745,1096,1412,1437]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">
Lezhawa 1973, as
<taxonomicName id="4CB54D4BF813FFFF86AFB4EFFB81FA50" box="[956,1096,1412,1437]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF86AFB4EFFB81FA50" box="[956,1096,1412,1437]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
</bibRefCitation>
).—Mtskheta-Mtianeti region: Kazbek (ZMZ 514683/2); valley of Andaki river 2.2 km upstream of its confluence with the Argun river,
<geoCoordinate id="EE81500FF813FFFF8584B4A6FED0FA29" box="[151,281,1484,1508]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="42.652225">42°39'08''N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8433B4A6FE57FA28" box="[288,414,1484,1509]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="45.191113">45°11'28''E</geoCoordinate>
(FW 36/5); Stepantsminda (= Kazbegi),
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<geoCoordinate id="EE81500FF813FFFF86DFB4A6FBECFA29" box="[972,1061,1484,1508]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.65">44°39'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8127B4A6FBADFA29" box="[1076,1124,1484,1508]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
26/5); gorge of Argun river between Shatili and Georgitsminda,
<geoCoordinate id="EE81500FF813FFFF8750B485FD0CF9C5" box="[579,709,1519,1544]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="42.6375">42°38'15''N</geoCoordinate>
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(FW 37/1); Sioni,
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<geoCoordinate id="EE81500FF813FFFF81B1B485FB35F9C5" box="[1186,1276,1519,1544]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.583332">44°35'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF801CB49AFAF6F9C5" box="[1295,1343,1520,1544]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
25/23); Kobi,
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<geoCoordinate id="EE81500FF813FFFF8443B77EFE60F9E1" box="[336,425,1556,1580]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.5">44°30'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF84ACB77EFE26F9E1" box="[447,495,1556,1580]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
27/25); Kumlistsikhe,
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<geoCoordinate id="EE81500FF813FFFF8667B77EFC04F9E1" box="[884,973,1556,1580]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.483334">44°29'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF86F7B77EFBDDF9E1" box="[996,1044,1556,1580]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
21/8).—Tbilisi: Tbilisi,
<geoCoordinate id="EE81500FF813FFFF802CB77EFA55F9E1" box="[1343,1436,1556,1580]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.7">41°42'N</geoCoordinate>
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(
<collectionCode id="EDA4AE0DF813FFFF85ECB752FEF0F99D" LSID="urn:lsid:biocol.org:col:34838" box="[255,313,1592,1616]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="79">SMF</collectionCode>
54026/1).—Kakheti region: Dzhvarboseli,
<geoCoordinate id="EE81500FF813FFFF8632B75DFCB4F99D" box="[801,893,1591,1616]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="42.416668">42°25'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8697B75DFC17F99D" box="[900,990,1591,1616]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="45.5">45°30'E</geoCoordinate>
(IUC 388/n, voucher to
<bibRefCitation id="EF244B39F813FFFF81E2B75DFA51F99D" author="Lezhawa" box="[1265,1432,1591,1616]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1962) K izucheniyu nazemnoy malakofauny gornoy Tusheti. Soobshcheniya Akademii Nauk Gruzinskoy SSR, 29, 327 - 332." type="journal article" year="1962">Lezhawa 1962</bibRefCitation>
,
<bibRefCitation id="EF244B39F813FFFF8584B736FF1BF9B9" author="Lezhawa" box="[151,210,1628,1652]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">1973</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF85EEB737FE43F9B8" box="[253,394,1628,1653]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF85EEB737FE43F9B8" box="[253,394,1628,1653]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
); Verkhne Omalo: banks of Tushetski Alazani river,
<geoCoordinate id="EE81500FF813FFFF86F7B736FB88F9B9" box="[996,1089,1628,1652]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="42.366665">42°22'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF815BB736FB6BF9B8" box="[1096,1186,1628,1653]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="45.633335">45°38'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF81A1B736FECEF955" author="Lezhawa" pageId="4" pageNumber="79" refString="Lezhawa, G. (1962) K izucheniyu nazemnoy malakofauny gornoy Tusheti. Soobshcheniya Akademii Nauk Gruzinskoy SSR, 29, 327 - 332." type="journal article" year="1962">
