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<mods:title id="EBD832BB883E0E6CFC352192DC2D692E">Systematics of Indobaetis Müller-Liebenau &amp; Morihara 1982, and related implications for some other Baetidae genera (Ephemeroptera)</mods:title>
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<treatment id="A40F87A394753B72FF58FE52FEB7A745" ID-DOI="http://doi.org/10.5281/zenodo.6137746" ID-GBIF-Taxon="119511064" ID-Zenodo-Dep="6137746" LSID="urn:lsid:plazi:treatment:A40F87A394753B72FF58FE52FEB7A745" httpUri="http://treatment.plazi.org/id/A40F87A394753B72FF58FE52FEB7A745" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214">
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<paragraph id="2C1936B594753B71FF58FE52FD27A589" blockId="5.[151,725,474,534]" box="[151,725,474,500]" pageId="5" pageNumber="214">
<heading id="775181D994753B71FF58FE52FD27A589" bold="true" box="[151,725,474,500]" fontSize="11" level="1" pageId="5" pageNumber="214" reason="1">
<taxonomicName id="EBA64D3694753B71FF58FE52FD27A589" ID-CoL="5483" ID-ENA="1569278" authority="Muller-Liebenau &amp; Morihara 1982" authorityName="Muller-Liebenau &amp; Morihara" authorityYear="1982" box="[151,725,474,500]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FF58FE52FD27A589" bold="true" box="[151,725,474,500]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF58FE52FEE4A589" bold="true" box="[151,278,474,500]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
<bibRefCitation id="48374B4494753B71FED2FE52FD27A589" author="Muller-Liebenau" box="[285,725,474,500]" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
</emphasis>
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<paragraph id="2C1936B594753B71FF58FE75FEEBA66B" blockId="5.[151,725,474,534]" box="[151,281,509,534]" pageId="5" pageNumber="214">
(
<figureCitation id="B49D2A3094753B71FF50FE75FEE2A66B" box="[159,272,509,534]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStart-10="FIGURES 60 – 65" captionStart-11="FIGURES 66 – 70" captionStart-12="FIGURES 71 – 75" captionStart-2="FIGURES 11 – 17" captionStart-3="FIGURES 18 – 22" captionStart-4="FIGURES 23 – 29" captionStart-5="FIGURES 30 – 35" captionStart-6="FIGURES 36 – 39" captionStart-7="FIGURES 40 – 45" captionStart-8="FIGURES 46 – 52" captionStart-9="FIGURES 53 – 59" captionStartId-0="1.[151,275,1369,1394]" captionStartId-1="2.[151,264,1953,1976]" captionStartId-10="13.[151,264,1684,1707]" captionStartId-11="14.[151,264,1648,1671]" captionStartId-12="16.[151,264,1726,1749]" captionStartId-2="3.[151,264,1827,1850]" captionStartId-3="4.[151,264,1610,1633]" captionStartId-4="7.[151,264,1729,1752]" captionStartId-5="8.[151,264,1685,1708]" captionStartId-6="9.[151,264,1558,1581]" captionStartId-7="10.[151,264,1358,1381]" captionStartId-8="11.[151,264,1945,1968]" captionStartId-9="12.[151,264,1810,1833]" captionTargetBox-0="[168,1419,157,1326]" captionTargetBox-1="[159,1427,193,1930]" captionTargetBox-10="[151,1436,197,1662]" captionTargetBox-11="[151,1436,447,1623]" captionTargetBox-12="[151,1436,195,1701]" captionTargetBox-2="[151,1436,197,1805]" captionTargetBox-3="[152,1436,197,1589]" captionTargetBox-4="[158,1424,204,1705]" captionTargetBox-5="[151,1436,195,1661]" captionTargetBox-6="[151,1436,195,1508]" captionTargetBox-7="[151,1436,198,1334]" captionTargetBox-8="[167,1424,193,1919]" captionTargetBox-9="[151,1435,195,1772]" captionTargetId-0="figure@1.[168,1420,157,1348]" captionTargetId-1="figure@2.[159,1427,193,1932]" captionTargetId-10="figure@13.[151,1436,197,1663]" captionTargetId-11="figure@14.[151,1436,447,1624]" captionTargetId-12="figure@16.[151,1436,195,1703]" captionTargetId-2="figure@3.[151,1436,197,1807]" captionTargetId-3="figure@4.[151,1436,197,1590]" captionTargetId-4="figure@7.[151,1436,195,1705]" captionTargetId-5="figure@8.[151,1436,195,1662]" captionTargetId-6="figure@9.[151,1436,195,1538]" captionTargetId-7="figure@10.[151,1436,195,1335]" captionTargetId-8="figure@11.[167,1424,187,1929]" captionTargetId-9="figure@12.[151,1436,193,1789]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-10="13" captionTargetPageId-11="14" captionTargetPageId-12="16" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." captionText-1="FIGURES 5 – 10. Indobaetis costai. 5, fore tarsus of female subimago; 6 – 8, fore tarsus of male subimago and its enlarged base; 9, scales on abdominal tergum; 10, apex of left maxilla, dorsal view. Abbreviations: ds 1 and ds 2, first and second dentisetae." captionText-10="FIGURES 60 – 65. Indobaetis microfolius sp. n. 60, abdominal sterna and terga VI – VII; 61, denticles on anterior-lateral corner of sternum VII; 62, tergalius VII (ventral view); 63, surface of tergum VI; 64, 65, posterior margin and scales on abdominal tergum VI." captionText-11="FIGURES 66 – 70. Indobaetis microfolius sp. n. (holotype). 