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<document id="26AFE7A85D04E156CA79AFD57601E34B" ID-CLB-Dataset="138346" ID-DOI="10.11646/phytotaxa.561.1.6" ID-GBIF-Dataset="76da0309-11a1-46bb-95bb-75a4412fb0bb" ID-ISSN="1179-3163" ID-Zenodo-Dep="7052827" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1662448843418" checkinUser="plazi" docAuthor="Li, Tian, Xu, Lian-Sheng &amp; Chen, You-Sheng" docDate="2022" docId="03A787F66771FFF1FF75FF47FE11FC69" docLanguage="en" docName="phytotaxa.561.1.6.pdf" docOrigin="Phytotaxa 561 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.561.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Saussurea khunjerabensis Y. S. Chen 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="72" masterDocId="FF9EFF8E6775FFF6FFFDFFB3FFB1FFE1" masterDocTitle="Saussurea khunjerabensis (Asteraceae, Cardueae), a new species from Pamir" masterLastPageNumber="74" masterPageNumber="65" pageNumber="69" updateTime="1699376808871" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A56301D7418706739F4C9A8AE36ECB0E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F48EBC8AEBFE7FC1B75533D4C38224CB">
<mods:title id="DBAFBCC77C23093CB2C90FC334520D49">Saussurea khunjerabensis (Asteraceae, Cardueae), a new species from Pamir</mods:title>
</mods:titleInfo>
<mods:name id="30B6828EA9F4B7FDCCD26DF1B911D13C" type="personal">
<mods:role id="7E5451C7484FF63C2675E156E6799DFC">
<mods:roleTerm id="6E749EA2E69E67B6A3B5B243B07FF634">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BDA0101FD92F5B8C69DA5CBE48237873">Li, Tian</mods:namePart>
<mods:nameIdentifier id="925C247890062C124808DAB2DC27B486" type="ORCID">0000-0001-8872-7711</mods:nameIdentifier>
<mods:affiliation id="B1A664A78C37D03595741EB53BFDA3FC">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, China. &amp; litian 207 @ mails. ucas. ac. cn; https: // orcid. org / 0000 - 0001 - 8872 - 7711</mods:affiliation>
<mods:nameIdentifier id="3032DD4A741AED5C5AED415B5050D958" type="email">litian207@mails.ucas.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="EE13448FBEF7EBF9EF93684898DEA0B1" type="personal">
<mods:role id="60E4CDF630EBEC258AD35B8C63921BE2">
<mods:roleTerm id="EEA02E0BE826FB07B02412F33D14DB75">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61BBB96F0CC694D4C30A424C2F3D94EC">Xu, Lian-Sheng</mods:namePart>
<mods:nameIdentifier id="CD32DBC84CF352AE845A10DBBE265E29" type="ORCID">0000-0002-9874-9452</mods:nameIdentifier>
<mods:affiliation id="8DA25C7B2EFD9EB1B3C9B57938C144F6">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, China. &amp; xuliansheng @ ibcas. ac. cn; https: // orcid. org / 0000 - 0002 - 9874 - 9452</mods:affiliation>
<mods:nameIdentifier id="F2930839EB986BF8933FA613DE28310F" type="email">xuliansheng@ibcas.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="81975FCFF2853B5E938566838D24BE18" type="personal">
<mods:role id="B2D7DDBA743729470426C8012F892269">
<mods:roleTerm id="7D403D764B4D5BC73ABF97ABD76F469B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0660F08268F0EBD748024FC99067E942">Chen, You-Sheng</mods:namePart>
<mods:nameIdentifier id="50A29692775E4F85FB275CE53B29093B" type="ORCID">0000-0002-7729-1075</mods:nameIdentifier>
<mods:affiliation id="D089DF6EBE475E3FA69FECD6666DD875">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, China. &amp; yschen @ scbg. ac. cn; https: // orcid. org / 0000 - 0002 - 7729 - 1075</mods:affiliation>
<mods:nameIdentifier id="67FFB66E74BB43B5D32B0B47FBD52829" type="email">yschen@scbg.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="64F5F89D94C718E3D0C60473C42F96DF">text</mods:typeOfResource>
<mods:relatedItem id="C6D33FE5F50C6646DEEBA4093A6BC7B8" type="host">
<mods:titleInfo id="3C7F7EE0E4E98556783CEB7AE10991CE">
<mods:title id="02FA421CC803B2B42788482251E265C1">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0AD36D003642A5B72EAB27148C881E9B">
<mods:date id="90F83192F590AD3F31E101719BB112BE">2022</mods:date>
<mods:detail id="A5A3D3D6BAF3A4FC0C6CA1918651807F" type="pubDate">
<mods:number id="F264828300CF9190EDB26E7AA5646195">2022-09-06</mods:number>
</mods:detail>
<mods:detail id="6E8C303A68605535E037004FE3914797" type="volume">
<mods:number id="977BE19B0CC3B9D5E6101DC220212316">561</mods:number>
</mods:detail>
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<mods:number id="84D04809DBCD4352D450A5AE13434235">1</mods:number>
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<mods:start id="0E2767D5E2EB3560F654C6041AED55EA">65</mods:start>
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<mods:url id="5E51B3120CBE8B8AF21EE211AB55442D">http://dx.doi.org/10.11646/phytotaxa.561.1.6</mods:url>
</mods:location>
<mods:classification id="8CE25480F048E653094651D6140B5E8D">journal article</mods:classification>
<mods:identifier id="D7E23285034C55C4DBDB298A6DD257BF" type="CLB-Dataset">138346</mods:identifier>
<mods:identifier id="834F49AF5384EBD918AE832E7546529F" type="DOI">10.11646/phytotaxa.561.1.6</mods:identifier>
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<mods:identifier id="8D15D528559AFB609A6618A3D462A225" type="Zenodo-Dep">7052827</mods:identifier>
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<treatment id="03A787F66771FFF1FF75FF47FE11FC69" ID-DOI="http://doi.org/10.5281/zenodo.7052829" ID-GBIF-Taxon="199964706" ID-Zenodo-Dep="7052829" LSID="urn:lsid:plazi:treatment:03A787F66771FFF1FF75FF47FE11FC69" httpUri="http://treatment.plazi.org/id/03A787F66771FFF1FF75FF47FE11FC69" lastPageId="7" lastPageNumber="72" pageId="4" pageNumber="69">
<subSubSection id="C314656B6771FFF2FF75FF47FCA0FEEF" box="[136,785,244,271]" pageId="4" pageNumber="69" type="nomenclature">
<paragraph id="8BB136E06771FFF2FF75FF47FCA0FEEF" blockId="4.[136,785,244,271]" box="[136,785,244,271]" pageId="4" pageNumber="69">