Lezhawa 1962, as
<taxonomicName id="4CB54D4BF813FFFF8094B737FECEF955" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF8094B737FECEF955" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
</bibRefCitation>
); Akhmeta,
<geoCoordinate id="EE81500FF813FFFF848FB7EAFE30F955" box="[412,505,1664,1688]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="42.033333">42°02'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8717B715FD94F955" box="[516,605,1663,1688]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="45.2">45°12'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF8763B715FC16F955" author="Lezhawa" box="[624,991,1663,1688]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">
Lezhawa 1973, as
<taxonomicName id="4CB54D4BF813FFFF865CB7EBFC16F955" box="[847,991,1663,1688]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF865CB7EBFC16F955" box="[847,991,1663,1688]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
</bibRefCitation>
); Qvareli,
<geoCoordinate id="EE81500FF813FFFF8172B715FB74F955" box="[1121,1213,1663,1688]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.95">41°57'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF81DAB715FAEBF955" box="[1225,1314,1663,1688]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="45.816666">45°49'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF8026B715FE40F970" author="Lezhawa" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">
Lezhawa 1973, as
<taxonomicName id="4CB54D4BF813FFFF85EEB7CFFE40F970" box="[253,393,1700,1725]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF85EEB7CFFE40F970" box="[253,393,1700,1725]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
</bibRefCitation>
); Lagodekhi,
<geoCoordinate id="EE81500FF813FFFF873AB7CEFD4CF970" box="[553,645,1700,1725]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.9">41°54'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF879EB7CEFD2FF970" box="[653,742,1700,1725]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="46.333332">46°20'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF87E6B7CEFCECF971" box="[757,805,1700,1724]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
8/1).—Samtskhe-Javakheti region: Borjomi, Banikhevi valley,
<geoCoordinate id="EE81500FF813FFFF85FFB7ADFE80F92D" box="[236,329,1735,1760]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.866665">41°52'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8446B7A2FE67F92D" box="[341,430,1736,1760]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.383335">43°23'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF84D2B7A2FE38F92D" box="[449,497,1736,1760]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
5/1); Abastumani,
<geoCoordinate id="EE81500FF813FFFF87CBB7ADFCFDF92D" box="[728,820,1735,1760]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.75">41°45'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8653B7ADFC50F92D" box="[832,921,1735,1760]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="42.833332">42°50'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF86BEB7A2FC2EF92D" LSID="urn:lsid:biocol.org:col:34838" box="[941,999,1736,1760]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="79">SMF</collectionCode>
2127/3, voucher to
<bibRefCitation id="EF244B39F813FFFF81C9B7A2FA46F92D" author="Boettger" box="[1242,1423,1735,1760]" pageId="4" pageNumber="79" refString="Boettger, O. (1886 a) Die Binnenmollusken des Talysch-Gebietes. Nach neuesten Materialien bearbeitet. In: Radde, G. (Ed.), Die Fauna und Flora des sudwestlichen Caspi-Gebietes. Wissenschaftliche Beitrage zu den Reisen an der persischrussischen Grenze. Brockhaus, Leipzig, pp. 255 - 350, pls. 2 - 3. http: // dx. doi. org / 10.5962 / bhl. title. 36963" type="book chapter" year="1886" yearSuffix="a">Boettger 1886a</bibRefCitation>
); Sapara Monastery,
<geoCoordinate id="EE81500FF813FFFF8463B786FE04F8C8" box="[368,461,1772,1797]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.6">41°36'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF84C6B786FDE7F8C9" box="[469,558,1772,1796]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.033333">43°02'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF872DB786FDA7F8C9" box="[574,622,1772,1796]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
20/1); Vardzia
<quantity id="4C4D9B2DF813FFFF860CB786FC93F8C8" box="[799,858,1772,1797]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="79" unit="km" value="2.0">2 km</quantity>
towards Tmogvi,
