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<document id="72C912409C17E68C3421CC390C39BDF6" ID-DOI="10.11646/zootaxa.3619.5.3" ID-GBIF-Dataset="01f367b1-2419-457b-912c-e01b0a1aef19" ID-ISSN="1175-5326" ID-Zenodo-Dep="221066" ID-ZooBank="36A7C0C3-3961-47F3-A0D0-E338BF0E15D6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578735267" checkinUser="plazi" docAuthor="Morgulis, Elizabeth" docDate="2013" docId="03A787B23C67FFB3EBEDFF2DFD511693" docLanguage="en" docName="zt03619p553.pdf" docOrigin="Zootaxa 3619 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otites grata Loew" docType="treatment" docVersion="9" lastPageNumber="545" masterDocId="FF9EFFCA3C64FFB6EB7AFFBAFF9B1647" masterDocTitle="Review of Otites Latreille (Diptera: Ulidiidae) from Israel with two new species and notes on biology and behavior" masterLastPageNumber="553" masterPageNumber="541" pageNumber="544" updateTime="1698323758385" updateUser="plazi">
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<mods:title id="89ED0085DBAB49588CD7AC38E6DD3FA1">Review of Otites Latreille (Diptera: Ulidiidae) from Israel with two new species and notes on biology and behavior</mods:title>
</mods:titleInfo>
<mods:name id="C68762B31DDB50E1881A3BD5B6EB298B" type="personal">
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<mods:namePart id="C0B87AE7BADFBC2BF01E4081F864CEF3">Morgulis, Elizabeth</mods:namePart>
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<mods:date id="585BA52C2405BB4A091D0FBBDA80E12F">2013</mods:date>
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<treatment id="03A787B23C67FFB3EBEDFF2DFD511693" ID-DOI="http://doi.org/10.5281/zenodo.6151583" ID-GBIF-Taxon="119560927" ID-Zenodo-Dep="6151583" LSID="urn:lsid:plazi:treatment:03A787B23C67FFB3EBEDFF2DFD511693" httpUri="http://treatment.plazi.org/id/03A787B23C67FFB3EBEDFF2DFD511693" lastPageId="5" lastPageNumber="545" pageId="3" pageNumber="544">
<subSubSection id="C314652F3C67FFB5EBEDFF2DFE59175F" pageId="3" pageNumber="544" type="nomenclature">
<paragraph id="8BB136A43C67FFB5EBEDFF2DFE1916F6" blockId="3.[151,462,151,210]" box="[151,386,151,177]" pageId="3" pageNumber="544">
<heading id="D0F981C83C67FFB5EBEDFF2DFE1916F6" bold="true" box="[151,386,151,177]" fontSize="11" level="1" pageId="3" pageNumber="544" reason="1">
<taxonomicName id="4C0E4D273C67FFB5EBEDFF2DFE1916F6" ID-CoL="4B739" authority="Loew" authorityName="Loew" box="[151,386,151,177]" class="Insecta" family="Ulidiidae" genus="Otites" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="grata">
<emphasis id="B97AEAB63C67FFB5EBEDFF2DFE1916F6" bold="true" box="[151,386,151,177]" pageId="3" pageNumber="544">
<emphasis id="B97AEAB63C67FFB5EBEDFF2DFEB816F6" bold="true" box="[151,291,151,177]" italics="true" pageId="3" pageNumber="544">Otites grata</emphasis>
(Loew)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB136A43C67FFB5EBEDFF00FE551695" blockId="3.[151,462,151,210]" box="[151,462,186,210]" pageId="3" pageNumber="544">
(
<figureCitation id="13352A213C67FFB5EBE5FF00FF7C1695" box="[159,231,186,210]" captionStart="FIGURES 2 4" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[236,1253,237,1844]" captionTargetId="figure@8.[179,1409,193,1932]" captionTargetPageId="8" captionText="FIGURES 2 4. Head, lateral view. 2. Otites grata; dashed line shows gena height. 3. O. nox n. sp. 4. O. vitalyi n. sp. FIGURES 5 7. Epandrium, posterior view, cerci removed. 5. O. grata. 6. O. nox n. sp. 7. O. vitalyi n. sp." httpUri="https://zenodo.org/record/221068/files/figure.png" pageId="3" pageNumber="544">Figs 2</figureCitation>
, 5, 8, 11, 14, 17, 20)
</paragraph>
<paragraph id="8BB136A43C67FFB5EBEDFEB8FE59175F" blockId="3.[151,627,258,311]" box="[151,450,258,280]" pageId="3" pageNumber="544">
<taxonomicName id="4C0E4D273C67FFB5EBEDFEB8FE25175F" authority="Loew 1856: 50" authorityName="Loew" authorityPageNumber="50" authorityYear="1856" box="[151,446,258,280]" class="Insecta" family="Platystomatidae" genus="Ortalis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="grata">
<emphasis id="B97AEAB63C67FFB5EBEDFEB8FE87175F" box="[151,284,258,280]" italics="true" pageId="3" pageNumber="544">Ortalis grata</emphasis>
Loew 1856: 50
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C314652F3C67FFB5EBEDFE98FDE81770" box="[151,627,289,311]" pageId="3" pageNumber="544" type="description">
<paragraph id="8BB136A43C67FFB5EBEDFE98FDE81770" blockId="3.[151,627,258,311]" box="[151,627,289,311]" pageId="3" pageNumber="544">
<taxonomicName id="4C0E4D273C67FFB5EBEDFE98FE4A1770" authority="Hennig 1939: 31" authorityName="Hennig" authorityPageNumber="31" authorityYear="1939" box="[151,465,289,311]" class="Insecta" family="Ulidiidae" genus="Otites" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="grata">
<emphasis id="B97AEAB63C67FFB5EBEDFE98FE8A1770" box="[151,273,290,311]" italics="true" pageId="3" pageNumber="544">Otites grata</emphasis>
: Hennig 1939: 31
</taxonomicName>
; Soós 1984: 48.