66 – 69, details of larval middle leg; 70, abdominal sterna and terga of male subimago, spread on slide." captionText-12="FIGURES 71 – 75. Indobaetis microfolius sp. n. 71, hypopharynx and superlingua; 72 – 75, abdominal segment X with paracercus and bases of cerci: 72, 74, 75, ventral view; 73, dorsal view." captionText-2="FIGURES 11 – 17. Indobaetis costai, middle leg of subimago and its details. Abbreviations: bml, blunt microlepides; mt, microtrichiae; pml, pointed microlepides; t 1 &amp; 2, fused first and second tarsomeres; t 3 – t 5, tarsomeres 3 – 5." captionText-3="FIGURES 18 – 22. Indobaetis costai. 18, subimaginal exuviae of left half of mesonotum; 19, subimaginal exuviae of left subalar sclerite, postsubalar sclerite and lateropostnotal crest; 20, abdominal segments I – VI of male imago, spread on slide; 21, genitals of male imago (intersegmental ventral muscles taken away and unpaired sterno-styligeral muscle visible though integument); 22, part of larval fore leg." captionText-4="FIGURES 23 – 29. Indobaetis costai. 23 – 25, male imago: 23, wing; 24, genitals (ventral view, muscles of middle and left parts shown by interrupted lines); 25, head and dorsal outline of pronotum and anteronotal protuberance, lateral view. 26, head of female imago, dorsal view. 27, subimaginal exuviae of right unistyliger and gonostylus; 28, subimaginal genitals developing under larval cuticle; 29, the same at earlier phase. Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sternostyligeral muscle; m. IX – X, intersegmental ventral muscles going from base of 9 th abdominal segment to base of 10 th abdominal segment; us, unistyliger." captionText-5="FIGURES 30 – 35. Indobaetis costai, eggs and their enlarged micropilar areas." captionText-6="FIGURES 36 – 39. Posterior view on 10 th abdominal segment (cerci and paracercus taken away). 36, Nigrobaetis (Takobia) maxillaris; 37, undescribed species of Papuanatula from Baliem Valley (Papua province of Indonesia); 38, 39, two specimens of Indobaetis costai." captionText-7="FIGURES 40 – 45. Indobaetis spp., larvae. 40, I. starmuehlneri, head; 41 – 44, I. costai: 41, head; 42, claw of left leg, ventral view; 43, exuviae of right half of pronotum and mesonotum; 44, exuviae of frons. 45, I. microfolius sp. n., the same." captionText-8="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." captionText-9="FIGURES 53 – 59. Indobaetis microfolius sp. n. 53, margins of larval abdominal segments IV – VI with tergalii; 54, genitals of male subimago (sterno-styligeral muscle and styliger-gonostylar muscles shown by interrupted lines); 55, larval head; 56, wing venation; 57, head of male subimago; 58, larval claw, posterior view; 59, exuviae of right half of larval pronotum and mesonotum (53, 54, 56 – 58, holotype). Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sterno-styligeral muscle; us, unistyliger." httpUri-0="https://zenodo.org/record/250182/files/figure.png" httpUri-1="https://zenodo.org/record/250183/files/figure.png" httpUri-10="https://zenodo.org/record/250192/files/figure.png" httpUri-11="https://zenodo.org/record/250193/files/figure.png" httpUri-12="https://zenodo.org/record/250194/files/figure.png" httpUri-2="https://zenodo.org/record/250184/files/figure.png" httpUri-3="https://zenodo.org/record/250185/files/figure.png" httpUri-4="https://zenodo.org/record/250186/files/figure.png" httpUri-5="https://zenodo.org/record/250187/files/figure.png" httpUri-6="https://zenodo.org/record/250188/files/figure.png" httpUri-7="https://zenodo.org/record/250189/files/figure.png" httpUri-8="https://zenodo.org/record/250190/files/figure.png" httpUri-9="https://zenodo.org/record/250191/files/figure.png" pageId="5" pageNumber="214">Figs 2–75</figureCitation>
)
</paragraph>
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<subSubSection id="64BC653E94753B72FF58FDCEFAEAA5AD" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214" type="materials_examined">
<paragraph id="2C1936B594753B71FF58FDCEFC7EA623" blockId="5.[151,1436,581,2010]" box="[151,908,581,606]" pageId="5" pageNumber="214">
<typeStatus id="F31D881794753B71FF58FDCEFF3DA623" box="[151,207,582,606]" pageId="5" pageNumber="214">Type</typeStatus>
species: 
<taxonomicName id="EBA64D3694753B71FEFAFDCDFC75A623" authority="Muller-Liebenau &amp; Morihara 1982" authorityName="Muller-Liebenau &amp; Morihara" authorityYear="1982" box="[309,903,581,606]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FEFAFDCDFE07A620" box="[309,501,581,606]" italics="true" pageId="5" pageNumber="214">Indobaetis costai</emphasis>
<bibRefCitation id="48374B4494753B71FE33FDCDFC75A623" author="Muller-Liebenau" box="[508,903,581,606]" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="2C1936B594753B71FF08FDE1FEF9A6DB" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
General characteristic given to 
<taxonomicName id="EBA64D3694753B71FDE4FDE1FD53A6FF" box="[555,673,617,642]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FDE4FDE1FD53A6FF" box="[555,673,617,642]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="48374B4494753B71FD01FDE1FB9FA6FF" author="Muller-Liebenau" box="[718,1133,617,642]" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara (1982)</bibRefCitation>
should be modified as the following.