<geoCoordinate id="EE81500FF813FFFF8139B786FB67F8C8" box="[1066,1198,1772,1797]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="41.38222">41°22'56&quot;N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF81A5B786FAF0F8C8" box="[1206,1337,1772,1797]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="43.2975">43°17'51&quot;E</geoCoordinate>
,
<quantity id="4C4D9B2DF813FFFF8055B786FA55F8C9" box="[1350,1436,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="4" pageNumber="79" unit="m" value="1240.0">1240 m</quantity>
a.s.l. (
<collectionCode id="EDA4AE0DF813FFFF85CFB67AFED5F8E5" box="[220,284,1808,1832]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="79">ZMH</collectionCode>
<date id="FF0B1008F813FFFF8435B665FEB6F8E5" box="[294,383,1807,1832]" pageId="4" pageNumber="79" value="87-10-06">100687</date>
/3); canyon of Kura river near Vardzia,
<geoCoordinate id="EE81500FF813FFFF865AB67AFC6CF8E5" box="[841,933,1808,1832]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.366665">41°22'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF86BCB665FBC1F8E5" box="[943,1032,1807,1832]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.25">43°15'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF8108B67AFB82F8E5" box="[1051,1099,1808,1832]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="79">ZIN</collectionCode>
6/1).—Kvemo Kartli region: Martqopi,
<geoCoordinate id="EE81500FF813FFFF841EB65EFEA0F881" box="[269,361,1844,1868]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.783333">41°47'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8463B65EFE00F880" box="[368,457,1844,1869]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="45.016666">45°01'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF84CBB65EFDB4F881" author="Boettger" box="[472,637,1844,1868]" pageId="4" pageNumber="79" refString="Boettger, O. (1880) Kaukasische Mollusken, gesammelt von Herrn Hans Leder, z. Z. in Tiflis. Jahrbucher der deutschen malakozoologischen Gesellschaft, 7, 109 - 150, pl. 4." type="journal article" year="1880">Boettger 1880</bibRefCitation>
); Manglisi,
<geoCoordinate id="EE81500FF813FFFF8611B65EFC97F881" box="[770,862,1844,1868]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.7">41°42'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8676B65EFC76F881" box="[869,959,1844,1868]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.383335">44°23'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF813FFFF86DEB65EFBCEF881" LSID="urn:lsid:biocol.org:col:34838" box="[973,1031,1844,1868]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="79">SMF</collectionCode>
2124/51;
<collectionCode id="EDA4AE0DF813FFFF816BB65EFB04F881" box="[1144,1229,1844,1868]" httpUri="http://grbio.org/cool/2k0w-p5dh" name="Zoological Institute, Polish Academy of Sciences" pageId="4" pageNumber="79">IZPAN</collectionCode>
); Tsqneti,
<geoCoordinate id="EE81500FF813FFFF802CB65EFA55F881" box="[1343,1436,1844,1868]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.7">41°42'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8584B632FF39F8BD" box="[151,240,1880,1904]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.7">44°42'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF85ECB63DFE61F8BD" author="Lezhawa" box="[255,424,1879,1904]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
,
<bibRefCitation id="EF244B39F813FFFF84A7B632FE26F8BD" author="Lezhawa" box="[436,495,1880,1904]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">1973</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF870AB633FD6CF8BD" box="[537,677,1879,1904]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF870AB633FD6CF8BD" box="[537,677,1879,1904]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
); Betania monastery,
<geoCoordinate id="EE81500FF813FFFF868FB632FBD7F8BD" box="[924,1054,1880,1904]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="41.69167">41°41'30''N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF8135B63DFB6CF8BD" box="[1062,1189,1879,1904]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="44.61111">44°36'40''E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF81A7B63DFECEF858" author="Lezhawa" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">
Lezhawa 1966, as
<taxonomicName id="4CB54D4BF813FFFF8094B633FECEF858" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF8094B633FECEF858" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
</bibRefCitation>
); Rekha,
<geoCoordinate id="EE81500FF813FFFF8466B616FE1BF859" box="[373,466,1916,1940]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.683334">41°41'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF84CAB616FDFBF858" box="[473,562,1916,1941]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="43.85">43°51'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF8752B616FD23F858" author="Lezhawa" box="[577,746,1916,1941]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