</paragraph>
</subSubSection>
<subSubSection id="C314652F3C67FFB5EBEDFED9FA821166" pageId="3" pageNumber="544" type="materials_examined">
<paragraph id="8BB136A43C67FFB5EBEDFED9FB43129B" blockId="3.[151,1437,355,2006]" pageId="3" pageNumber="544">
<emphasis id="B97AEAB63C67FFB5EBEDFED9FD3F173B" bold="true" box="[151,676,355,380]" pageId="3" pageNumber="544">
Material examined.
<collectingCountry id="F31976343C67FFB5EAF6FED9FE62173B" box="[396,505,355,380]" name="Israel" pageId="3" pageNumber="544">ISRAEL</collectingCountry>
: Har Hermon
</emphasis>
: Har Hermon [Hermon],
<date id="FFB010643C67FFB5E8B7FEDEFBA6173B" box="[973,1085,356,380]" pageId="3" pageNumber="544" value="1998-05-12">12.v.1998</date>
, J. Atlani (1Ƥ), [Mt. Hermon]
<date id="FFB010643C67FFB5EBEDFE3DFE9417D8" box="[151,271,391,415]" pageId="3" pageNumber="544" value="1973-04-23">23.iv.1973</date>
, D. Furth (23, 1Ƥ), [Mt. Hermon]
<quantity id="4CF69B413C67FFB5E9D4FE3DFD6117D8" box="[686,762,391,415]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="544" unit="m" value="800.0">800 m</quantity>
,
<date id="FFB010643C67FFB5E872FE3DFC1A17D8" box="[776,897,391,415]" pageId="3" pageNumber="544" value="1978-04-27">27.iv.1978</date>
, D. Furth (2Ƥ),
<date id="FFB010643C67FFB5EF3BFE3DFB2317D8" box="[1089,1208,391,415]" pageId="3" pageNumber="544" value="1973-04-23">23.iv.1973</date>
(13, 3Ƥ),
<date id="FFB010643C67FFB5EE46FE3DFA0717D8" box="[1340,1436,391,415]" pageId="3" pageNumber="544" value="1979-05-03">3.v.1979</date>
(1Ƥ); Majdal Shams,
<date id="FFB010643C67FFB5EAD8FE16FD951783" box="[418,526,428,452]" pageId="3" pageNumber="544" value="1992-06-09">9.vi.1992</date>
, A. Freidberg (13),
<date id="FFB010643C67FFB5E868FE16FC191783" box="[786,898,428,452]" pageId="3" pageNumber="544" value="1998-05-12">12.v.1998</date>
, A. Freidberg (13); Majdal Shams [Magdel Shames],
<date id="FFB010643C67FFB5EA76FE75FEE717A0" box="[268,380,463,487]" pageId="3" pageNumber="544" value="1998-05-12">12.v.1998</date>
, N. Yaacov (1Ƥ); Mezudat Nimrod [Golan Qala'at Nimrod],
<date id="FFB010643C67FFB5EF1FFE75FB4E17A0" box="[1125,1237,463,487]" pageId="3" pageNumber="544" value="1983-05-19">19.v.1983</date>
, I. Yarom (13); Mezudat Nimrod [Qala'at Nimrod],
<date id="FFB010643C67FFB5E957FE4EFD14144B" box="[557,655,500,524]" pageId="3" pageNumber="544" value="1987-05-06">6.v.1987</date>
, A. Shlagman (1Ƥ);
<emphasis id="B97AEAB63C67FFB5E801FE49FBAB144B" bold="true" box="[891,1072,499,524]" pageId="3" pageNumber="544">Golan Heights:</emphasis>
Panyas [Banias],
<date id="FFB010643C67FFB5EF86FE4EFAEE144B" box="[1276,1397,500,524]" pageId="3" pageNumber="544" value="1977-04-19">19.iv.1977</date>
, D. Simon (13),
<date id="FFB010643C67FFB5EA55FDADFE3C1468" box="[303,423,535,559]" pageId="3" pageNumber="544" value="1983-04-12">12.iv.1983</date>
, A. Freidberg (1Ƥ);
<emphasis id="B97AEAB63C67FFB5E9EFFDA2FCD31477" bold="true" box="[661,840,535,560]" pageId="3" pageNumber="544">Upper Galilee:</emphasis>
Metula,
<date id="FFB010643C67FFB5E8CAFDADFB881468" box="[944,1043,535,559]" pageId="3" pageNumber="544" value="1987-05-06">6.v.1987</date>
, A. Freidberg (1Ƥ); Kefar Gil'adi,
<date id="FFB010643C67FFB5EBEDFD86FE971413" box="[151,268,572,596]" pageId="3" pageNumber="544" value="1956-04-11">11.iv.1956</date>
(13); Ma'ayan Barukh [Mayan Baruch],
<date id="FFB010643C67FFB5E999FD86FCC41413" box="[739,863,572,596]" pageId="3" pageNumber="544" value="1973-03-17">17.iii.1973</date>
, D. Furth (13),
<date id="FFB010643C67FFB5EF62FD86FB0A1413" box="[1048,1169,572,596]" pageId="3" pageNumber="544" value="1974-04-26">26.iv.1974</date>
, D. Furth (1),
<date id="FFB010643C67FFB5EE4FFD86FA0C1413" box="[1333,1431,572,596]" pageId="3" pageNumber="544" value="2006-05-01">1.v.2006</date>