</paragraph>
<paragraph id="2C1936B594753B71FF08FD39FDFCA6B7" blockId="5.[151,1436,581,2010]" box="[199,526,689,714]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FD39FDFCA6B7" bold="true" box="[199,526,689,714]" pageId="5" pageNumber="214">
Apomorphies of 
<taxonomicName id="EBA64D3694753B71FE41FD39FDF5A6B7" box="[398,519,689,714]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FE41FD39FDF5A6B7" bold="true" box="[398,519,689,714]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="2C1936B594753B71FF08FD5DFF4DA727" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FD5DFF15A693" bold="true" box="[199,231,725,750]" pageId="5" pageNumber="214">(1)</emphasis>
Each mandible has incisor and kinetodontium completely fused together forming an integral plate with cutting edge widened, evenly dentate and vaulted ventrally, so that its part corresponding to incisor is less projected medially than its part corresponding to kinetodontium (
<figureCitation id="B49D2A3094753B71FCDDFC95FC64A74B" box="[786,918,797,822]" captionStart="FIGURES 46 – 52" captionStartId="11.[151,264,1945,1968]" captionTargetBox="[167,1424,193,1919]" captionTargetId="figure@11.[167,1424,187,1929]" captionTargetPageId="11" captionText="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." httpUri="https://zenodo.org/record/250190/files/figure.png" pageId="5" pageNumber="214">Figs 46–48</figureCitation>
; 
<bibRefCitation id="48374B4494753B71FC6BFC95FAC5A74B" author="Muller-Liebenau" box="[932,1335,797,822]" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
: 
<figureCitation id="B49D2A3094753B71FA8BFC95FA7BA74B" box="[1348,1417,797,822]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,275,1369,1394]" captionTargetBox="[168,1419,157,1326]" captionTargetId="figure@1.[168,1420,157,1348]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." httpUri="https://zenodo.org/record/250182/files/figure.png" pageId="5" pageNumber="214">Figs 1</figureCitation>
f, 2f).
</paragraph>
<paragraph id="2C1936B594753B71FF08FCEDFDA0A7DF" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FCEDFF1AA703" bold="true" box="[199,232,869,894]" pageId="5" pageNumber="214">(2)</emphasis>
Prostheca of right mandible is slender and pointed, not bifurcate (
<figureCitation id="B49D2A3094753B71FC08FCEDFBBAA703" box="[967,1096,869,894]" captionStart="FIGURES 46 – 52" captionStartId="11.[151,264,1945,1968]" captionTargetBox="[167,1424,193,1919]" captionTargetId="figure@11.[167,1424,187,1929]" captionTargetPageId="11" captionText="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." httpUri="https://zenodo.org/record/250190/files/figure.png" pageId="5" pageNumber="214">Figs 47, 48</figureCitation>
; 
<bibRefCitation id="48374B4494753B71FB9BFCEDFF21A7DF" author="Muller-Liebenau" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
: 
<figureCitation id="B49D2A3094753B71FF12FC01FED2A7DF" box="[221,288,905,930]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,275,1369,1394]" captionTargetBox="[168,1419,157,1326]" captionTargetId="figure@1.[168,1420,157,1348]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." httpUri="https://zenodo.org/record/250182/files/figure.png" pageId="5" pageNumber="214">Figs 1</figureCitation>
f, 2f) (see Discussion, 2.4).