,
<bibRefCitation id="EF244B39F813FFFF87E5B616FCF8F859" author="Lezhawa" box="[758,817,1916,1940]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">1973</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF8648B617FC2EF858" box="[859,999,1916,1941]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF8648B617FC2EF858" box="[859,999,1916,1941]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
); Kojori
<geoCoordinate id="EE81500FF813FFFF8143B616FB65F859" box="[1104,1196,1916,1940]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.666668">41°40'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF81A7B616FAC4F859" box="[1204,1293,1916,1940]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.7">44°42'E</geoCoordinate>
(IUC 403/2, voucher to
<bibRefCitation id="EF244B39F813FFFF8407B6CAFE72F874" author="Lezhawa" box="[276,443,1952,1977]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF84F7B6CBFDA6F874" box="[484,623,1952,1977]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF84F7B6CBFDA6F874" box="[484,623,1952,1977]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
); Akhalsopeli,
<geoCoordinate id="EE81500FF813FFFF860FB6CAFC56F875" box="[796,927,1952,1976]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="15" value="41.65">41°39'00''N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF86B5B6CAFBECF875" box="[934,1061,1952,1976]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="15" value="44.341667">44°20'30''E</geoCoordinate>
(IUC 394/4, voucher to
<bibRefCitation id="EF244B39F813FFFF8024B6CAFF1BF810" author="Lezhawa" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
,
<bibRefCitation id="EF244B39F813FFFF85CEB6AEFED1F811" author="Lezhawa" box="[221,280,1988,2012]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">1973</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF8452B6AFFE05F810" box="[321,460,1988,2013]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF8452B6AFFE05F810" box="[321,460,1988,2013]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
); Kiketi,
<geoCoordinate id="EE81500FF813FFFF872AB6AEFD5CF811" box="[569,661,1988,2012]" direction="north" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="41.65">41°39'N</geoCoordinate>
<geoCoordinate id="EE81500FF813FFFF878FB6AEFD3CF811" box="[668,757,1988,2012]" direction="east" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="44.65">44°39'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F813FFFF8617B6AEFC64F810" author="Lezhawa" box="[772,941,1988,2013]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1966) Nazemnye mollyuski. In: Fauna bespozvonochnykh Trialetskogo khrebta, Tbilisi, pp. 9 - 20." type="book chapter" year="1966">Lezhawa 1966</bibRefCitation>
,
<bibRefCitation id="EF244B39F813FFFF86ABB6AEFC3AF811" author="Lezhawa" box="[952,1011,1988,2012]" pageId="4" pageNumber="79" refString="Lezhawa, G. (1973) Khmeletis moluskebi. In: Sakartvelis tskhovelta samqaro IV. Metsniereba, Tbilisi, pp. 20 - 68." type="book chapter" year="1973">1973</bibRefCitation>
, as
<taxonomicName id="4CB54D4BF813FFFF810FB6AFFB61F810" box="[1052,1192,1988,2013]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="79" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="B9C1EADAF813FFFF810FB6AFFB61F810" box="[1052,1192,1988,2013]" italics="true" pageId="4" pageNumber="79">V. substriata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B0A36C8F812FFFE85D4B1FDFEA3FE39" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="80">
<collectingCountry id="F3A27658F812FFFE85D4B1FDFEE5FF7D" box="[199,300,151,176]" name="Armenia" pageId="5" pageNumber="80">Armenia</collectingCountry>
, Shirak Province: Ashotsk (= Gukasyan), Akhuryan gorge,
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<geoCoordinate id="EE81500FF812FFFE8173B1FDFB16FF7D" box="[1120,1247,151,176]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="43.858334">43°51'30''E</geoCoordinate>
(SCZENASRA/ 11); Amasia,
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<geoCoordinate id="EE81500FF812FFFE8485B1D6FE26FF19" box="[406,495,188,212]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="43.783333">43°47'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F812FFFE84ECB1D6FD18FF18" author="Akramowski" box="[511,721,188,213]" pageId="5" pageNumber="80" refString="Akramowski, N. N. (1976) Fauna Armyanskoy SSR. Mollyuski (Mollusca). Akademiya nauk Armyanskoy SSR, Institut Zoologii, Yerevan, 268 pp., 16 pls." type="book" year="1976">Akramowski 1976</bibRefCitation>
).—Tavush Province: Ijevan,