, L. Friedman (1Ƥ), [HaTanur],
<date id="FFB010643C67FFB5E97AFDE5FDE31430" box="[512,632,607,631]" pageId="3" pageNumber="544" value="1983-04-13">13.iv.1983</date>
, A. Freidberg (223, 13Ƥ), I. Yarom (53, 7Ƥ); Nahal 'Iyyon Nature Reserve [Tanur],
<date id="FFB010643C67FFB5EA22FD3EFE2014DB" box="[344,443,644,668]" pageId="3" pageNumber="544" value="1987-05-06">6.v.1987</date>
, F. Kaplan (1Ƥ); Nahal 'Iyyon, haTanur,
<quantity id="4CF69B413C67FFB5E8FBFD3EFC5214DB" box="[897,969,644,668]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="544" unit="m" value="430.0">430 m</quantity>
,
<geoCoordinate id="EE3A50633C67FFB5E8AEFD3EFBC414DB" box="[980,1119,644,668]" direction="north" orientation="latitude" pageId="3" pageNumber="544" precision="1" value="33.267735">33º16.064'N</geoCoordinate>
<geoCoordinate id="EE3A50633C67FFB5EF1CFD3EFB6B14DB" box="[1126,1264,644,668]" direction="east" orientation="longitude" pageId="3" pageNumber="544" precision="1" value="35.575">35º34.500'E</geoCoordinate>
,
<date id="FFB010643C67FFB5EF81FD3EFAED14DB" box="[1275,1398,644,668]" pageId="3" pageNumber="544" value="2011-03-15">15.iii.2011</date>
, A. Freidberg (13), L. Bodner (13); E. Morgulis (63, 4Ƥ); Qiryat Shemona,
<date id="FFB010643C67FFB5E8B0FD1DFBB614F8" box="[970,1069,679,703]" pageId="3" pageNumber="544" value="1990-05-03">3.v.1990</date>
, J. Halperin (13); Emeq haHula,
<date id="FFB010643C67FFB5EBEDFD76FE8814A3" box="[151,275,716,740]" pageId="3" pageNumber="544" value="1981-03-17">17.iii.1981</date>
, A. Freidberg (13); Avivim,
<date id="FFB010643C67FFB5E921FD76FD5714A3" box="[603,716,716,740]" pageId="3" pageNumber="544" value="1981-05-18">18.v.1981</date>
, A. Freidberg (133, 13Ƥ),
<date id="FFB010643C67FFB5E885FD76FBEE14A3" box="[1023,1141,716,740]" pageId="3" pageNumber="544" value="1981-04-18">18.iv.1981</date>
(1Ƥ); Goren,
<date id="FFB010643C67FFB5EE6FFD76FAE414A3" box="[1301,1407,716,740]" pageId="3" pageNumber="544" value="1987-04-08">8.iv.1987</date>
, I. Nussbaum (13); Nahal Dishon [N. Dishon],
<date id="FFB010643C67FFB5E9F7FD55FC9E1540" box="[653,773,751,775]" pageId="3" pageNumber="544" value="1983-04-14">14.iv.1983</date>
, Y. Zvik (2Ƥ); Nahal Dalton [Dalton],
<date id="FFB010643C67FFB5EFC3FD55FAAA1540" box="[1209,1329,751,775]" pageId="3" pageNumber="544" value="1974-04-25">25.iv.1974</date>
, D. Furth (23, 2Ƥ); Upper Nahal Keziv [Up. N. Cziv],
<date id="FFB010643C67FFB5E9C6FCAEFC84156B" box="[700,799,788,812]" pageId="3" pageNumber="544" value="1983-05-03">3.v.1983</date>
, Y. Zvik (1Ƥ); Upper Nahal Keziv [upper N. Keziv],
<date id="FFB010643C67FFB5EBEDFC8DFF621508" box="[151,249,823,847]" pageId="3" pageNumber="544" value="1983-05-03">3.v.1983</date>
, A. Freidberg (13); Ma'alot-Tarshiha,
<date id="FFB010643C67FFB5E9C7FC8DFCAD1508" box="[701,822,823,847]" pageId="3" pageNumber="544" value="2005-04-26">26.iv.2005</date>
, M. Meir (1); Ga'aton [Gaaton],
<date id="FFB010643C67FFB5EFC2FC8DFAAA1508" box="[1208,1329,823,847]" pageId="3" pageNumber="544" value="1961-04-25">25.iv.1961</date>
, B. Galil (1Ƥ); Nahal Keziv [N. Keziv],
<date id="FFB010643C67FFB5E975FCE6FDE21533" box="[527,633,860,884]" pageId="3" pageNumber="544" value="1978-04-04">4.iv.1978</date>
, D. Furth (13); Har Meron,
<date id="FFB010643C67FFB5E8A2FCE6FBD21533" box="[984,1097,860,884]" pageId="3" pageNumber="544" value="1967-05-17">17.v.1967</date>
, J. Kugler (1Ƥ); Har Meron [Meiron],
<date id="FFB010643C67FFB5EA6AFCC5FE1215D0" box="[272,393,895,919]" pageId="3" pageNumber="544" value="1972-04-20">20.iv.1972</date>
, J. Kugler (33); Har Meron [Meron],
<date id="FFB010643C67FFB5E81BFCC5FC4615D0" box="[865,989,895,919]" pageId="3" pageNumber="544" value="1958-03-31">31.iii.1958</date>