</paragraph>
<paragraph id="2C1936B594753B71FF08FC25FB52A02B" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FC25FF1AA7BB" bold="true" box="[199,232,941,966]" pageId="5" pageNumber="214">(3)</emphasis>
Maxilla has 2 dentisetae: 1st dentiseta (most distal) is thick and adjacent to maxillary canines (as in many other 
<taxonomicName id="EBA64D3694753B71FF17FC59FECDA797" box="[216,319,977,1002]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
); 2nd dentiseta is sharply arched and pressed to the first dentiseta, so that its middle part is hidden between the first dentiseta and the canines; 3rd dentiseta is absent; setae of median-dorsal row form a compact bunch, basally adjacent to the second dentiseta and distally diverging from it (
<figureCitation id="B49D2A3094753B71FBCBFB91FBA9A04F" box="[1028,1115,1049,1074]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[159,1427,193,1930]" captionTargetId="figure@2.[159,1427,193,1932]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Indobaetis costai. 5, fore tarsus of female subimago; 6 – 8, fore tarsus of male subimago and its enlarged base; 9, scales on abdominal tergum; 10, apex of left maxilla, dorsal view. Abbreviations: ds 1 and ds 2, first and second dentisetae." httpUri="https://zenodo.org/record/250183/files/figure.png" pageId="5" pageNumber="214">Figs 10</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FBA7FB91FB75A04F" box="[1128,1159,1049,1074]" captionStart="FIGURES 46 – 52" captionStartId="11.[151,264,1945,1968]" captionTargetBox="[167,1424,193,1919]" captionTargetId="figure@11.[167,1424,187,1929]" captionTargetPageId="11" captionText="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." httpUri="https://zenodo.org/record/250190/files/figure.png" pageId="5" pageNumber="214">51</figureCitation>
). In other 
<taxonomicName id="EBA64D3694753B71FB34FB91FAAFA04F" box="[1275,1373,1049,1074]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
three dentisetae, initial for Tridentiseta 
<bibRefCitation id="48374B4494753B71FDDEFBB5FD24A02B" author="Kluge" box="[529,726,1085,1110]" pageId="5" pageNumber="214" refString="Kluge, N. J., Studemann, D., Landolt, P. &amp; Gonser, T. (1995) A reclassification of Siphlonuroidea (Ephemeroptera). Mitteilungen der Schweizerischen entomologischen Gesellschaf, 68, 103 - 132." type="journal article" year="1995">
Kluge 
<emphasis id="1ED2EAA794753B71FD93FBB6FD7FA02B" box="[604,653,1085,1110]" italics="true" pageId="5" pageNumber="214">et al</emphasis>
. 1995
</bibRefCitation>
, are retained (
<bibRefCitation id="48374B4494753B71FCBCFBB5FB61A02B" author="Kluge" box="[883,1171,1085,1110]" pageId="5" pageNumber="214" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp." type="book" year="2004">
Kluge 2004: 
<figureCitation id="B49D2A3094753B71FBC8FBB5FBAEA02B" box="[1031,1116,1085,1110]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[158,1424,204,1705]" captionTargetId="figure@7.[151,1436,195,1705]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Indobaetis costai. 23 – 25, male imago: 23, wing; 24, genitals (ventral view, muscles of middle and left parts shown by interrupted lines); 25, head and dorsal outline of pronotum and anteronotal protuberance, lateral view. 26, head of female imago, dorsal view. 27, subimaginal exuviae of right unistyliger and gonostylus; 28, subimaginal genitals developing under larval cuticle; 29, the same at earlier phase. Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sternostyligeral muscle; m. IX – X, intersegmental ventral muscles going from base of 9 th abdominal segment to base of 10 th abdominal segment; us, unistyliger." httpUri="https://zenodo.org/record/250186/files/figure.png" pageId="5" pageNumber="214">Figs 28</figureCitation>
B, D
</bibRefCitation>
).
</paragraph>
<paragraph id="2C1936B594753B71FF08FBE9FDE4A0E3" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FBE9FF1AA007" bold="true" box="[199,232,1121,1146]" pageId="5" pageNumber="214">(4)</emphasis>
Larval tibia has a regular row of stout setae on outer margin, different from setae of a regular row on outer margin of femur (
<figureCitation id="B49D2A3094753B71FE92FB0DFE41A0E3" box="[349,435,1157,1182]" captionStart="FIGURES 18 – 22" captionStartId="4.[151,264,1610,1633]" captionTargetBox="[152,1436,197,1589]" captionTargetId="figure@4.[151,1436,197,1590]" captionTargetPageId="4" captionText="FIGURES 18 – 22. Indobaetis costai. 18, subimaginal exuviae of left half of mesonotum; 19, subimaginal exuviae of left subalar sclerite, postsubalar sclerite and lateropostnotal crest; 20, abdominal segments I – VI of male imago, spread on slide; 21, genitals of male imago (intersegmental ventral muscles taken away and unpaired sterno-styligeral muscle visible though integument); 22, part of larval fore leg." httpUri="https://zenodo.org/record/250185/files/figure.png" pageId="5" pageNumber="214">Figs 22</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FE70FB0DFDFBA0E3" box="[447,521,1157,1182]" captionStart="FIGURES 66 – 70" captionStartId="14.[151,264,1648,1671]" captionTargetBox="[151,1436,447,1623]" captionTargetId="figure@14.[151,1436,447,1624]" captionTargetPageId="14" captionText="FIGURES 66 – 70. Indobaetis microfolius sp. n. (holotype). 66 – 69, details of larval middle leg; 70, abdominal sterna and terga of male subimago, spread on slide." httpUri="https://zenodo.org/record/250193/files/figure.png" pageId="5" pageNumber="214">67, 68</figureCitation>
).