<geoCoordinate id="EE81500FF812FFFE8132B1D6FBB4FF18" box="[1057,1149,188,213]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.883335">40°53'N</geoCoordinate>
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(SCZENASRA/ 2); Ijevan, Tissovaya Forest (SCZENASRA/1); Gandzakar: middle part of S slope of Mount Budur,
<geoCoordinate id="EE81500FF812FFFE800AB1B5FA55FF35" box="[1305,1436,223,248]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="15" value="40.841667">40°50'30''N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8584B06EFEDFFED0" box="[151,278,260,285]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="45.175003">45°10'30''E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE8439B06EFE4AFED1" box="[298,387,260,284]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="80">ZMMU</collectionCode>
Lc-27381/2); Dilijan, left bank of Agstafa river (
<collectionCode id="EDA4AE0DF812FFFE86C7B06EFBE4FED1" box="[980,1069,260,284]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="80">ZMMU</collectionCode>
Lc-7654/11, 7655/26); Dilijan, Shamakhyan,
<geoCoordinate id="EE81500FF812FFFE845EB04DFE06FE8D" box="[333,463,295,320]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="15" value="40.7625">40°45'45''N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE84FEB04DFDA4FE8D" box="[493,621,295,320]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="45.83611">45°50'10''E</geoCoordinate>
(SCZENASRA/1); Dilijan, Papanino,
<geoCoordinate id="EE81500FF812FFFE816FB042FB36FE8D" box="[1148,1279,296,320]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="15" value="40.74167">40°44'30''N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE800EB04DFA55FE8D" box="[1309,1436,295,320]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="45.845833">45°50'45''E</geoCoordinate>
(SCZENASRA/2).—Kotayk Province: Tsaghkadzor, Mount Teghenis,
<geoCoordinate id="EE81500FF812FFFE86FFB026FB81FEA9" box="[1004,1096,332,356]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.533333">40°32'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8148B026FB7DFEA9" box="[1115,1204,332,356]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="44.65">44°39'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE81DDB026FAC7FEA9" box="[1230,1294,332,356]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="5" pageNumber="80">ZMH</collectionCode>
80614/22); Tsaghkadzor,
<geoCoordinate id="EE81500FF812FFFE8453B01AFE54FE45" box="[320,413,368,392]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.533333">40°32'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE84A1B01AFDC2FE45" box="[434,523,368,392]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="44.716667">44°43'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F812FFFE8734B005FCCCFE45" author="Akramowski" box="[551,773,367,392]" pageId="5" pageNumber="80" refString="Akramowski, N. N. (1968) Nekotorye dopolneniya k faune mollyuskov Armenii. Vklyuchaya novosti dlya fauny Zakavkazya i SSSR. In: Mollyuski i ikh pol v yekosistemakh. Avtoreferaty dokladov, Nauka, Leningrad, 150 pp. [pp. 26 - 27]" type="book chapter" year="1968">Akramowski 1968</bibRefCitation>
).—Gegharkunik Province: Tsapatagh (= Babadzhan),
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<geoCoordinate id="EE81500FF812FFFE8430B0FEFE6BFE60" box="[291,418,404,429]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="45.475002">45°28'30''E</geoCoordinate>
(SCZENASRA/1).—Ararat Province: Khosrov Forest,
<geoCoordinate id="EE81500FF812FFFE810CB0FEFBB5FE61" box="[1055,1148,404,428]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.033333">40°02'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8196B0FEFB17FE60" box="[1157,1246,404,429]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="44.916668">44°55'E</geoCoordinate>
(SCZENASRA/ 1).—Syunik Province: Tatev,
<geoCoordinate id="EE81500FF812FFFE84F6B0D2FDA2FE1D" box="[485,619,440,464]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="15" value="39.383335">39°23'00”N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8761B0DDFD3DFE1D" box="[626,756,439,464]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="46.24167">46°14'30”E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE8610B0D2FC95FE1D" box="[771,860,440,464]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="80">ZMMU</collectionCode>
Lc-22197/1, 27358/10); Kajaran,
<geoCoordinate id="EE81500FF812FFFE81CCB0D2FAF2FE1D" box="[1247,1339,440,464]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="39.15">39°09'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8050B0DDFA55FE1D" box="[1347,1436,439,464]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.15">46°09'E</geoCoordinate>
(SCZENASRA/2).