, J. Kugler (13); Har Meron reserve, camping under Kefar Meron,
<quantity id="4CF69B413C67FFB5EA8AFC1EFDA015FB" box="[496,571,932,956]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="544" unit="m" value="600.0">600 m</quantity>
,
<geoCoordinate id="EE3A50633C67FFB5E930FC1EFD3D15FB" box="[586,678,932,956]" direction="north" orientation="latitude" pageId="3" pageNumber="544" precision="925" value="32.966667">32°58'N</geoCoordinate>
<geoCoordinate id="EE3A50633C67FFB5E9CAFC1EFC9715FB" box="[688,780,932,956]" direction="east" orientation="longitude" pageId="3" pageNumber="544" precision="925" value="35.433334">35°26'E</geoCoordinate>
,
<date id="FFB010643C67FFB5E860FC1EFC0815FB" box="[794,915,932,956]" pageId="3" pageNumber="544" value="2002-04-25">25.iv.2002</date>
, A. Freidberg (113, 15Ƥ), L. Friedman (13, 3Ƥ); Har Gilon [Har Gilon],
<date id="FFB010643C67FFB5EA8FFC7DFDEB1598" box="[501,624,967,991]" pageId="3" pageNumber="544" value="1969-03-15">15.iii.1969</date>
, J. Kugler (13);
<emphasis id="B97AEAB63C67FFB5E839FC72FA9B15A7" bold="true" box="[835,1280,967,992]" pageId="3" pageNumber="544">
<collectingCountry id="F31976343C67FFB5E839FC72FC0315A7" box="[835,920,968,992]" name="Jordan" pageId="3" pageNumber="544">Jordan</collectingCountry>
Valley and Southern Golan:
</emphasis>
Kare Deshe,
<date id="FFB010643C67FFB5EBEDFC56FE891243" box="[151,274,1004,1028]" pageId="3" pageNumber="544" value="1974-03-22">22.iii.1974</date>
, D. Furth (13, 1Ƥ); Huqoq Huquq,
<date id="FFB010643C67FFB5E9C1FC56FCAC1243" box="[699,823,1004,1028]" pageId="3" pageNumber="544" value="1981-03-17">17.iii.1981</date>
, A. Freidberg (43, 3Ƥ), T. Furman (1Ƥ); 'En Ravid,
<date id="FFB010643C67FFB5EBEDFBB5FE9F1260" box="[151,260,1039,1063]" pageId="3" pageNumber="544" value="1969-03-08">8.iii.1969</date>
, J. Kugler (13),
<date id="FFB010643C67FFB5EABBFBB5FDAC1260" box="[449,567,1039,1063]" pageId="3" pageNumber="544" value="1969-04-19">19.iv.1969</date>
(23),
<date id="FFB010643C67FFB5E9F8FBB5FD711260" box="[642,746,1039,1063]" pageId="3" pageNumber="544" value="1978-04-03">3.iv.1978</date>
D. Furth (13),
<date id="FFB010643C67FFB5E8E0FBB5FC601260" box="[922,1019,1039,1063]" pageId="3" pageNumber="544" value="1983-05-03">3.v.1983</date>
I. Yarom (13); Ginnosar,
<date id="FFB010643C67FFB5EE59FBB5FA001260" box="[1315,1435,1039,1063]" pageId="3" pageNumber="544" value="1958-03-15">15.iii.1958</date>
(2Ƥ);
<emphasis id="B97AEAB63C67FFB5EBA6FB89FE17120B" bold="true" box="[220,396,1075,1100]" pageId="3" pageNumber="544">Carmel Ridge:</emphasis>
Haifa,
<date id="FFB010643C67FFB5EA9AFB8EFDD0120B" box="[480,587,1076,1100]" pageId="3" pageNumber="544" value="1972-04-01">1.iv.1972</date>
, J. Kugler (23); Karmel [Carmel],
<date id="FFB010643C67FFB5E8A3FB8EFBCA120B" box="[985,1105,1076,1100]" pageId="3" pageNumber="544" value="1992-04-28">28.iv.1992</date>
, A. Shahar (1Ƥ); Nahal Oren [n. Oranim],
<date id="FFB010643C67FFB5EA55FBEDFE331228" box="[303,424,1111,1135]" pageId="3" pageNumber="544" value="1959-04-21">21.iv.1959</date>
, J. Krystal (13); Nahal Oren [N. Oren],
<date id="FFB010643C67FFB5E8F2FBEDFC6E1228" box="[904,1013,1111,1135]" pageId="3" pageNumber="544" value="1979-03-03">3.iii.1979</date>
, D. Furth (13),
<date id="FFB010643C67FFB5EFC2FBEDFABB1228" box="[1208,1312,1111,1135]" pageId="3" pageNumber="544" value="1978-04-03">3.iv.1978</date>
(23, 4Ƥ); Nahal Yoqne'am [N. Yokneam],
<date id="FFB010643C67FFB5E973FBC6FD1E12D3" box="[521,645,1148,1172]" pageId="3" pageNumber="544" value="1974-03-22">22.iii.1974</date>
, D. Furth (13),
<date id="FFB010643C67FFB5E83AFBC6FC3C12D3" box="[832,935,1148,1172]" pageId="3" pageNumber="544" value="1968-04-06">6.iv.1968</date>