<heading id="D0F9818C6771FFF2FF75FF47FCA0FEEF" box="[136,785,244,271]" fontSize="11" level="2" pageId="4" pageNumber="69" reason="3">
<taxonomicName id="4C0E4D636771FFF2FF75FF47FD89FEEF" ID-CoL="9VGDN" authority="Y. S. Chen" authorityName="Y. S. Chen" authorityYear="2022" box="[136,568,244,271]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis" status="sp. nov.">
<emphasis id="B97AEAF26771FFF2FF75FF47FE09FEEE" bold="true" box="[136,440,244,271]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
Y. S. Chen
</taxonomicName>
,
<emphasis id="B97AEAF26771FFF2FDB9FF47FD2BFEEF" bold="true" box="[580,666,244,270]" italics="true" pageId="4" pageNumber="69">
<taxonomicNameLabel id="A24957896771FFF2FDB9FF47FD2BFEEF" box="[580,666,244,270]" pageId="4" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13352A656771FFF2FD57FF47FD5FFEEF" box="[682,750,244,270]" captionStart="FIGURE 2" captionStartId="3.[136,229,2004,2026]" captionTargetBox="[237,1350,403,1979]" captionTargetId="figure-92@3.[237,1350,403,1979]" captionTargetPageId="3" captionText="FIGURE 2. Living plants of Saussurea khunjerabensis. A–C. Habitat of S. khunjerabensis. All photos by Xu-Guang Yan from Khunjerab pass, Taxkorgan county, China." figureDoi="http://doi.org/10.5281/zenodo.7052833" httpUri="https://zenodo.org/record/7052833/files/figure.png" pageId="4" pageNumber="69">Fig. 2</figureCitation>
,
<figureCitation id="13352A656771FFF2FD07FF47FCB9FEEF" box="[762,776,244,270]" captionStart="FIGURE 3" captionStartId="5.[136,229,1985,2007]" captionTargetBox="[200,1384,257,1891]" captionTargetId="figure-19@5.[151,1436,190,1961]" captionTargetPageId="5" captionText="FIGURE 3. Saussurea khunjerabensis. A. Habit; B. Style branches; C. Corolla; D.Achene with pappus; E. Inner pappus; F. Outer pappus; G. Anther; H. Phyllaries (from left to right: outer to inner). Drawn by Miss Shu-Han Li, based on the holotype." figureDoi="http://doi.org/10.5281/zenodo.7052835" httpUri="https://zenodo.org/record/7052835/files/figure.png" pageId="4" pageNumber="69">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FEF3FE7EFE98" pageId="4" pageNumber="69" type="materials_examined">
<paragraph id="8BB136E06771FFF2FF75FEF3FE7EFE98" blockId="4.[136,1452,320,378]" pageId="4" pageNumber="69">
<emphasis id="B97AEAF26771FFF2FF75FEF3FF73FEB4" bold="true" box="[136,194,320,341]" pageId="4" pageNumber="69">Type:</emphasis>
—
<materialsCitation id="3B663CBD6771FFF2FF26FEF3FE7AFE98" ID-GBIF-Occurrence="3898374301" collectingDate="2000-07-22" collectionCode="IBSC, SHI, XJBI" collectorName="Ping Yan &amp; Hui Sun" country="China" elevation="5000" location="Taxkorgan county" municipality="Khunjerab" pageId="4" pageNumber="69" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F31976706771FFF2FF26FEF3FE98FEB4" box="[219,297,320,341]" name="China" pageId="4" pageNumber="69">CHINA</collectingCountry>
.
<location id="8ED1603B6771FFF2FEC9FEF3FE52FEB4" LSID="urn:lsid:plazi:treatment:03A787F66771FFF1FF75FF47FE11FC69:8ED1603B6771FFF2FEC9FEF3FE52FEB4" box="[308,483,320,342]" country="China" municipality="Khunjerab" name="Taxkorgan county" pageId="4" pageNumber="69">Taxkorgan county</location>
,
<collectingMunicipality id="6BD5AC9A6771FFF2FE12FEF3FDE5FEB7" box="[495,596,320,342]" pageId="4" pageNumber="69">Khunjerab</collectingMunicipality>
pass, alpine scree slopes,
<quantity id="4CF69B056771FFF2FCA4FEF3FC17FEB4" box="[857,934,320,342]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="69" unit="m" value="5000.0">
<elevation id="0023D1D36771FFF2FCA4FEF3FC17FEB4" box="[857,934,320,342]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="69" unit="m" value="5000.0">5000 m</elevation>
</quantity>
,
<date id="FFB010206771FFF2FC4CFEF3FB82FEB4" box="[945,1075,320,342]" pageId="4" pageNumber="69" value="2000-07-22">
<collectingDate id="EFF4E9C86771FFF2FC4CFEF3FB82FEB4" box="[945,1075,320,342]" pageId="4" pageNumber="69" value="2000-07-22">22 July 2000</collectingDate>
</date>
,
<emphasis id="B97AEAF26771FFF2FBC2FEF3FAF0FEB4" box="[1087,1345,320,341]" italics="true" pageId="4" pageNumber="69">
<collectorName id="26FB53366771FFF2FBC2FEF3FB26FEB4" box="[1087,1175,320,341]" pageId="4" pageNumber="69">Ping Yan</collectorName>
&amp;
<collectorName id="26FB53366771FFF2FB44FEF3FAB8FEB4" box="[1209,1289,320,341]" pageId="4" pageNumber="69">Hui Sun</collectorName>
3527
</emphasis>
(
<typeStatus id="54B588426771FFF2FAB2FEF3FA19FEB7" box="[1359,1448,320,342]" pageId="4" pageNumber="69" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED1FAE256771FFF2FF40FED7FF44FE98" box="[189,245,356,377]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="4" pageNumber="69" type="Herbarium">IBSC</collectionCode>
; isotypes:
<collectionCode id="ED1FAE256771FFF2FEA6FED7FE35FE98" box="[347,388,356,377]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14032" lsid="urn:lsid:biocol.org:col:14032" name="Shihezi Agricultural College" pageId="4" pageNumber="69" type="Herbarium">SHI</collectionCode>
,
<collectionCode id="ED1FAE256771FFF2FE70FED7FE75FE98" box="[397,452,356,377]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14783" lsid="urn:lsid:biocol.org:col:14783" name="Xinjiang Institute of Ecology and Geography" pageId="4" pageNumber="69" type="Herbarium">XJBI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FE1BFC81FC77" pageId="4" pageNumber="69" type="description">
<paragraph id="8BB136E06771FFF2FF75FE1BFC81FC77" blockId="4.[136,1452,424,919]" pageId="4" pageNumber="69">
Herbs perennial,
<quantity id="4CF69B056771FFF2FEB1FE1BFE0AFE22" box="[332,443,424,451]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.5" metricValueMin="1.1" pageId="4" pageNumber="69" unit="cm" value="18.0" valueMax="25.0" valueMin="11.0">11–25 cm</quantity>
high, monocarpic. Caudex stout, unbranched. Stem solitary, ca.