</paragraph>
<paragraph id="2C1936B594753B71FF08FB21FEA1A09B" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FB21FF1AA0BF" bold="true" box="[199,232,1193,1218]" pageId="5" pageNumber="214">(5)</emphasis>
Larval claw has distal denticle enlarged, with at least one strong seta posteriad of it (
<figureCitation id="B49D2A3094753B71FB19FB21FAC3A0BF" box="[1238,1329,1193,1218]" captionStart="FIGURES 40 – 45" captionStartId="10.[151,264,1358,1381]" captionTargetBox="[151,1436,198,1334]" captionTargetId="figure@10.[151,1436,195,1335]" captionTargetPageId="10" captionText="FIGURES 40 – 45. Indobaetis spp., larvae. 40, I. starmuehlneri, head; 41 – 44, I. costai: 41, head; 42, claw of left leg, ventral view; 43, exuviae of right half of pronotum and mesonotum; 44, exuviae of frons. 45, I. microfolius sp. n., the same." httpUri="https://zenodo.org/record/250189/files/figure.png" pageId="5" pageNumber="214">Figs 42</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FAF0FB21FAAEA0BF" box="[1343,1372,1193,1218]" captionStart="FIGURES 53 – 59" captionStartId="12.[151,264,1810,1833]" captionTargetBox="[151,1435,195,1772]" captionTargetId="figure@12.[151,1436,193,1789]" captionTargetPageId="12" captionText="FIGURES 53 – 59. Indobaetis microfolius sp. n. 53, margins of larval abdominal segments IV – VI with tergalii; 54, genitals of male subimago (sterno-styligeral muscle and styliger-gonostylar muscles shown by interrupted lines); 55, larval head; 56, wing venation; 57, head of male subimago; 58, larval claw, posterior view; 59, exuviae of right half of larval pronotum and mesonotum (53, 54, 56 – 58, holotype). Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sterno-styligeral muscle; us, unistyliger." httpUri="https://zenodo.org/record/250191/files/figure.png" pageId="5" pageNumber="214">58</figureCitation>
) (see Discussion, 2.7).
</paragraph>
<paragraph id="2C1936B594753B71FF08FB79FF15A153" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FB79FF1AA177" bold="true" box="[199,232,1265,1290]" pageId="5" pageNumber="214">(6)</emphasis>
<collectingCountry id="54B1762594753B71FF3FFB79FEDAA177" box="[240,296,1265,1290]" name="India" pageId="5" pageNumber="214">Hind</collectingCountry>
wings are absent; larva has no vestiges of hind protoptera (
<bibRefCitation id="48374B4494753B71FC01FB79FA92A177" author="Muller-Liebenau" box="[974,1376,1265,1290]" pageId="5" pageNumber="214" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
: 
<figureCitation id="B49D2A3094753B71FAA3FB79FF51A153" captionStart="FIGURES 1 – 4" captionStartId="1.[151,275,1369,1394]" captionTargetBox="[168,1419,157,1326]" captionTargetId="figure@1.[168,1420,157,1348]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." httpUri="https://zenodo.org/record/250182/files/figure.png" pageId="5" pageNumber="214">
Figs 
<date id="5818107594753B71FF58FA9DFF51A153" box="[151,163,1301,1326]" pageId="5" pageNumber="214">1</date>
</figureCitation>
i, 
<quantity id="EB5E9B5094753B71FF74FA9EFF28A153" box="[187,218,1302,1326]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="5" pageNumber="214" unit="g" value="2.0">2g</quantity>
).
</paragraph>
<paragraph id="2C1936B594753B71FF08FAB1FC43A12C" blockId="5.[151,1436,581,2010]" box="[199,945,1337,1362]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FAB1FF1AA12F" bold="true" box="[199,232,1337,1362]" pageId="5" pageNumber="214">(7)</emphasis>
Tergalii of first pair are lost, only 6 pairs of tergalii are present.
