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<document id="DC913A42B6E1F5E200F2B2D4FDC93844" ID-CLB-Dataset="8672" ID-DOI="10.5281/zenodo.4468383" ID-GBIF-Dataset="5c5c8aad-73f1-4a26-bdca-d1d02ba3fec1" ID-Zenodo-Dep="4468383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611044739228" checkinUser="plazi" docAuthor="Martínez, Leonel, Brescovit, Antonio D., Villarreal, Eduardo &amp; Oliveira, Luiz Fernando M." docDate="2021" docId="A86D27653D77FF91FF1DF90EFCB0FE1F" docLanguage="en" docName="zootaxa.4914.1.1.pdf" docOrigin="Zootaxa 4914 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Patrera Simon 1903" docType="treatment" docVersion="13" lastPageNumber="5" masterDocId="54545F1D3D73FF94FF8AFFC9FFEBFF96" masterDocTitle="An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia" masterLastPageNumber="64" masterPageNumber="1" pageNumber="5" updateTime="1698924792971" updateUser="ExternalLinkService">
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<mods:title id="5FD6AA3AA527B55AC7C60474EBC0C59C">An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia</mods:title>
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<mods:namePart id="2C2C774649ABBF79FC71BEF277055605">Martínez, Leonel</mods:namePart>
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<mods:namePart id="172256568E15A237F1F27715FE0DEEA6">Brescovit, Antonio D.</mods:namePart>
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<mods:nameIdentifier id="EC1667BB6E9A1BAAA136EF9BDF590DEE" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="6F8C6B73B2CBB7094B6A008281E394DE">Villarreal, Eduardo</mods:namePart>
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<mods:nameIdentifier id="D012526982A060D055FDB4A373D93CA4" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E96642C4B5760DBFF5717E9DAFEEE9DD">Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:title id="6CBE251DB868C6799CAF222C2F9CA524">Zootaxa</mods:title>
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<mods:date id="F9700D1FF1484A833CC61DD811E44BE1">2021</mods:date>
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<treatment id="A86D27653D77FF91FF1DF90EFCB0FE1F" ID-DOI="http://doi.org/10.5281/zenodo.4469556" ID-GBIF-Taxon="177121388" ID-Zenodo-Dep="4469556" LSID="urn:lsid:plazi:treatment:A86D27653D77FF91FF1DF90EFCB0FE1F" httpUri="http://treatment.plazi.org/id/A86D27653D77FF91FF1DF90EFCB0FE1F" lastPageId="5" lastPageNumber="5" pageId="4" pageNumber="5">
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<taxonomicName id="E7C4EDF03D77FF90FF1DF90EFE78F976" ID-CoL="6JL2" ID-ENA="2938911" authority="Simon, 1903" authorityName="Simon" authorityYear="1903" box="[151,403,1734,1761]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D77FF90FF1DF90EFE78F976" bold="true" box="[151,403,1734,1761]" pageId="4" pageNumber="5">
<emphasis id="12B04A613D77FF90FF1DF90EFF19F977" bold="true" box="[151,242,1735,1761]" italics="true" pageId="4" pageNumber="5">Patrera</emphasis>
<bibRefCitation id="4455EB823D77FF90FF70F90FFE78F976" author="Simon, E." box="[250,403,1734,1760]" pageId="4" pageNumber="5" pagination="21" refId="ref41646" refString="Simon, E. (1903) Descriptions d'arachnides nouveaux. Annales de la Societe Entomologique de Belgique, 47, 21 ‾ 39. https: // doi. org / 10.5962 / bhl. part. 25299" type="journal article" year="1903">Simon, 1903</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitationGroup id="00D4B15D3D77FF90FF1DF8C5FC12F8D4" pageId="4" pageNumber="5">