<quantity id="4CF69B056771FFF2FB60FE1BFB67FE22" box="[1181,1238,424,451]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="69" unit="cm" value="2.0">2 cm</quantity>
in diameter, erect, simple, usually hidden by reflexed leaves, basally covered with fibrous remains of petioles; stem, cauline leaves and synflorescence covered with dense white or yellowish arachnoid lanate. Rosette and stem leaves petiolate; petioles
<quantity id="4CF69B056771FFF2FF75FDA7FF6AFDCE" box="[136,219,532,559]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="4" pageNumber="69" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 cm</quantity>
long; leaf blade linear or linear-oblong,
<quantity id="4CF69B056771FFF2FD5CFDA7FD45FDCE" box="[673,756,532,559]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="4" pageNumber="69" unit="cm" value="5.5" valueMax="9.0" valueMin="2.0">2–9 cm</quantity>
long,
<quantity id="4CF69B056771FFF2FCC6FDA7FC29FDCE" box="[827,920,532,559]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="4" pageNumber="69" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 mm</quantity>
wide, both surfaces light green, base attenuate; leaf blade margin 2–8 pinnately lobed or pinnatifid, lobe linear,
<quantity id="4CF69B056771FFF2FC90FD8BFC7DFDB2" box="[877,972,568,595]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="4" pageNumber="69" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 mm</quantity>
long, apex acute. Uppermost stem leaves linear, sessile,
<quantity id="4CF69B056771FFF2FED7FDEFFECAFD96" box="[298,379,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="69" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long,
<quantity id="4CF69B056771FFF2FE3DFDEFFDADFD96" box="[448,540,604,631]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="69" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
wide, usually reflexed, both surfaces densely white or yellowish arachnoid lanate, margin entire, apex acute. Capitula numerous, densely congested on the top of stem in a hemispheric synflorescence
<quantity id="4CF69B056771FFF2FF75FD17FF5BFD5E" box="[136,234,676,703]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="69" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
in diameter, exposed at flowering time, sessile. Involucre cylindric,
<quantity id="4CF69B056771FFF2FC0AFD17FBE4FD5E" box="[1015,1109,676,703]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="4" pageNumber="69" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
in diameter. Phyllaries in 3–4 rows, membranous, apex acute; outer phyllaries linear-lanceolate,
<quantity id="4CF69B056771FFF2FC96FD7BFC52FD02" box="[875,995,712,739]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="69" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
long, ca.
<quantity id="4CF69B056771FFF2FBB2FD7BFB3EFD02" box="[1103,1167,712,739]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="69" unit="mm" value="3.0">3 mm</quantity>
wide, purple and densely white lanate; middle phyllaries triangular-lanceolate,
<quantity id="4CF69B056771FFF2FD24FD5FFCE0FCE6" box="[729,849,748,775]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="69" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
long, ca.
<quantity id="4CF69B056771FFF2FC41FD5FFC4DFCE6" box="[956,1020,748,775]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="69" unit="mm" value="2.0">2 mm</quantity>
wide, purple and densely white lanate; inner phyllaries narrowly triangular-lanceolate,
<quantity id="4CF69B056771FFF2FD5EFCA3FCADFCCA" box="[675,796,784,811]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="4" pageNumber="69" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
long, ca.
<quantity id="4CF69B056771FFF2FC71FCA3FC52FCCA" box="[908,995,784,811]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="69" unit="mm" value="1.5">1.5 mm</quantity>
wide, apically purple and densely white lanate. Receptacle bristles ca.
<quantity id="4CF69B056771FFF2FE2AFC87FDAAFCAE" box="[471,539,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="69" unit="mm" value="3.0">3 mm</quantity>
. Corolla purple,
<quantity id="4CF69B056771FFF2FD28FC87FCFCFCAE" box="[725,845,820,847]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="69" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
long, tube ca.
<quantity id="4CF69B056771FFF2FC12FC87FBF4FCAE" box="[1007,1093,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="69" unit="mm" value="7.5">7.5 mm</quantity>
long, limb ca.
<quantity id="4CF69B056771FFF2FB17FC87FA9AFCAE" box="[1258,1323,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="69" unit="mm" value="8.0">8 mm</quantity>
long, lobes ca.