</paragraph>
<paragraph id="2C1936B594753B71FF08FAD5FCD8A1E7" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FAD5FF1AA10B" bold="true" box="[199,232,1373,1398]" pageId="5" pageNumber="214">(8)</emphasis>
In mature larva buds of subimaginal gonostyli are folded by « 
<taxonomicName id="EBA64D3694753B71FC26FAD5FB9FA10B" box="[1001,1133,1373,1398]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FC26FAD5FB9FA10B" box="[1001,1133,1373,1398]" italics="true" pageId="5" pageNumber="214">Labiobaetis</emphasis>
</taxonomicName>
- 
<typeStatus id="F31D881794753B71FBBBFAD6FB55A10B" box="[1140,1191,1374,1398]" pageId="5" pageNumber="214">type</typeStatus>
» (
<figureCitation id="B49D2A3094753B71FB06FAD5FAABA10B" box="[1225,1369,1373,1398]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[158,1424,204,1705]" captionTargetId="figure@7.[151,1436,195,1705]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Indobaetis costai. 23 – 25, male imago: 23, wing; 24, genitals (ventral view, muscles of middle and left parts shown by interrupted lines); 25, head and dorsal outline of pronotum and anteronotal protuberance, lateral view. 26, head of female imago, dorsal view. 27, subimaginal exuviae of right unistyliger and gonostylus; 28, subimaginal genitals developing under larval cuticle; 29, the same at earlier phase. Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sternostyligeral muscle; m. IX – X, intersegmental ventral muscles going from base of 9 th abdominal segment to base of 10 th abdominal segment; us, unistyliger." httpUri="https://zenodo.org/record/250186/files/figure.png" pageId="5" pageNumber="214">Figs 28, 29</figureCitation>
); this character is known only for 
<taxonomicName id="EBA64D3694753B71FE1DFA0AFDDEA1E4" box="[466,556,1410,1433]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FE1DFA0AFDDEA1E4" box="[466,556,1410,1433]" italics="true" pageId="5" pageNumber="214">I. costai</emphasis>
</taxonomicName>
(see Discussion, 2.10)
</paragraph>
<paragraph id="2C1936B594753B71FF08FA2DFCB7A257" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08FA2DFF1AA1C3" bold="true" box="[199,232,1445,1470]" pageId="5" pageNumber="214">(9)</emphasis>
On fore leg of female subimago and imago 1st tarsomere is completely fused with 2nd tarsomere, so the tarsus is 4-segmented (
<figureCitation id="B49D2A3094753B71FE56FA41FDF2A19F" box="[409,512,1481,1506]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStartId-0="1.[151,275,1369,1394]" captionStartId-1="2.[151,264,1953,1976]" captionTargetBox-0="[168,1419,157,1326]" captionTargetBox-1="[159,1427,193,1930]" captionTargetId-0="figure@1.[168,1420,157,1348]" captionTargetId-1="figure@2.[159,1427,193,1932]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." captionText-1="FIGURES 5 – 10. Indobaetis costai. 5, fore tarsus of female subimago; 6 – 8, fore tarsus of male subimago and its enlarged base; 9, scales on abdominal tergum; 10, apex of left maxilla, dorsal view. Abbreviations: ds 1 and ds 2, first and second dentisetae." httpUri-0="https://zenodo.org/record/250182/files/figure.png" httpUri-1="https://zenodo.org/record/250183/files/figure.png" pageId="5" pageNumber="214">Figs 2–5</figureCitation>
), like tarsi of middle and hind legs (
<figureCitation id="B49D2A3094753B71FC55FA41FC1EA19F" box="[922,1004,1481,1506]" captionStart="FIGURES 11 – 17" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[151,1436,197,1805]" captionTargetId="figure@3.[151,1436,197,1807]" captionTargetPageId="3" captionText="FIGURES 11 – 17. Indobaetis costai, middle leg of subimago and its details. Abbreviations: bml, blunt microlepides; mt, microtrichiae; pml, pointed microlepides; t 1 &amp; 2, fused first and second tarsomeres; t 3 – t 5, tarsomeres 3 – 5." httpUri="https://zenodo.org/record/250184/files/figure.png" pageId="5" pageNumber="214">Fig. 11</figureCitation>
). This character is known for 
<taxonomicName id="EBA64D3694753B71FA8EFA42FA6EA19C" box="[1345,1436,1482,1505]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FA8EFA42FA6EA19C" box="[1345,1436,1482,1505]" italics="true" pageId="5" pageNumber="214">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694753B71FF07FA66FEADA27B" box="[200,351,1517,1542]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794753B71FF07FA66FEADA27B" box="[200,351,1517,1542]" italics="true" pageId="5" pageNumber="214">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94753B71FEA7FA66FE6FA278" box="[360,413,1518,1541]" pageId="5" pageNumber="214" rank="species">sp.n.</taxonomicNameLabel>
In most other 
<taxonomicName id="EBA64D3694753B71FD87FA65FD58A27B" box="[584,682,1517,1542]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
fore tarsus of female subimago and imago has 1st tarsomere well separated from 2nd tarsomere (
<figureCitation id="B49D2A3094753B71FE3BF999FDC6A257" box="[500,564,1553,1578]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,275,1369,1394]" captionTargetBox="[168,1419,157,1326]" captionTargetId="figure@1.[168,1420,157,1348]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Fore tarsus of female. 1, female subimago of Nigrobaetis (Takobia) muticus; 2, female subimago of Indobaetis costai; 3, imago of the same specimen; 4, female subimaginal tarsus of Indobaetis microfolius sp. n. visible through larval cuticle. Abbreviations: t 1 – t 5, tarsomeres 1 – 5; t 1 &amp; 2, fused first and second tarsomeres." httpUri="https://zenodo.org/record/250182/files/figure.png" pageId="5" pageNumber="214">Fig. 1</figureCitation>
) (see Discussion, 2.12).