<treatmentCitation id="A165B0623D77FF90FF1DF8C5FE7FF8B5" author="Simon, E." box="[151,404,1803,1827]" page="30" pageId="4" pageNumber="5" year="1903">
<taxonomicName id="E7C4EDF03D77FF90FF1DF8C5FE7FF8B5" ID-CoL="6JL2" authority="Simon, 1903: 30" authorityName="Simon" authorityPageNumber="30" authorityYear="1903" box="[151,404,1803,1827]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D77FF90FF1DF8C5FF0EF8B5" box="[151,229,1804,1827]" italics="true" pageId="4" pageNumber="5">Patrera</emphasis>
<bibRefCitation id="4455EB823D77FF90FF60F8C2FE7FF8B5" author="Simon, E." box="[234,404,1803,1827]" pageId="4" pageNumber="5" pagination="21" refId="ref41646" refString="Simon, E. (1903) Descriptions d'arachnides nouveaux. Annales de la Societe Entomologique de Belgique, 47, 21 ‾ 39. https: // doi. org / 10.5962 / bhl. part. 25299" type="journal article" year="1903">Simon, 1903: 30</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<typeStatus id="FF7F28D13D77FF90FE17F8C5FE22F8B2" box="[413,457,1804,1828]" pageId="4" pageNumber="5">type</typeStatus>
species:
<treatmentCitation id="A165B0623D77FF90FDA9F8C5FC94F8B5" author="Simon, E." box="[547,895,1803,1827]" page="30" pageId="4" pageNumber="5" year="1903">
<taxonomicName id="E7C4EDF03D77FF90FDA9F8C5FC94F8B5" ID-CoL="765NH" authority="Simon, 1903: 30" authorityName="Simon. Additionally" authorityPageNumber="30" authorityYear="1903" box="[547,895,1803,1827]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fulvastra">
<emphasis id="12B04A613D77FF90FDA9F8C5FD3AF8B5" box="[547,721,1803,1827]" italics="true" pageId="4" pageNumber="5">Patrera fulvastra</emphasis>
<bibRefCitation id="4455EB823D77FF90FD5CF8C2FC94F8B5" author="Simon, E." box="[726,895,1803,1827]" pageId="4" pageNumber="5" pagination="21" refId="ref41646" refString="Simon, E. (1903) Descriptions d'arachnides nouveaux. Annales de la Societe Entomologique de Belgique, 47, 21 ‾ 39. https: // doi. org / 10.5962 / bhl. part. 25299" type="journal article" year="1903">Simon, 1903: 30</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="A165B0623D77FF90FC03F8C2FB60F8B5" author="Petrunkevitch, A." box="[905,1163,1803,1827]" page="505" pageId="4" pageNumber="5" year="1911">
<bibRefCitation id="4455EB823D77FF90FC03F8C2FB60F8B5" author="Petrunkevitch, A." box="[905,1163,1803,1827]" pageId="4" pageNumber="5" pagination="1" refId="ref41410" refString="Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1 ‾ 791." type="journal article" year="1911">Petrunkevitch (1911: 505</bibRefCitation>
</treatmentCitation>
; 1928: 163);
<treatmentCitation id="A165B0623D77FF90FA84F8C5FF1BF8D4" author="Roewer, C. F." page="544" pageId="4" pageNumber="5" year="1954">
<bibRefCitation id="4455EB823D77FF90FA84F8C5FF1BF8D4" author="Roewer, C. F." pageId="4" pageNumber="5" refId="ref41464" refString="Roewer, C. F. (1954) Katalog der Araneae von 1758 bis 1940. Vol. 2. Institut royal des sciences naturelles de Belgique, Bruxelles, 923 pp." type="book" year="1954">Roewer 1954: 544</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="A165B0623D77FF90FF71F8E3FE2BF8D4" author="Bonnet, P." box="[251,448,1834,1858]" page="3439" pageId="4" pageNumber="5" year="1958">
<bibRefCitation id="4455EB823D77FF90FF71F8E3FE2BF8D4" author="Bonnet, P." box="[251,448,1834,1858]" pageId="4" pageNumber="5" pagination="3027" refId="ref40702" refString="Bonnet, P. (1958) Bibliographia araneorum. Douladoure Toulouse, 2 (4), 3027 ‾ 4230." type="journal article" year="1958">Bonnet 1958: 3439</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="A165B0623D77FF90FE40F8E3FD67F8D4" author="Brescovit, A. D." box="[458,652,1834,1858]" page="31" pageId="4" pageNumber="5" year="1997">