<quantity id="4CF69B056771FFF2FF4FFCEBFF47FC92" box="[178,246,856,883]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="69" unit="mm" value="4.0">4 mm</quantity>
long. Achenes cylindroid, brown, obconic,
<quantity id="4CF69B056771FFF2FD0EFCEBFCE2FC92" box="[755,851,856,883]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="69" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, ribbed, glabrous. Pappus straw-colored; outer bristles
<quantity id="4CF69B056771FFF2FF1DFCCFFE8EFC76" box="[224,319,892,919]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="69" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, scabrid; inner bristles
<quantity id="4CF69B056771FFF2FDBEFCCFFD0FFC76" box="[579,702,892,919]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="69" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
, plumose.
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FC77FD7BFBE3" box="[136,714,964,1027]" pageId="4" pageNumber="69" type="biology_ecology">
<paragraph id="8BB136E06771FFF2FF75FC77FEB5FC3F" blockId="4.[136,714,964,1027]" box="[136,260,964,990]" pageId="4" pageNumber="69">
<heading id="D0F9818C6771FFF2FF75FC77FEB5FC3F" bold="true" box="[136,260,964,990]" fontSize="11" level="1" pageId="4" pageNumber="69" reason="1">
<emphasis id="B97AEAF26771FFF2FF75FC77FEB5FC3F" bold="true" box="[136,260,964,990]" pageId="4" pageNumber="69">Phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E06771FFF2FF75FC5BFD7BFBE3" blockId="4.[136,714,964,1027]" box="[136,714,1000,1027]" pageId="4" pageNumber="69">Flowering and fruiting from late July to September.</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FB83FB37FB8F" box="[136,1158,1072,1135]" pageId="4" pageNumber="69" type="etymology">
<paragraph id="8BB136E06771FFF2FF75FB83FEB8FBAB" blockId="4.[136,1158,1072,1135]" box="[136,265,1072,1098]" pageId="4" pageNumber="69">
<heading id="D0F9818C6771FFF2FF75FB83FEB8FBAB" bold="true" box="[136,265,1072,1098]" fontSize="11" level="1" pageId="4" pageNumber="69" reason="1">
<emphasis id="B97AEAF26771FFF2FF75FB83FEB8FBAB" bold="true" box="[136,265,1072,1098]" pageId="4" pageNumber="69">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E06771FFF2FF75FBE7FB37FB8F" blockId="4.[136,1158,1072,1135]" box="[136,1158,1108,1135]" pageId="4" pageNumber="69">
<taxonomicName id="4C0E4D636771FFF2FF75FBE7FE1AFB8F" authorityName="Y. S. Chen" authorityYear="2022" box="[136,427,1108,1134]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26771FFF2FF75FBE7FE1AFB8F" box="[136,427,1108,1134]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
</taxonomicName>
is named after its
<typeStatus id="54B588426771FFF2FD84FBE6FD18FB8E" box="[633,681,1109,1135]" pageId="4" pageNumber="69">type</typeStatus>
locality, Khunjerab pass,
<collectingRegion id="49CAF8026771FFF2FC36FBE7FB81FB8F" box="[971,1072,1108,1134]" country="China" name="Xinjiang Uygur" pageId="4" pageNumber="69">Xinjiang</collectingRegion>
,
<collectingCountry id="F31976706771FFF2FBC6FBE7FB33FB8F" box="[1083,1154,1108,1134]" name="China" pageId="4" pageNumber="69">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FB2FFE9AFAC2" pageId="4" pageNumber="69" type="distribution">
<paragraph id="8BB136E06771FFF2FF75FB2FFE1AFB57" blockId="4.[136,1451,1180,1315]" box="[136,427,1180,1206]" pageId="4" pageNumber="69">
<heading id="D0F9818C6771FFF2FF75FB2FFE1AFB57" bold="true" box="[136,427,1180,1206]" fontSize="11" level="1" pageId="4" pageNumber="69" reason="1">
<emphasis id="B97AEAF26771FFF2FF75FB2FFE1AFB57" bold="true" box="[136,427,1180,1206]" pageId="4" pageNumber="69">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E06771FFF2FF75FB73FE9AFAC2" blockId="4.[136,1451,1180,1315]" pageId="4" pageNumber="69">
<taxonomicName id="4C0E4D636771FFF2FF75FB73FE1FFB3B" authorityName="Y. S. Chen" authorityYear="2022" box="[136,430,1216,1242]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26771FFF2FF75FB73FE1FFB3B" box="[136,430,1216,1242]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
</taxonomicName>
is currently only known from Khunjerab pass, Taxkorgan county, southwestern
<collectingRegion id="49CAF8026771FFF2FAB4FB73FF5CFB1F" country="China" name="Xinjiang Uygur" pageId="4" pageNumber="69">Xinjiang province</collectingRegion>
,
<collectingCountry id="F31976706771FFF2FF05FB57FE8FFB1F" box="[248,318,1252,1278]" name="China" pageId="4" pageNumber="69">China</collectingCountry>
, and adjacent Khunjerab pass area in
<collectingCountry id="F31976706771FFF2FD1AFB57FCF7FB1F" box="[743,838,1252,1278]" name="Pakistan" pageId="4" pageNumber="69">Pakistan</collectingCountry>
(
<figureCitation id="13352A656771FFF2FCA8FB57FC2BFB1E" box="[853,922,1252,1279]" captionStart="FIGURE 5" captionStartId="7.[136,229,2053,2075]" captionTargetBox="[170,1351,948,2024]" captionTargetId="figure-293@7.[151,1436,926,2030]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Saussurea khunjerabensis (●)." figureDoi="http://doi.org/10.5281/zenodo.7052839" httpUri="https://zenodo.org/record/7052839/files/figure.png" pageId="4" pageNumber="69">Fig. 5</figureCitation>
). It grows on alpine scree slopes at altitudes of 4500~
<quantity id="4CF69B056771FFF2FF32FABBFE96FAC2" box="[207,295,1288,1315]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="69" unit="m" value="5000.0">5000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF2FF75FAE3FEA1F9A3" pageId="4" pageNumber="69" type="conservation">
<paragraph id="8BB136E06771FFF2FF75FAE3FEC4FA8B" blockId="4.[136,1452,1360,1602]" box="[136,373,1360,1386]" pageId="4" pageNumber="69">
<heading id="D0F9818C6771FFF2FF75FAE3FEC4FA8B" bold="true" box="[136,373,1360,1386]" fontSize="11" level="1" pageId="4" pageNumber="69" reason="1">
<emphasis id="B97AEAF26771FFF2FF75FAE3FEC4FA8B" bold="true" box="[136,373,1360,1386]" pageId="4" pageNumber="69">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E06771FFF2FF75FAC7FEA1F9A3" blockId="4.[136,1452,1360,1602]" pageId="4" pageNumber="69">
<taxonomicName id="4C0E4D636771FFF2FF75FAC7FE1BFA6F" authorityName="Y. S. Chen" authorityYear="2022" box="[136,426,1396,1422]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26771FFF2FF75FAC7FE1BFA6F" box="[136,426,1396,1422]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
</taxonomicName>
is only found in the Khunjerab pass border area between
<collectingCountry id="F31976706771FFF2FBDAFAC7FBDBFA6F" box="[1063,1130,1396,1422]" name="China" pageId="4" pageNumber="69">China</collectingCountry>
and
<collectingCountry id="F31976706771FFF2FB63FAC7FAB0FA6F" box="[1182,1281,1396,1422]" name="Pakistan" pageId="4" pageNumber="69">Pakistan</collectingCountry>
. This species is severely overcollected by local people as “snow lotus”. Many species of
<emphasis id="B97AEAF26771FFF2FC4CFA2BFC77FA53" box="[945,966,1432,1458]" italics="true" pageId="4" pageNumber="69">S.</emphasis>
subg.