</paragraph>
<paragraph id="2C1936B594753B71FF08F9BDFC9BA233" blockId="5.[151,1436,581,2010]" box="[199,873,1589,1614]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08F9BDFC9BA233" bold="true" box="[199,873,1589,1614]" pageId="5" pageNumber="214">
Character of 
<taxonomicName id="EBA64D3694753B71FEA8F9BDFE12A233" box="[359,480,1589,1614]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FEA8F9BDFE12A233" bold="true" box="[359,480,1589,1614]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
</taxonomicName>
of unclear phylogenetic polarity.
</emphasis>
</paragraph>
<paragraph id="2C1936B594753B71FF08F9D1FEA8A2C7" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08F9D1FF04A20F" bold="true" box="[199,246,1625,1650]" pageId="5" pageNumber="214">(10)</emphasis>
In subimago, 1st and 5th tarsomeres are covered mainly with pointed microlepides, other tarsomeres mainly with blunt microlepides (
<figureCitation id="B49D2A3094753B71FDCAF9F5FD98A2EB" box="[517,618,1661,1686]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[159,1427,193,1930]" captionTargetId="figure@2.[159,1427,193,1932]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Indobaetis costai. 5, fore tarsus of female subimago; 6 – 8, fore tarsus of male subimago and its enlarged base; 9, scales on abdominal tergum; 10, apex of left maxilla, dorsal view. Abbreviations: ds 1 and ds 2, first and second dentisetae." httpUri="https://zenodo.org/record/250183/files/figure.png" pageId="5" pageNumber="214">Figs 5–8</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FDB9F9F6FD4DA2EB" box="[630,703,1662,1686]" captionStart="FIGURES 11 – 17" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[151,1436,197,1805]" captionTargetId="figure@3.[151,1436,197,1807]" captionTargetPageId="3" captionText="FIGURES 11 – 17. Indobaetis costai, middle leg of subimago and its details. Abbreviations: bml, blunt microlepides; mt, microtrichiae; pml, pointed microlepides; t 1 &amp; 2, fused first and second tarsomeres; t 3 – t 5, tarsomeres 3 – 5." httpUri="https://zenodo.org/record/250184/files/figure.png" pageId="5" pageNumber="214">11–17</figureCitation>
); this character is known for 
<taxonomicName id="EBA64D3694753B71FBCAF9F6FBADA2E8" box="[1029,1119,1662,1685]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FBCAF9F6FBADA2E8" box="[1029,1119,1662,1685]" italics="true" pageId="5" pageNumber="214">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694753B71FB59F9F6FADEA2EB" box="[1174,1324,1661,1686]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794753B71FB59F9F6FADEA2EB" box="[1174,1324,1661,1686]" italics="true" pageId="5" pageNumber="214">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94753B71FAFCF9F6FA95A2E8" box="[1331,1383,1662,1685]" pageId="5" pageNumber="214" rank="species">sp.n.</taxonomicNameLabel>
(see Discussion, 2.13)
</paragraph>
<paragraph id="2C1936B594753B71FF08F94DFDE3A2A3" blockId="5.[151,1436,581,2010]" box="[199,529,1733,1758]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08F94DFDE3A2A3" bold="true" box="[199,529,1733,1758]" pageId="5" pageNumber="214">
Plesiomorphy of 
<taxonomicName id="EBA64D3694753B71FE5EF94DFDF8A2A3" box="[401,522,1733,1758]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FE5EF94DFDF8A2A3" bold="true" box="[401,522,1733,1758]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="2C1936B594753B71FF08F961FDA1A337" blockId="5.[151,1436,581,2010]" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08F961FF01A37F" bold="true" box="[199,243,1769,1794]" pageId="5" pageNumber="214">(11)</emphasis>
Sterno-styligeral muscle is retained (
<figureCitation id="B49D2A3094753B71FD78F961FCE6A37F" box="[695,788,1769,1794]" captionStart="FIGURES 18 – 22" captionStartId="4.[151,264,1610,1633]" captionTargetBox="[152,1436,197,1589]" captionTargetId="figure@4.[151,1436,197,1590]" captionTargetPageId="4" captionText="FIGURES 18 – 22. Indobaetis costai. 18, subimaginal exuviae of left half of mesonotum; 19, subimaginal exuviae of left subalar sclerite, postsubalar sclerite and lateropostnotal crest; 20, abdominal segments I – VI of male imago, spread on slide; 21, genitals of male imago (intersegmental ventral muscles taken away and unpaired sterno-styligeral muscle visible though integument); 22, part of larval fore leg." httpUri="https://zenodo.org/record/250185/files/figure.png" pageId="5" pageNumber="214">Figs 21</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FCE8F962FCB7A37F" box="[807,837,1770,1794]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[158,1424,204,1705]" captionTargetId="figure@7.[151,1436,195,1705]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Indobaetis costai. 23 – 25, male imago: 23, wing; 24, genitals (ventral view, muscles of middle and left parts shown by interrupted lines); 25, head and dorsal outline of pronotum and anteronotal protuberance, lateral view. 26, head of female imago, dorsal view. 27, subimaginal exuviae of right unistyliger and gonostylus; 28, subimaginal genitals developing under larval cuticle; 29, the same at earlier phase. Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sternostyligeral muscle; m. IX – X, intersegmental ventral muscles going from base of 9 th abdominal segment to base of 10 th abdominal segment; us, unistyliger." httpUri="https://zenodo.org/record/250186/files/figure.png" pageId="5" pageNumber="214">24</figureCitation>