J. Kugler (13, 1Ƥ);
<emphasis id="B97AEAB63C67FFB5EFE2FBC6FAD712D3" bold="true" box="[1176,1356,1147,1172]" pageId="3" pageNumber="544">Lower Galilee:</emphasis>
Qiryat Tiv'on [K. Tivon],
<date id="FFB010643C67FFB5EA11FB25FE7D12F0" box="[363,486,1183,1207]" pageId="3" pageNumber="544" value="1955-03-25">25.iii.1955</date>
, L. Fishelsohn (13); Qiryat Tiv'on [Tivon],
<date id="FFB010643C67FFB5E8ADFB25FBC812F0" box="[983,1107,1183,1207]" pageId="3" pageNumber="544" value="1956-03-22">22.iii.1956</date>
, J. Kugler (1Ƥ),
<date id="FFB010643C67FFB5EE6BFB25FAE012F0" box="[1297,1403,1183,1207]" pageId="3" pageNumber="544" value="1975-04-02">2.iv.1975</date>
, F. Kaplan (2Ƥ), M. Kaplan (13, 1Ƥ);
<emphasis id="B97AEAB63C67FFB5E952FB79FD45129B" bold="true" box="[552,734,1219,1244]" pageId="3" pageNumber="544">Yizre'el Valley:</emphasis>
Nahalal,
<date id="FFB010643C67FFB5E833FB7EFC5A129B" box="[841,961,1220,1244]" pageId="3" pageNumber="544" value="1941-04-17">17.iv.1941</date>
, Ch. Bytinski-Salz (1Ƥ).
</paragraph>
<paragraph id="8BB136A43C67FFB5EBBDFB5DFA821166" blockId="3.[151,1437,355,2006]" pageId="3" pageNumber="544">
<emphasis id="B97AEAB63C67FFB5EBBDFB5DFE211347" bold="true" box="[199,442,1255,1280]" pageId="3" pageNumber="544">Redescription. Head</emphasis>
(
<figureCitation id="13352A213C67FFB5EAB3FB5DFD9512B8" box="[457,526,1255,1279]" captionStart="FIGURES 2 4" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[236,1253,237,1844]" captionTargetId="figure@8.[179,1409,193,1932]" captionTargetPageId="8" captionText="FIGURES 2 4. Head, lateral view. 2. Otites grata; dashed line shows gena height. 3. O. nox n. sp. 4. O. vitalyi n. sp. FIGURES 5 7. Epandrium, posterior view, cerci removed. 5. O. grata. 6. O. nox n. sp. 7. O. vitalyi n. sp." httpUri="https://zenodo.org/record/221068/files/figure.png" pageId="3" pageNumber="544">Fig. 2</figureCitation>
): Structure: 1.101.21 times as high as long. Frons posteriorly 1.021.15 times as wide as long, and anteriorly, at lunule level, 1.301.55 times as wide as long. Face straight. Carina flattened, about 0.550.60 times as wide as antennal groove at same level; carina protrusion about 0.8 times as long as parafacial at narrowest level. Eye 1.321.44 times as high as long. Gena 0.510.65 times as high as eye. Fronto-orbital plate at antenna insertion level 0.620.72 times as long as gena height. Pedicel 0.680.82 times as long as high at apex and 0.500.65 times as long as 1st flagellomere; first flagellomere 1.261.35 times as long as high. Color and vestiture: Occiput black dorsally, brown-orange ventrally, gray microtrichose; vertex orange medially, black laterally, gray microtrichose. Ocellar triangle black, gray microtrichose. Face mostly brown and indistinctly silvery-white microtrichose, except antennal groove brown to black. Gena yellow anteriorly, brownish posteriorly, slightly white microtrichose. Postgena brownish-beige, white microtrichose. Scape and pedicel orange-brown; 1st flagellomere orange, dorsoapically infuscate; arista orange to brown. Clypeus black, gray microtrichose. Palp yellow to orange, apically often infuscate; proboscis dark brown. Chaetotaxy: Medial vertical seta 1.041.16 times as long as lateral vertical seta; ocellar and postocellar setae each 0.400.57 times as long as medial vertical seta; posterior orbital seta 1.31.4 times as long as anterior orbital seta and 0.660.76 times as long as medial vertical seta. Frons densely setulose, setulae 0.30.4 times as long as posterior orbital seta. Gena with long black setulae and 24 setae. Vibrissal angle, postgena and occiput long setulose. Labium with long mixed black and yellow setulae.