</paragraph>
<paragraph id="8B0A36C8F812FFFE85D4B095FB16FCD8" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="80">
<collectingCountry id="F3A27658F812FFFE85D4B095FE8BFDD5" box="[199,322,511,536]" name="Azerbaijan" pageId="5" pageNumber="80">Azerbaijan</collectingCountry>
: Zaqatala State Reserve, Mount Ağkiman,
<geoCoordinate id="EE81500FF812FFFE862BB36AFC5DFDD5" box="[824,916,512,536]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="41.816666">41°49'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE868DB095FC3EFDD5" box="[926,1015,511,536]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.516666">46°31'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE811BB36AFBF1FDD5" box="[1032,1080,512,536]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
17/1); Zaqatala State Reserve, Mount Vergetel, near ledge,
<geoCoordinate id="EE81500FF812FFFE84FEB34EFD80FDF1" box="[493,585,548,572]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="41.733334">41°44'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8744B34EFD79FDF0" box="[599,688,548,573]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.433334">46°26'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE87D7B34EFD3DFDF1" box="[708,756,548,572]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
16/27); Qusar,
<geoCoordinate id="EE81500FF812FFFE86A6B34EFBDBFDF0" box="[949,1042,548,573]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="41.433334">41°26'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE810CB34EFBBEFDF0" box="[1055,1143,548,573]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="48.433334">48°26'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE819EB34EFB0EFDF1" LSID="urn:lsid:biocol.org:col:34838" box="[1165,1223,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="80">SMF</collectionCode>
2128, voucher to
<bibRefCitation id="EF244B39F812FFFE8584B322FE80FDAD" author="Boettger" box="[151,329,583,608]" pageId="5" pageNumber="80" refString="Boettger, O. (1886 b) Neuntes Verzeichniss (IX) von Mollusken der Kaukasuslander nach Sendungen des Hrn. Hans Leder, z. Z. in Helenendorf bei Elisabetpol (Transkaukasien). Jahrbucher der deutschen malakozoologischen Gesellschaft, 13, 121 - 156, pl. 3." type="journal article" year="1886" yearSuffix="b">Boettger 1886b</bibRefCitation>
); Şəki, Kiş river,
<geoCoordinate id="EE81500FF812FFFE871EB32DFDA3FDAD" box="[525,618,583,608]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="41.266666">41°16'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8762B322FD02FDAD" box="[625,715,584,608]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="47.2">47°12'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE87C9B322FCC3FDAD" box="[730,778,584,608]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
24/4); western branch of Kürəkçay river near fish-factory,
<geoCoordinate id="EE81500FF812FFFE8584B306FF3DFD48" box="[151,244,620,645]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.433334">40°26'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE85E9B306FE9AFD48" box="[250,339,620,645]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.316666">46°19'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE8472B306FE58FD49" box="[353,401,620,644]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
11/5); road to Lake Göygöl
<quantity id="4C4D9B2DF812FFFE87DDB306FCCEFD48" box="[718,775,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="80" unit="km" value="1.0">1 km</quantity>
before the lake,
<geoCoordinate id="EE81500FF812FFFE86D3B306FB8CFD48" box="[960,1093,620,645]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="15" value="40.425003">40°25'30”N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE815FB306FB04FD48" box="[1100,1229,620,645]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="15" value="46.333332">46°20'00”E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE81CFB306FAC5FD49" box="[1244,1292,620,644]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
14/17); road to Lake Göygöl,
<quantity id="4C4D9B2DF812FFFE8445B3FAFE62FD65" box="[342,427,655,680]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="80" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">
2
<emphasis id="B9C1EADAF812FFFE8477B3E5FEBBFD65" bold="true" box="[356,370,655,680]" pageId="5" pageNumber="80"></emphasis>
3 km
</quantity>
before fish-factory,