<bibRefCitation id="4455EB823D77FF90FE40F8E3FD67F8D4" author="Brescovit, A. D." box="[458,652,1834,1858]" pageId="4" pageNumber="5" pagination="1 - 187" refId="ref40616" refString="Brescovit, A. D. (1997) Revis &quot; o de Anyphaeninae Bertkau a nivel de generos na regi &quot; o neotropical (Araneae, Anyphaenidae). Revista Brasileira de Zoologia, 13, 1 - 187." type="journal article" year="1997">Brescovit 1997: 31</bibRefCitation>
</treatmentCitation>
̅36;
<treatmentCitation id="A165B0623D77FF90FD4AF8E3FC27F8D4" ID-DOI="http://doi.org/10.5281/zenodo.6081753" ID-Zenodo-Dep="6081753" author="Duperre, N. &amp; Tapia, E." box="[704,972,1834,1858]" httpUri="http://treatment.plazi.org/id/F319EF090703FB18FD267D278A58C69C" page="30" pageId="4" pageNumber="5" year="2016">
<bibRefCitation id="4455EB823D77FF90FD4AF8E3FC27F8D4" author="Duperre, N. &amp; Tapia, E." box="[704,972,1834,1858]" pageId="4" pageNumber="5" pagination="1 - 50" refId="ref40935" refString="Duperre, N. &amp; Tapia, E. (2016) Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Choco forest of Ecuador, with the description of thirteen new species. European Journal of Taxonomy, 255, 1 - 50. https: // doi. org / 10.5852 / ejt. 2016.255" type="journal article" year="2016">Dupérré &amp; Tapia 2016: 30</bibRefCitation>
</treatmentCitation>
̅44.
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</paragraph>
</subSubSection>
<subSubSection id="68DEC5F83D77FF91FF1DF8B9FCB0FE1F" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="207B96733D77FF91FF1DF8B9FCB0FE1F" blockId="4.[151,1437,1904,2039]" lastBlockId="5.[151,1437,151,393]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="12B04A613D77FF90FF1DF8B9FE6DF81C" bold="true" box="[151,390,1904,1930]" pageId="4" pageNumber="5">Emended diagnosis.</emphasis>
<taxonomicName id="E7C4EDF03D77FF90FE07F8B8FE08F81C" authorityName="Simon" authorityYear="1903" box="[397,483,1905,1930]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D77FF90FE07F8B8FE08F81C" box="[397,483,1905,1930]" italics="true" pageId="4" pageNumber="5">Patrera</emphasis>
</taxonomicName>
species can be distinguished by the following characters combination: carapace subrectangular, tracheal spiracle between the middle of the abdomen and the epigastric groove, lateral margin of endites concave, male palp with subtegular projection, prominent in
<taxonomicName id="E7C4EDF03D77FF90FC00F871FC05F844" authorityName="Simon. Additionally" authorityYear="1903" box="[906,1006,1976,2002]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fulvastra">
<emphasis id="12B04A613D77FF90FC00F871FC05F844" box="[906,1006,1976,2002]" italics="true" pageId="4" pageNumber="5">fulvastra</emphasis>
</taxonomicName>
and
<taxonomicName id="E7C4EDF03D77FF90FBADF871FB85F844" authorityName="Martínez &amp; Brescovit &amp; Villarreal &amp; Oliveira" authorityYear="2021" box="[1063,1134,1976,2002]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="12B04A613D77FF90FBADF871FB85F844" box="[1063,1134,1976,2002]" italics="true" pageId="4" pageNumber="5">florezi</emphasis>
</taxonomicName>
species groups or reduced in
<taxonomicName id="E7C4EDF03D77FF90FF3FF815FF15F860" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[181,254,2012,2038]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philipi">
<emphasis id="12B04A613D77FF90FF3FF815FF15F860" box="[181,254,2012,2038]" italics="true" pageId="4" pageNumber="5">philipi</emphasis>