</paragraph>
</subSubSection>
<subSubSection id="C314652F3C67FFB2EBB3F894FD7012CD" lastPageId="4" lastPageNumber="545" pageId="3" pageNumber="544" type="description">
<paragraph id="8BB136A43C67FFB5EBB3F894FA061192" blockId="3.[151,1437,355,2006]" pageId="3" pageNumber="544">
<emphasis id="B97AEAB63C67FFB5EBB3F894FEB91101" bold="true" box="[201,290,1838,1862]" pageId="3" pageNumber="544">Thorax</emphasis>
: Color and vestiture: Black. Scutum gray microtrichose except 1 medial and 2 lateral longitudinal stripes (anterior and posterior to transverse suture) with brownish microtrichia: medial stripe extends from anterior margin of scutum to level of anterior dorsocentral seta, anterior lateral stripe triangular, posterior lateral stripe extends from transverse suture to level slightly posterior to anterior dorsocentral seta. Anepisternum, anepimeron and katepisternum subshining, slightly gray microtrichose, remaining pleura silvery-gray microtrichose.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBEDFF2DFA821693" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">Chaetotaxy: Two supra-alar, 1 postalar, 1 intra-alar, 2 dorsocentral, 1 acrostichal, 1 proepisternal, 24 anepisternal, and 1 katepisternal setae present. Scutum, postpronotal lobe, katepisternum and anepisternum setulose.</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FF5AFCB11723" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FF5AFE9A16BF" bold="true" box="[201,257,224,248]" pageId="4" pageNumber="545">Legs</emphasis>
: Coxae and trochanters brown; femora mostly brown to black, except apical 0.16 yellow; foretibia mostly brown to black, except basal 0.16 yellow, midtibia and hindtibia mostly yellow, except apical 0.25 brown to black; tarsi mostly yellow, except basal fore tarsomere often brown to black. Coxae and trochanters densely gray microtrichose, remaining parts slightly gray microtrichose.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FED5FEDA1429" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FED5FE9217CF" bold="true" box="[201,265,367,392]" pageId="4" pageNumber="545">Wing</emphasis>
(
<figureCitation id="13352A213C60FFB2EA60FED5FEF517C0" box="[282,366,367,391]" captionStart="FIGURES 17 19" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[262,1299,215,1895]" captionTargetId="figure@10.[247,1330,193,1905]" captionTargetPageId="10" captionText="FIGURES 17 19. Wings. 17. Otites grata. 18. O. nox n. sp. 19. O. vitalyi n. sp. Abbreviations: BB—basal band, SBB—subbasal band, PB—pterostigmal band, SAB—subapical band, AB—apical band." httpUri="https://zenodo.org/record/221070/files/figure.png" pageId="4" pageNumber="545">Fig. 17</figureCitation>
): Pattern: Brown to black spotted pattern present: SBB extending from costal vein to vein Cu1, narrowed and faded posteriorly; PB as two spots, one at apex of pterostigma and adjacent part of cell r1, and second at crossvein R-M, spots often vaguely connected; SAB present at vein DM-Cu and at apex of cell r1, reaching AB anteriorly. Venation: Veins yellow to orange except sections within dark bands. Cell r4+5 0.580.70 times as wide at apex as at widest point. Crossvein R-M aligned proximal to apex of vein R1. Vein Cu2 curved, not forming posteroapical lobe. Calypteres yellowish white. Halter base brown, stem and knob yellow to orange. Wing length
<quantity id="4CF69B413C60FFB2EBEDFDECFEA61429" box="[151,317,598,622]" metricMagnitude="-3" metricUnit="m" metricValue="6.08" metricValueMax="8.24" metricValueMin="3.92" pageId="4" pageNumber="545" unit="mm" value="6.08" valueMax="8.24" valueMin="3.92">3.928.24 mm</quantity>
.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FDC3FEEA149D" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FDC3FEA514D5" bold="true" box="[201,318,633,658]" pageId="4" pageNumber="545">Abdomen</emphasis>
: Black. Syntergite 1+2 with transverse, medially widened, stripe of silvery-gray microtrichia; each of tergites 15 with medially widened basal stripe of dense silvery-gray microtrichia; remaining parts indistinctly gray microtrichose.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FD5CFCA315F5" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FD5CFE1014B9" bold="true" box="[201,395,741,766]" pageId="4" pageNumber="545">Male terminalia</emphasis>
: Epandrium (Fig. 5) as high as wide, slightly rhombic in posterior view; medial surstylus fully fused with epandrium, bearing numerous prensisetae; lateral surstylus slim and elongate. Phallus (
<figureCitation id="13352A213C60FFB2EE37FCB0FA081565" box="[1357,1427,778,802]" captionStart="FIGURES 8 10" captionStartId="9.[151,264,1953,1976]" captionTargetBox="[255,1331,294,1888]" captionTargetId="figure@9.[179,1409,193,1932]" captionTargetPageId="9" captionText="FIGURES 8 10. Phallus. 8. Otites grata. 9. O. nox n. sp. 10. O. vitalyi n. sp." httpUri="https://zenodo.org/record/221069/files/figure.png" pageId="4" pageNumber="545">Fig. 8</figureCitation>