<emphasis id="B97AEAF26771FFF2FBF2FA2AFB3CFA53" box="[1039,1165,1433,1458]" italics="true" pageId="4" pageNumber="69">Eriocoryne</emphasis>
are included in the IUCN red list due to their small geographic ranges, few and severely fragmented populations, and in continuing decline due to excessive medicinal collection. According to the IUCN Red List Categories and Criteria (
<bibRefCitation id="EF9F4B116771FFF2FB47FA53FE6DF9FF" author="IUCN Standards &amp; Petitions Subcommittee" pageId="4" pageNumber="69" refId="ref4108" refString="IUCN Standards and Petitions Subcommittee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. IUCN Standards and Petitions Subcommittee." type="book" year="2019">IUCN Standards and Petitions Subcommittee 2019</bibRefCitation>
), the conservation status of this species should be assessed as Critically Endangered (CR, B2ab).
</paragraph>
</subSubSection>
<subSubSection id="C314656B6771FFF1FF75F9C3FF4FFD51" lastPageId="7" lastPageNumber="72" pageId="4" pageNumber="69" type="discussion">
<paragraph id="8BB136E06771FFF2FF75F9C3FDCBF96A" blockId="4.[136,1452,1648,2071]" box="[136,634,1648,1675]" pageId="4" pageNumber="69">
<emphasis id="B97AEAF26771FFF2FF75F9C3FDC3F96B" bold="true" box="[136,626,1648,1674]" pageId="4" pageNumber="69">Similar species and phylogenetic position</emphasis>
:
</paragraph>
<paragraph id="8BB136E06771FFF1FF75F927FF17FE8C" blockId="4.[136,1452,1648,2071]" lastBlockId="7.[136,1452,158,688]" lastPageId="7" lastPageNumber="72" pageId="4" pageNumber="69">
<taxonomicName id="4C0E4D636771FFF2FF75F927FE1EF94F" authorityName="Y. S. Chen" authorityYear="2022" box="[136,431,1684,1710]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26771FFF2FF75F927FE1EF94F" box="[136,431,1684,1710]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
</taxonomicName>
belongs to
<emphasis id="B97AEAF26771FFF2FDC1F927FDE0F94F" box="[572,593,1684,1710]" italics="true" pageId="4" pageNumber="69">S.</emphasis>
subgen.
<emphasis id="B97AEAF26771FFF2FD40F926FC8AF94F" box="[701,827,1685,1710]" italics="true" pageId="4" pageNumber="69">Eriocoryne</emphasis>
because of its capitula being usually aggregated in a densely congested hemispheric synflorescence and hollow stems. In addition, the capitula is usually surrounded or half-surrounded by densely lanate subtending leaves.
<taxonomicName id="4C0E4D636771FFF2FD0AF96FFBAFF917" authorityName="Y. S. Chen" authorityYear="2022" box="[759,1054,1756,1782]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26771FFF2FD0AF96FFBAFF917" box="[759,1054,1756,1782]" italics="true" pageId="4" pageNumber="69">Saussurea khunjerabensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C0E4D636771FFF2FB43F96FFADAF917" box="[1214,1387,1756,1782]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="69" phylum="Tracheophyta" rank="species" species="simpsoniana">
<emphasis id="B97AEAF26771FFF2FB43F96FFADAF917" box="[1214,1387,1756,1782]" italics="true" pageId="4" pageNumber="69">S. simpsoniana</emphasis>
</taxonomicName>
(
<figureCitation id="13352A656771FFF2FA83F96FFF27F8FA" captionStart="FIGURE 5" captionStartId="7.[136,229,2053,2075]" captionTargetBox="[170,1351,948,2024]" captionTargetId="figure-293@7.[151,1436,926,2030]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Saussurea khunjerabensis (●)." figureDoi="http://doi.org/10.5281/zenodo.7052839" httpUri="https://zenodo.org/record/7052839/files/figure.png" pageId="4" pageNumber="69">Fig. 5</figureCitation>
) in its habit, distinct elongate erect flowering stems, uppermost linear stem leaves reflexed, numerous aggregated capitula exposed on stem apex, straw-colored pappus and purple corolla, but different by its stems usually unbranched and
<quantity id="4CF69B056771FFF2FF44F8FBFE99F882" box="[185,296,1864,1891]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.5" metricValueMin="1.1" pageId="4" pageNumber="69" unit="cm" value="18.0" valueMax="25.0" valueMin="11.0">11–25 cm</quantity>
tall (vs. usually branched and
<quantity id="4CF69B056771FFF2FD7AF8FBFD5BF882" box="[647,746,1864,1891]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="4" pageNumber="69" unit="cm" value="7.0" valueMax="12.0" valueMin="2.0">2–12 cm</quantity>
tall), rosette and stem leaves on both surfaces light green and densely white arachnoid lanate (vs. adaxially dark green or purplish, sparsely arachnoid; abaxially grayish white, arachnoid tomentose to lanate), rosette and stem leaf margins usually pinnately lobed or divided (vs. usually dentate to pinnately lobed, revolute), capitula in a hemispheric synflorescence
<quantity id="4CF69B056771FFF2FC74F807FC5AF82E" box="[905,1003,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="69" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
in diam. (vs.