</taxonomicName>
species group. Additionally, the latter group is provided with a sclerotized ventral tegular process (
<figureCitation id="B8FF8AF63D77FF91FAE0F815FF2CFF25" captionStart="FIGURE 22" captionStartId="42.[151,250,1809,1833]" captionTargetBox="[272,1303,181,1785]" captionTargetId="figure-27@42.[272,1316,181,1785]" captionTargetPageId="42" captionText="FIGURE 22A–D. Patrera armata (Chickering, 1940). Male (ICN-Ar-10608): A habitus, dorsal view; B Chelicerae ventral view; C left palp, retroventral view (Arrow: indicates projection basal of the tibia); D palp, ventral view; E palp, retrolateral view. Female (ICN-Ar-5481): B habitus, dorsal view; G epigynum, ventral view; F epigynum, dorsal view. Abbreviations: BE, base of the embolus; Cvp, cheliceral ventral projection; dps, dorsal projection of the subtegulum; E, embolus; FD, fertilization ducts; H, hood; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; ST, subtegulum; T, tegulum. Scale bars: A–B: 2mm. C: 1mm; D–F: 0.5mm; G–H: 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.4468433" httpUri="https://zenodo.org/record/4468433/files/figure.png" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Figs 22E</figureCitation>
̅F; 23C̅E; 24E̅G; 25C̅E; 26D̅F;
<bibRefCitation id="4455EB823D76FF91FDE7FF5EFC64FF27" author="Duperre, N. &amp; Tapia, E." box="[621,911,151,177]" pageId="5" pageNumber="6" pagination="1 - 50" refId="ref40935" refString="Duperre, N. &amp; Tapia, E. (2016) Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Choco forest of Ecuador, with the description of thirteen new species. European Journal of Taxonomy, 255, 1 - 50. https: // doi. org / 10.5852 / ejt. 2016.255" type="journal article" year="2016">Dupérré &amp; Tapia 2016:30</bibRefCitation>
̅41, figs 41, 46). Females can be distinguished from
<taxonomicName id="E7C4EDF03D76FF91FF5CFF72FE76FF43" authority="Brescovit" authorityName="Brescovit" box="[214,413,187,213]" class="Arachnida" family="Anyphaenidae" genus="Katissa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D76FF91FF5CFF72FEC1FF42" box="[214,298,187,212]" italics="true" pageId="5" pageNumber="6">Katissa</emphasis>
Brescovit
</taxonomicName>
and
<taxonomicName id="E7C4EDF03D76FF91FE5DFF72FCFCFF43" authority="Duperre &amp; Tapia" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[471,791,187,213]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D76FF91FE5DFF72FDA4FF43" box="[471,591,187,213]" italics="true" pageId="5" pageNumber="6">Shuyushka</emphasis>
Dupérré &amp; Tapia
</taxonomicName>
by having a distinct sclerotized hood situated on the anterior region of the epigynum, which can be triangular, transversal or scape-shaped (
<figureCitation id="B8FF8AF63D76FF91FBEEFF16FB57FF6D" box="[1124,1212,223,251]" captionStart="FIGURE 1" captionStartId="6.[151,250,1808,1832]" captionTargetBox="[237,1345,181,1784]" captionTargetId="figure-27@6.[237,1350,181,1785]" captionTargetPageId="6" captionText="FIGURE 1A–G. Patrera fulvastra Simon, 1903. Male (ICN-Ar-5581): A habitus, dorsal view; C left palp, retroventral view; D palp, ventral view; E palp, retrolateral view. Female (MCN 24208): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Abbreviations: A, atrium; BE, base of the embolus; CD, copulatory ducts; dps, dorsal projection of the subtegulum; H, hood; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; ST, subtegulum; SR, seminal receptacles; T, tegulum. Scale bars:A–B: 3mm; C: 0.8mm; D–E: 1mm; F–G: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4468385" httpUri="https://zenodo.org/record/4468385/files/figure.png" pageId="5" pageNumber="6">Figs 1F</figureCitation>