) laterally (on both sides) short setulose at basal 0.9 and long setulose at apical 0.1; basal setulae about half as long as phallus width, apical setulae about twice as long as phallus width. Phallus dorsomedially with scales at basal 0.9 and setulose at apical 0.1; medial setulae at apical 0.1 of phallus as long as lateral setulae at same place. Glans membranous and setulose, setulae as long as width of glans.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FC04FBA21259" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FC04FE3F1591" bold="true" box="[201,420,957,982]" pageId="4" pageNumber="545">Female terminalia</emphasis>
: Aculeus 5.55.8 times as long as wide. Cercal unit oval (
<figureCitation id="13352A213C60FFB2EF4DFC04FB111591" box="[1079,1162,958,982]" captionStart="FIGURES 11 13" captionStartId="9.[151,264,1983,2006]" captionText="FIGURES 11 13. Cercal unit. 11. O. grata. 12. O. nox n. sp. 13. O. vitalyi n. sp. FIGURES 14 16. Spermatheca. 14. O. grata. 15. O. nox n. sp. 16. O. vitalyi n. sp." pageId="4" pageNumber="545">Fig. 11</figureCitation>
), with well pronounced, long lateral groove. Sensilla: BV apical to BD. Pair of large campaniform sensilla present distal to BV. 3 spermathecae, wrinkled, about 12.5 times as long as wide at widest place (Fig. 14).
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FB90FD981205" blockId="4.[151,1437,151,2026]" box="[201,515,1066,1090]" pageId="4" pageNumber="545">
Body length
<quantity id="4CF69B413C60FFB2EA20FB90FE641205" box="[346,511,1066,1090]" metricMagnitude="-3" metricUnit="m" metricValue="6.84" metricValueMax="9.6" metricValueMin="4.08" pageId="4" pageNumber="545" unit="mm" value="6.84" valueMax="9.6" valueMin="4.08">4.089.60 mm</quantity>
.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FBF4FBC11221" blockId="4.[151,1437,151,2026]" box="[201,1114,1102,1126]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FBF4FF631221" bold="true" box="[201,248,1102,1126]" pageId="4" pageNumber="545">Egg</emphasis>
: Elongate, white, rice grain like, anterior end pointed, posterior end rounded.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FBC9FD7012CD" blockId="4.[151,1437,151,2026]" box="[201,747,1138,1162]" pageId="4" pageNumber="545">Measurments (mm): Length 0.91.1, width 0.25.</paragraph>
</subSubSection>
<subSubSection id="C314652F3C60FFB2EBB3FB2CFD4712E9" box="[201,732,1173,1198]" pageId="4" pageNumber="545" type="materials_examined">
<paragraph id="8BB136A43C60FFB2EBB3FB2CFD4712E9" blockId="4.[151,1437,151,2026]" box="[201,732,1173,1198]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FB2CFEFE12E9" bold="true" box="[201,357,1173,1198]" pageId="4" pageNumber="545">
<typeStatus id="54B588063C60FFB2EBB3FB2CFE9912E9" box="[201,258,1174,1198]" pageId="4" pageNumber="545">Type</typeStatus>
locality.
</emphasis>
Fiume (currently Rieka,
<collectingCountry id="F31976343C60FFB2E9FBFB2CFD4E12E9" box="[641,725,1174,1198]" name="Croatia" pageId="4" pageNumber="545">Croatia</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="C314652F3C60FFB2EBB3FB03FC30135D" pageId="4" pageNumber="545" type="distribution">
<paragraph id="8BB136A43C60FFB2EBB3FB03FAB21295" blockId="4.[151,1437,151,2026]" box="[201,1321,1209,1234]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FB03FEF91295" bold="true" box="[201,354,1209,1234]" pageId="4" pageNumber="545">Distribution.</emphasis>
<collectingCountry id="F31976343C60FFB2EA13FB00FE241295" box="[361,447,1210,1234]" name="Croatia" pageId="4" pageNumber="545">Croatia</collectingCountry>
;
<collectingCountry id="F31976343C60FFB2EAB0FB00FD811295" box="[458,538,1210,1234]" name="Turkey" pageId="4" pageNumber="545">Turkey</collectingCountry>
(Amasia),
<collectingCountry id="F31976343C60FFB2E9EDFB00FD491295" box="[663,722,1210,1234]" name="Syria" pageId="4" pageNumber="545">Syria</collectingCountry>
(Latakia),
<collectingCountry id="F31976343C60FFB2E834FB00FC2B1295" box="[846,944,1210,1234]" name="Lebanon" pageId="4" pageNumber="545">Lebanon</collectingCountry>
(Beyrout) (Hennig, 1939),
<collectingCountry id="F31976343C60FFB2EF9EFB00FABD1295" box="[1252,1318,1210,1234]" name="Israel" pageId="4" pageNumber="545">Israel</collectingCountry>
.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FB67FAB312B1" blockId="4.[151,1437,151,2026]" box="[201,1320,1245,1270]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FB67FE5712B1" bold="true" box="[201,460,1245,1270]" pageId="4" pageNumber="545">
Distribution in
<collectingCountry id="F31976343C60FFB2EAFAFB64FE5D12B1" box="[384,454,1246,1270]" name="Israel" pageId="4" pageNumber="545">Israel</collectingCountry>
.