<quantity id="4CF69B056771FFF2FB75F807FB40F82E" box="[1160,1265,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="4" pageNumber="69" unit="cm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 cm</quantity>
in diam.), white or yellowish lanate indumentum (vs. white or purple lanate indumentum), phyllaries linear-lanceolate to narrowly triangular-lanceolate (vs. narrowly triangular-ovate or narrowly ovate-elliptic to linear), ribbed and glabrous achene ca.
<quantity id="4CF69B056772FFF1FF4DFF2DFEBEFF58" box="[176,271,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="72" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long (vs. wrinkled achene ca.
<quantity id="4CF69B056772FFF1FD93FF2DFD7DFF58" box="[622,716,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="72" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long). According to
<emphasis id="B97AEAF26772FFF1FC47FF2DFBD4FF59" box="[954,1125,158,184]" italics="true" pageId="7" pageNumber="72">
Flora of
<collectingCountry id="F31976706772FFF1FBDCFF2DFBD4FF59" box="[1057,1125,158,184]" name="China" pageId="7" pageNumber="72">China</collectingCountry>
</emphasis>
(
<bibRefCitation id="EF9F4B116772FFF1FB8BFF2DFA2EFF59" author="Shi, C. &amp; Raab-Straube, E. von" box="[1142,1439,158,184]" pageId="7" pageNumber="72" pagination="56 - 149" refId="ref4548" refString="Shi, C. &amp; Raab-Straube, E. von (2011) Saussurea Candolle. In: Wu, Z. Y. &amp; Raven, P. H. (eds.) Flora of China, vols. 20 - 21. Science Press, Beijing &amp; Missouri Botanical Garden Press, pp. 56 - 149." type="book chapter" year="2011">Shi &amp; Raab-Straube 2011</bibRefCitation>
),
<taxonomicName id="4C0E4D636772FFF1FF75FF70FE82FF3D" box="[136,307,195,221]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="simpsoniana">
<emphasis id="B97AEAF26772FFF1FF75FF70FE82FF3D" box="[136,307,195,221]" italics="true" pageId="7" pageNumber="72">S. simpsoniana</emphasis>
</taxonomicName>
is mainly distributed in
<collectingRegion id="49CAF8026772FFF1FDB7FF71FD16FF3D" box="[586,679,194,220]" country="China" name="Qinghai" pageId="7" pageNumber="72">Qinghai</collectingRegion>
, southwestern
<collectingRegion id="49CAF8026772FFF1FCB2FF71FC05FF3D" box="[847,948,194,220]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="72">Xinjiang</collectingRegion>
, southern and southwestern
<collectingRegion id="49CAF8026772FFF1FB08FF71FA82FF3D" box="[1269,1331,194,220]" country="China" name="Xizang" pageId="7" pageNumber="72">Tibet</collectingRegion>
. However, we found that the specimens of
<taxonomicName id="4C0E4D636772FFF1FDF9FF54FD02FEE1" box="[516,691,231,257]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="simpsoniana">
<emphasis id="B97AEAF26772FFF1FDF9FF54FD02FEE1" box="[516,691,231,257]" italics="true" pageId="7" pageNumber="72">S. simpsoniana</emphasis>
</taxonomicName>
collected in
<collectingRegion id="49CAF8026772FFF1FCB3FF55FC01FEE1" box="[846,944,230,256]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="72">Xinjiang</collectingRegion>
are misidentified and actually belong to
<taxonomicName id="4C0E4D636772FFF1FA6BFF54FE83FEC5" authorityName="Y. S. Chen" authorityYear="2022" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26772FFF1FA6BFF54FE83FEC5" italics="true" pageId="7" pageNumber="72">S. khunjerabensis</emphasis>
</taxonomicName>
. Chen (2015) considered that
<taxonomicName id="4C0E4D636772FFF1FD7EFEB8FC9DFEC5" box="[643,812,267,293]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="simpsoniana">
<emphasis id="B97AEAF26772FFF1FD7EFEB8FC9DFEC5" box="[643,812,267,293]" italics="true" pageId="7" pageNumber="72">S. simpsoniana</emphasis>
</taxonomicName>
only occurs in
<collectingRegion id="49CAF8026772FFF1FC2AFEB9FBA0FEC5" box="[983,1041,266,292]" country="China" name="Xizang" pageId="7" pageNumber="72">Tibet</collectingRegion>
of
<collectingCountry id="F31976706772FFF1FBC8FEB9FBCAFEC5" box="[1077,1147,266,292]" name="China" pageId="7" pageNumber="72">China</collectingCountry>
,
<collectingCountry id="F31976706772FFF1FB7BFEB9FB7DFEC5" box="[1158,1228,266,292]" name="Nepal" pageId="7" pageNumber="72">Nepal</collectingCountry>
,
<collectingCountry id="F31976706772FFF1FB2BFEB9FAA1FEC5" box="[1238,1296,266,292]" name="India" pageId="7" pageNumber="72">India</collectingCountry>
and Kashmir. Therefore, the geographical distributions of these two species are different in
<collectingCountry id="F31976706772FFF1FC14FE9DFB9DFEA9" box="[1001,1068,302,328]" name="China" pageId="7" pageNumber="72">China</collectingCountry>
and may overlap in
<collectingCountry id="F31976706772FFF1FAEDFE9DFADEFEA9" box="[1296,1391,302,328]" name="Pakistan" pageId="7" pageNumber="72">Pakistan</collectingCountry>
(