<geoCoordinate id="EE81500FF812FFFE8783B3E5FD25FD65" box="[656,748,655,680]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.416668">40°25'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE87E7B3E5FC84FD65" box="[756,845,655,680]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.316666">46°19'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE864FB3FAFC45FD65" box="[860,908,656,680]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
15/115); north shore of Lake Göygöl,
<geoCoordinate id="EE81500FF812FFFE802CB3E5FA55FD65" box="[1343,1436,655,680]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.416668">40°25'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8584B3DEFF39FD00" box="[151,240,692,717]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.333332">46°20'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE8410B3DEFEFAFD01" box="[259,307,692,716]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
13/3); mountain range between Toğanalı and Lake Göygöl,
<geoCoordinate id="EE81500FF812FFFE8112B3DEFB94FD01" box="[1025,1117,692,716]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.4">40°24'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE817AB3DEFB0BFD00" box="[1129,1218,692,717]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.316666">46°19'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE81C6B3DEFACCFD01" box="[1237,1285,692,716]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
9/1); Mount Kəpəz,
<geoCoordinate id="EE81500FF812FFFE85E3B3B2FE85FD3D" box="[240,332,728,752]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="40.35">40°21'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8444B3BDFE78FD3D" box="[343,433,727,752]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.35">46°21'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE84D0B3B2FE3AFD3D" box="[451,499,728,752]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
10/2); mountain Sağsağan Qayası,
<geoCoordinate id="EE81500FF812FFFE8686B3BDFC38FD3D" box="[917,1009,727,752]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="39.966667">39°58'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE86EFB3BDFB9CFD3D" box="[1020,1109,727,752]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="46.6">46°36'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE817BB3B2FB51FD3D" box="[1128,1176,728,752]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="80">ZIN</collectionCode>
12/1); Lerik,
<geoCoordinate id="EE81500FF812FFFE8053B3BDFA55FD3D" box="[1344,1436,727,752]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="38.766666">38°46'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE8584B396FF39FCD8" box="[151,240,764,789]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="48.416668">48°25'E</geoCoordinate>
(
<collectionCode id="EDA4AE0DF812FFFE85ECB396FEF0FCD9" LSID="urn:lsid:biocol.org:col:34838" box="[255,313,764,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="80">SMF</collectionCode>
2126/2, voucher to
<bibRefCitation id="EF244B39F812FFFE870AB397FD00FCD8" author="Boettger" box="[537,713,764,789]" pageId="5" pageNumber="80" refString="Boettger, O. (1886 a) Die Binnenmollusken des Talysch-Gebietes. Nach neuesten Materialien bearbeitet. In: Radde, G. (Ed.), Die Fauna und Flora des sudwestlichen Caspi-Gebietes. Wissenschaftliche Beitrage zu den Reisen an der persischrussischen Grenze. Brockhaus, Leipzig, pp. 255 - 350, pls. 2 - 3. http: // dx. doi. org / 10.5962 / bhl. title. 36963" type="book chapter" year="1886" yearSuffix="a">Boettger 1886a</bibRefCitation>
); Lǝnkǝran,
<geoCoordinate id="EE81500FF812FFFE8646B396FC78FCD8" box="[853,945,764,789]" direction="north" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="38.75">38°45'N</geoCoordinate>
<geoCoordinate id="EE81500FF812FFFE86ABB396FBDBFCD8" box="[952,1042,764,789]" direction="east" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="48.85">48°51'E</geoCoordinate>
(
<bibRefCitation id="EF244B39F812FFFE8133B396FB1BFCD8" author="Boettger" box="[1056,1234,764,789]" pageId="5" pageNumber="80" refString="Boettger, O. (1886 a) Die Binnenmollusken des Talysch-Gebietes. Nach neuesten Materialien bearbeitet. In: Radde, G. (Ed.), Die Fauna und Flora des sudwestlichen Caspi-Gebietes. Wissenschaftliche Beitrage zu den Reisen an der persischrussischen Grenze. Brockhaus, Leipzig, pp. 255 - 350, pls. 2 - 3. http: // dx. doi. org / 10.5962 / bhl. title. 36963" type="book chapter" year="1886" yearSuffix="a">Boettger 1886a</bibRefCitation>
).