̅G; 6F̅G; 13F̅G; 14F̅G; 15F̅G; 16F̅G; 17F̅G; 29F̅G;
<bibRefCitation id="4455EB823D76FF91FDE5FECAFCBFFE8B" author="Brescovit, A. D." box="[623,852,259,285]" pageId="5" pageNumber="6" pagination="1 - 187" refId="ref40616" refString="Brescovit, A. D. (1997) Revis &quot; o de Anyphaeninae Bertkau a nivel de generos na regi &quot; o neotropical (Araneae, Anyphaenidae). Revista Brasileira de Zoologia, 13, 1 - 187." type="journal article" year="1997">Brescovit 1997: 146</bibRefCitation>
, figs 51̅52;
<bibRefCitation id="4455EB823D76FF91FC62FECAFAFBFE8B" author="Duperre, N. &amp; Tapia, E." box="[1000,1296,259,285]" pageId="5" pageNumber="6" pagination="1 - 50" refId="ref40935" refString="Duperre, N. &amp; Tapia, E. (2016) Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Choco forest of Ecuador, with the description of thirteen new species. European Journal of Taxonomy, 255, 1 - 50. https: // doi. org / 10.5852 / ejt. 2016.255" type="journal article" year="2016">Dupérré &amp; Tapia 2016: 30</bibRefCitation>
̅41, figs 44, 52, 58, 63). Additionaly,
<taxonomicName id="E7C4EDF03D76FF91FE3DFEEEFDE6FED6" authorityName="Simon" authorityYear="1903" box="[439,525,295,320]" class="Arachnida" family="Anyphaenidae" genus="Patrera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D76FF91FE3DFEEEFDE6FED6" box="[439,525,295,320]" italics="true" pageId="5" pageNumber="6">Patrera</emphasis>
</taxonomicName>
species also can be distinguished from those of
<taxonomicName id="E7C4EDF03D76FF91FBB6FEEEFB5FFED7" authorityName="Duperre &amp; Tapia" authorityYear="2016" box="[1084,1204,295,321]" class="Arachnida" family="Anyphaenidae" genus="Shuyushka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D76FF91FBB6FEEEFB5FFED7" box="[1084,1204,295,321]" italics="true" pageId="5" pageNumber="6">Shuyushka</emphasis>
</taxonomicName>
by the absence of a ventral patellar apophysis and from those of
<taxonomicName id="E7C4EDF03D76FF91FD01FE82FD0CFEF2" authorityName="Brescovit" authorityYear="1993" box="[651,743,331,356]" class="Arachnida" family="Anyphaenidae" genus="Lepajan" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="12B04A613D76FF91FD01FE82FD0CFEF2" box="[651,743,331,356]" italics="true" pageId="5" pageNumber="6">Lepajan</emphasis>
</taxonomicName>
by the presence of chilum (see
<bibRefCitation id="4455EB823D76FF91FBC0FE82FA80FEF3" author="Duperre, N. &amp; Tapia, E." box="[1098,1387,331,357]" pageId="5" pageNumber="6" pagination="1 - 50" refId="ref40935" refString="Duperre, N. &amp; Tapia, E. (2016) Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Choco forest of Ecuador, with the description of thirteen new species. European Journal of Taxonomy, 255, 1 - 50. https: // doi. org / 10.5852 / ejt. 2016.255" type="journal article" year="2016">Dupérré &amp; Tapia 2016:21</bibRefCitation>
̅29, figs 28̅29, 33̅34, 38̅39;
<bibRefCitation id="4455EB823D76FF91FE40FEA6FD4CFE1F" author="Brescovit, A. D." box="[458,679,366,393]" pageId="5" pageNumber="6" pagination="693" refId="ref40576" refString="Brescovit, A. D. (1993) Lepajan, um genero novo de aranhas neotropicais da familia Anyphaenidae (Arachnida, Araneae). Revista Brasileira de Entomologia, 37 (4), 693 ‾ 703." type="journal article" year="1993">Brescovit 1993:125</bibRefCitation>
̅128, figs 1̅8).
</paragraph>
</subSubSection>
</treatment>
</document>
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