<figureCitation id="13352A656772FFF1FA83FE9DFF29FE8C" captionStart="FIGURE 5" captionStartId="7.[136,229,2053,2075]" captionTargetBox="[170,1351,948,2024]" captionTargetId="figure-293@7.[151,1436,926,2030]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Saussurea khunjerabensis (●)." figureDoi="http://doi.org/10.5281/zenodo.7052839" httpUri="https://zenodo.org/record/7052839/files/figure.png" pageId="7" pageNumber="72">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF7166686770FFF3FF75F872FB00F81A" ID-DOI="http://doi.org/10.5281/zenodo.7052835" ID-Zenodo-Dep="7052835" httpUri="https://zenodo.org/record/7052835/files/figure.png" pageId="5" pageNumber="70" startId="5.[136,229,1985,2007]" targetBox="[200,1384,257,1891]" targetPageId="5">
<paragraph id="8BB136E06770FFF3FF75F872FB00F81A" blockId="5.[136,1452,1985,2043]" pageId="5" pageNumber="70">
<emphasis id="B97AEAF26770FFF3FF75F872FF4CF837" bold="true" box="[136,253,1985,2007]" pageId="5" pageNumber="70">FIGURE 3.</emphasis>
<taxonomicName id="4C0E4D636770FFF3FEFFF871FE4BF837" authorityName="Y. S. Chen" authorityYear="2022" box="[258,506,1985,2007]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="70" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26770FFF3FEFFF871FE4BF837" box="[258,506,1985,2007]" italics="true" pageId="5" pageNumber="70">Saussurea khunjerabensis</emphasis>
</taxonomicName>
. A. Habit; B. Style branches; C. Corolla; D. Achene with pappus; E. Inner pappus; F. Outer pappus; G. Anther; H. Phyllaries (from left to right: outer to inner). Drawn by Miss Shu-Han Li, based on the holotype.
</paragraph>
</caption>
<caption id="DF7166686773FFF0FF75F85FFC8BF7C6" ID-DOI="http://doi.org/10.5281/zenodo.7052837" ID-Zenodo-Dep="7052837" httpUri="https://zenodo.org/record/7052837/files/figure.png" pageId="6" pageNumber="71" startId="6.[136,229,2028,2050]" targetBox="[157,1430,190,2004]" targetPageId="6">
<paragraph id="8BB136E06773FFF0FF75F85FFC8BF7C6" blockId="6.[136,1452,2028,2087]" pageId="6" pageNumber="71">
<emphasis id="B97AEAF26773FFF0FF75F85FFF4CF7E0" bold="true" box="[136,253,2028,2050]" pageId="6" pageNumber="71">FIGURE 4.</emphasis>
Living plants of
<emphasis id="B97AEAF26773FFF0FE62F85EFD36F7E3" box="[415,647,2029,2050]" italics="true" pageId="6" pageNumber="71">
<taxonomicName id="4C0E4D636773FFF0FE62F85EFD35F7E3" box="[415,644,2029,2050]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="71" phylum="Tracheophyta" rank="species" species="simpsoniana">Saussurea simpsoniana</taxonomicName>
.
</emphasis>
A. Lanate indumentum; B–C. Capitulum; D. Habitat; from Humla district, Nepal,
<emphasis id="B97AEAF26773FFF0FA64F85EFF51F7C7" italics="true" pageId="6" pageNumber="71">S. R. Zhang</emphasis>
,
<emphasis id="B97AEAF26773FFF0FF11F7A2FE7AF7C4" box="[236,459,2064,2086]" italics="true" pageId="6" pageNumber="71">Y. S. Chen et al. H0866</emphasis>
(PE). All photos by You-Sheng Chen.
</paragraph>
</caption>
<paragraph id="8BB136E06772FFF1FF40FEC4FCA3FD89" blockId="7.[136,1452,158,688]" pageId="7" pageNumber="72">
<taxonomicName id="4C0E4D636772FFF1FF40FEC4FE53FE71" authorityName="Y. S. Chen" authorityYear="2022" box="[189,482,374,401]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26772FFF1FF40FEC4FE53FE71" box="[189,482,374,401]" italics="true" pageId="7" pageNumber="72">Saussurea khunjerabensis</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C0E4D636772FFF1FD50FEC4FCEDFE71" box="[685,860,374,401]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="gossipiphora">
<emphasis id="B97AEAF26772FFF1FD50FEC4FCEDFE71" box="[685,860,374,401]" italics="true" pageId="7" pageNumber="72">S. gossipiphora</emphasis>
</taxonomicName>
(
<figureCitation id="13352A656772FFF1FC91FEC5FC00FE71" box="[876,945,374,400]" captionStart="FIGURE 4" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[157,1430,190,2004]" captionTargetId="figure-25@6.[157,1430,190,2004]" captionTargetPageId="6" captionText="FIGURE 4. Living plants of Saussurea simpsoniana. A. Lanate indumentum; B–C. Capitulum; D. Habitat; from Humla district, Nepal, S. R. Zhang, Y. S. Chen et al. H0866 (PE). All photos by You-Sheng Chen." figureDoi="http://doi.org/10.5281/zenodo.7052837" httpUri="https://zenodo.org/record/7052837/files/figure.png" pageId="7" pageNumber="72">Fig. 4</figureCitation>
) in its habit, size, distinct elongate erect and unbranched flowering stems, rosette and lower stem leaves petiolate, white or yellowish lanate indumentum and purple corolla, but distinguished by its rosette and stem leaves on both surfaces light green and densely white or yellowish arachnoid lanate (vs. both surfaces green and sparsely pilose or glabrous), lower leaf blade margins pinnately lobed or divided (vs. sparsely denticulate), capitula exposed on stem apex (vs. capitula surrounded and exceeded by densely lanate bracts), phyllaries linear-lanceolate to narrowly triangular-lanceolate (vs. ovate or narrowly ovate-elliptic to linear), achenes brown and ribbed (vs. black and smooth).