</paragraph>
<paragraph id="8B0A36C8F812FFFE85D4B275FBF9FC90" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="80">
<emphasis id="B9C1EADAF812FFFE85D4B275FEE3FCF5" bold="true" box="[199,298,799,824]" pageId="5" pageNumber="80">Habitat.</emphasis>
The species lives mainly in deciduous or mixed mountain forests, but occurs also in fissures of rocks in the subalpine zone above the timberline as well as under
<taxonomicName id="4CB54D4BF812FFFE8620B22EFC11FC90" box="[819,984,836,861]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="B9C1EADAF812FFFE8620B22EFC11FC90" box="[819,984,836,861]" italics="true" pageId="5" pageNumber="80">Rhododendron</emphasis>
</taxonomicName>
shrubs.
</paragraph>
</subSubSection>
<subSubSection id="C3AF6543F812FFFE85D4B20DFE28FC21" pageId="5" pageNumber="80" type="distribution">
<paragraph id="8B0A36C8F812FFFE85D4B20DFE28FC21" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="80">
<emphasis id="B9C1EADAF812FFFE85D4B20DFE75FC4D" bold="true" box="[199,444,871,896]" pageId="5" pageNumber="80">
Distribution (
<figureCitation id="138E2A4DF812FFFE847BB20DFE67FC4D" box="[360,430,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[151,1436,1060,1831]" captionTargetId="figure@5.[151,1436,1060,1832]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Vertigo (Vertigo) nitidula (Mousson) (dots) and Vertigo (Vertigo) substriata (Jeffreys) (stars) in the Caucasus region (0.5 ° grid)." httpUri="https://zenodo.org/record/287229/files/figure.png" pageId="5" pageNumber="80">Fig. 3</figureCitation>
).
</emphasis>
<taxonomicName id="4CB54D4BF812FFFE84D0B203FDBBFC4D" box="[451,626,871,896]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF812FFFE84D0B203FDBBFC4D" box="[451,626,871,896]" italics="true" pageId="5" pageNumber="80">Vertigo nitidula</emphasis>
</taxonomicName>
is widely distributed in the Caucasian region, occurring on both sides of the Great Caucasus as well as in the Lesser Caucasus and the Talysh Mountains. The occurrences in the Lesser Caucasus and in the Talysh Mountains in southern
<collectingCountry id="F3A27658F812FFFE87EBB2C5FCBAFC05" box="[760,883,943,968]" name="Azerbaijan" pageId="5" pageNumber="80">Azerbaijan</collectingCountry>
indicate that the range of
<taxonomicName id="4CB54D4BF812FFFE81A6B2DBFAE5FC05" box="[1205,1324,943,968]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="80" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B9C1EADAF812FFFE81A6B2DBFAE5FC05" box="[1205,1324,943,968]" italics="true" pageId="5" pageNumber="80">V. nitidula</emphasis>
</taxonomicName>
probably extends into
<collectingCountry id="F3A27658F812FFFE8435B2BEFEBFFC20" box="[294,374,980,1005]" name="Turkey" pageId="5" pageNumber="80">Turkey</collectingCountry>
and
<collectingCountry id="F3A27658F812FFFE84BFB2BFFE14FC21" box="[428,477,981,1004]" name="Iran" pageId="5" pageNumber="80">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>