</emphasis>
All parts north to and including Samaria (North to +32° 32' 31.86&quot; latitude).
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FABBFC30135D" blockId="4.[151,1437,151,2026]" box="[201,939,1281,1306]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FABBFE6B135D" bold="true" box="[201,496,1281,1306]" pageId="4" pageNumber="545">
Adult activity (in
<collectingCountry id="F31976343C60FFB2EAE6FABBFE7F135D" box="[412,484,1281,1306]" name="Israel" pageId="4" pageNumber="545">Israel</collectingCountry>
).
</emphasis>
Middle of March to beginning of June.
</paragraph>
</subSubSection>
<subSubSection id="C314652F3C60FFB3EBB3FA9FFD511693" lastPageId="5" lastPageNumber="546" pageId="4" pageNumber="545" type="biology_ecology">
<paragraph id="8BB136A43C60FFB2EBB3FA9FFEFE1389" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
<emphasis id="B97AEAB63C60FFB2EBB3FA9FFE431379" bold="true" box="[201,472,1317,1342]" pageId="4" pageNumber="545">Biology and behavior.</emphasis>
No apparent courtship was observed. Copulation position: The male mounts the female, grasps her aculeus with his surstyli and inserts his phallus. During insertion, the male usually places his foretarsi on the female's scutum, sometimes on her head, his midtarsi on her scutellum or abdominal syntergite 1+2, and his hindtarsi on the substrate. Both male and female usually extend their proboscises slowly several times during copulation.
</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3FA60FC791019" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">When copulation ends, the male remains in the same position on the female, pulling his abdomen away and slightly upward from her, often while holding her aculeus with one of his hindtarsi. Sometimes the phallus is fully withdrawn from the female's ovipositor at this point, whereas in other cases the male dismounts the female, faces the other direction, and withdraws the rest of his phallus completely. (n=4).</paragraph>
<paragraph id="8BB136A43C60FFB2EBB3F9D0FC6B111D" blockId="4.[151,1437,151,2026]" pageId="4" pageNumber="545">
Mating trophallaxis (
<figureCitation id="13352A213C60FFB2EACFF9D0FD8E10C5" box="[437,533,1642,1666]" captionStart="FIGURE 20. A" captionStartId="11.[151,250,1539,1562]" captionTargetBox="[192,1417,242,1450]" captionTargetId="figure@11.[151,1436,193,1518]" captionTargetPageId="11" captionText="FIGURE 20. A schematic presentation of female Otites grata expelling and consuming substance after mating. a d. Female expelling substance from aculeus. e f. Female depositing substance on substrate with the help of hindlegs. g. Female backing up in search of the substance (arrow shows the direction of movement). h j. Female consuming the substance." httpUri="https://zenodo.org/record/221071/files/figure.png" pageId="4" pageNumber="545">Fig. 20a</figureCitation>
j): After mating, the female starts grooming herself with special attention to the abdomen. She rubs it with her hindlegs, stroking it from base to apex. This grooming process is interrupted by occasional contractions of the abdomen and extensions of the ovipositor and/or the proboscis. After 1520 min, a droplet of opaque white substance appears at the tip of the aculeus. The droplet grows larger and, after about 10 seconds to 2.5 minutes, is deposited on the substrate, often with the help of the hindlegs. The female then reverses 14 steps to a position in which her proboscis comes in touch with the droplet, places her labellum on it and consumes it. Consumption of the droplet lasts approximately 46 min (n=3).
</paragraph>
<paragraph id="8BB136A43C60FFB3EBB3F8DCFD511693" blockId="4.[151,1437,151,2026]" lastBlockId="5.[151,1436,151,212]" lastPageId="5" lastPageNumber="546" pageId="4" pageNumber="545">
Oviposition and immature stages: Females laid eggs in narrow crevices such as the net that covered the ventilation openings of the cages, cracks of tree bark, cuts in cardboard and in the leaf sheaths of dry
<taxonomicName id="4C0E4D273C60FFB2EE61F830FF621181" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="4" pageNumber="545" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B97AEAB63C60FFB2EE61F830FF621181" italics="true" pageId="4" pageNumber="545">Phragmites australis</emphasis>
</taxonomicName>
. The number of eggs per batch was 1058 (number of batches =7). Eggs hatched 814 days after oviposition. First-instar larvae were able to skip by curving their body and grabbing their anal area with the mandibles (apparently flexing the muscles and) releasing the grip and pushing away from the substrate. Rearing attempts of the larvae to adults were unsuccessful.
</paragraph>
</subSubSection>
</treatment>
</document>