</paragraph>
<paragraph id="8BB136E06772FFF1FF40FDC1FF4FFD51" blockId="7.[136,1452,158,688]" pageId="7" pageNumber="72">
Morphological differences among
<taxonomicName id="4C0E4D636772FFF1FDC3FDC0FCD1FD6D" authorityName="Y. S. Chen" authorityYear="2022" box="[574,864,626,653]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="khunjerabensis">
<emphasis id="B97AEAF26772FFF1FDC3FDC0FCD1FD6D" box="[574,864,626,653]" italics="true" pageId="7" pageNumber="72">Saussurea khunjerabensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E4D636772FFF1FC91FDC0FBA4FD6D" box="[876,1045,627,653]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="simpsoniana">
<emphasis id="B97AEAF26772FFF1FC91FDC0FBA4FD6D" box="[876,1045,627,653]" italics="true" pageId="7" pageNumber="72">S. simpsoniana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D636772FFF1FBB4FDC0FB47FD6D" box="[1097,1270,626,653]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="72" phylum="Tracheophyta" rank="species" species="gossipiphora">
<emphasis id="B97AEAF26772FFF1FBB4FDC0FB47FD6D" box="[1097,1270,626,653]" italics="true" pageId="7" pageNumber="72">S. gossipiphora</emphasis>
</taxonomicName>
are summarised in
<tableCitation id="C68C035B6772FFF1FF59FD25FF4BFD51" box="[164,250,662,688]" captionStart="Table 2" captionStartId="8.[136,199,160,184]" captionTargetBox="[144,1443,197,674]" captionTargetPageId="8" captionText="Table 2. Morphological comparisons between Saussurea khunjerabensis sp. nov. and related species." httpUri="http://table.plazi.org/id/DF716668677DFFFEFF75FF13FB0CFF59" pageId="7" pageNumber="72" tableUuid="DF716668677DFFFEFF75FF13FB0CFF59">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C314656B6772FFF1FF75FD6DFE11FC69" pageId="7" pageNumber="72" type="materials_examined">
<paragraph id="8BB136E06772FFF1FF75FD6DFD23FD19" blockId="7.[136,1452,734,905]" box="[136,658,734,760]" pageId="7" pageNumber="72">
<heading id="D0F9818C6772FFF1FF75FD6DFD23FD19" bold="true" box="[136,658,734,760]" fontSize="11" level="1" pageId="7" pageNumber="72" reason="1">
<emphasis id="B97AEAF26772FFF1FF75FD6DFD23FD19" bold="true" box="[136,658,734,760]" pageId="7" pageNumber="72">
Additional specimens examined (
<typeStatus id="54B588426772FFF1FDECFD6DFD3AFD19" box="[529,651,734,760]" pageId="7" pageNumber="72" type="paratype">paratypes</typeStatus>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E06772FFF1FF75FCB1FE11FC69" blockId="7.[136,1452,734,905]" pageId="7" pageNumber="72">
<collectingCountry id="F31976706772FFF1FF75FCB1FF55FCFD" box="[136,228,770,796]" name="China" pageId="7" pageNumber="72">CHINA</collectingCountry>
. Taxkorgan county, Khunjerab pass,
<quantity id="4CF69B056772FFF1FD7DFCB1FD66FCFC" box="[640,727,770,797]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="72" unit="m" value="4800.0">4800 m</quantity>
,
<date id="FFB010206772FFF1FD1EFCB1FC03FCFD" box="[739,946,770,796]" pageId="7" pageNumber="72" value="2013-09-04">4 September 2013</date>
,
<emphasis id="B97AEAF26772FFF1FC40FCB0FB22FCFD" box="[957,1171,770,797]" italics="true" pageId="7" pageNumber="72">Y. S. Chen 13-2280</emphasis>
(PE); Taxkorgan county, Khunjerab pass,
<quantity id="4CF69B056772FFF1FEB5FC95FE10FCA0" box="[328,417,806,833]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="7" pageNumber="72" unit="m" value="4700.0">4700 m</quantity>
,
<date id="FFB010206772FFF1FE50FC95FDF3FCA1" box="[429,578,806,832]" pageId="7" pageNumber="72" value="1974-08">August 1974</date>
,
<emphasis id="B97AEAF26772FFF1FDB2FC94FD6FFCA1" box="[591,734,806,832]" italics="true" pageId="7" pageNumber="72">K. J. Feng 1</emphasis>
(HNWP); Taxkorgan county, Shuibulanggou,
<quantity id="4CF69B056772FFF1FB0CFC95FAFBFCA0" box="[1265,1354,806,833]" metricMagnitude="3" metricUnit="m" metricValue="4.52" pageId="7" pageNumber="72" unit="m" value="4520.0">4520 m</quantity>
,
<date id="FFB010206772FFF1FAA5FC95FF72FC85" pageId="7" pageNumber="72" value="1978-07-15">15 July 1978</date>
,
<emphasis id="B97AEAF26772FFF1FF30FCF8FE71FC85" box="[205,448,842,868]" italics="true" pageId="7" pageNumber="72">
<collectingRegion id="49CAF8026772FFF1FF30FCF8FE9CFC85" box="[205,301,843,868]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="72">Xinjiang</collectingRegion>
Exped. 1465
</emphasis>
(PE).
<collectingCountry id="F31976706772FFF1FDFAFCF9FD21FC85" box="[519,656,842,868]" name="Pakistan" pageId="7" pageNumber="72">PAKISTAN</collectingCountry>
. Khunjerab National Park, Khunjerab pass,
<quantity id="4CF69B056772FFF1FB85FCF9FB7EFC84" box="[1144,1231,842,869]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="72" unit="m" value="4800.0">4800 m</quantity>
,
<date id="FFB010206772FFF1FB27FCF9FA16FC85" box="[1242,1447,842,868]" pageId="7" pageNumber="72" value="2013-09-04">4 September 2013</date>
,
<emphasis id="B97AEAF26772FFF1FF75FCDCFEEFFC69" box="[136,350,878,905]" italics="true" pageId="7" pageNumber="72">Y. S. Chen 13-2279</emphasis>
(PE).
</paragraph>
</subSubSection>
</treatment>
</document>
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