, 
<figureCitation id="B49D2A3094753B71FC97F961FC8AA37F" box="[856,888,1769,1794]" captionStart="FIGURES 53 – 59" captionStartId="12.[151,264,1810,1833]" captionTargetBox="[151,1435,195,1772]" captionTargetId="figure@12.[151,1436,193,1789]" captionTargetPageId="12" captionText="FIGURES 53 – 59. Indobaetis microfolius sp. n. 53, margins of larval abdominal segments IV – VI with tergalii; 54, genitals of male subimago (sterno-styligeral muscle and styliger-gonostylar muscles shown by interrupted lines); 55, larval head; 56, wing venation; 57, head of male subimago; 58, larval claw, posterior view; 59, exuviae of right half of larval pronotum and mesonotum (53, 54, 56 – 58, holotype). Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sterno-styligeral muscle; us, unistyliger." httpUri="https://zenodo.org/record/250191/files/figure.png" pageId="5" pageNumber="214">54</figureCitation>
); this character is known for 
<taxonomicName id="EBA64D3694753B71FB29F962FABAA37C" box="[1254,1352,1770,1793]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FB29F962FABAA37C" box="[1254,1352,1770,1793]" italics="true" pageId="5" pageNumber="214">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694753B71FA43F962FEE4A35B" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794753B71FA43F962FEE4A35B" italics="true" pageId="5" pageNumber="214">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94753B71FED2F886FEA0A358" box="[285,338,1806,1829]" pageId="5" pageNumber="214" rank="species">sp.n.</taxonomicNameLabel>
(in contrast to 
<taxonomicName id="EBA64D3694753B71FE30F885FCD8A358" authority="Baetosternata" authorityName="Baetosternata" box="[511,810,1805,1830]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FE30F885FD71A35B" box="[511,643,1805,1830]" italics="true" pageId="5" pageNumber="214">Labiobaetis</emphasis>
, Baetosternata
</taxonomicName>
and some other taxa, whose sterno-styligeral muscle is completely lost—see Discussion, 2.11).
</paragraph>
<paragraph id="2C1936B594753B72FF08F8DDFAEAA5AD" blockId="5.[151,1436,581,2010]" lastBlockId="6.[151,1437,151,824]" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214">
<emphasis id="1ED2EAA794753B71FF08F8DDFCE9A313" bold="true" box="[199,795,1877,1902]" pageId="5" pageNumber="214">
Characters variable among species of 
<taxonomicName id="EBA64D3694753B71FD55F8DDFCE1A313" box="[666,787,1877,1902]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FD55F8DDFCE1A313" bold="true" box="[666,787,1877,1902]" italics="true" pageId="5" pageNumber="214">Indobaetis</emphasis>
</taxonomicName>
.
</emphasis>
Larval habitus varies from 
<taxonomicName id="EBA64D3694753B71FB90F8DDFB10A313" box="[1119,1250,1877,1902]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FB90F8DDFB10A313" box="[1119,1250,1877,1902]" italics="true" pageId="5" pageNumber="214">Nigrobaetis</emphasis>
</taxonomicName>
-like in 
<taxonomicName id="EBA64D3694753B71FAF0F8DEFA6EA310" box="[1343,1436,1878,1901]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794753B71FAF0F8DEFA6EA310" box="[1343,1436,1878,1901]" italics="true" pageId="5" pageNumber="214">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694753B71FF09F8F2FE88A3EF" box="[198,378,1913,1938]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794753B71FF09F8F2FF3DA3EC" box="[198,207,1914,1937]" italics="true" pageId="5" pageNumber="214">I</emphasis>
. 
<emphasis id="1ED2EAA794753B71FF12F8F1FE88A3EF" box="[221,378,1913,1938]" italics="true" pageId="5" pageNumber="214">starmuehlneri</emphasis>
</taxonomicName>
(i.e., with head bowed ventrally, head and thorax laterally compressed, three caudalii and their primary swimming setae well developed) to 
<taxonomicName id="EBA64D3694753B71FD5BF815FCFAA3CB" box="[660,776,1949,1974]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794753B71FD5BF815FCFAA3CB" box="[660,776,1949,1974]" italics="true" pageId="5" pageNumber="214">Acentrella</emphasis>
</taxonomicName>
-like in 
<taxonomicName id="EBA64D3694753B71FCADF816FC0BA3CB" box="[866,1017,1949,1974]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794753B71FCADF816FC0BA3CB" box="[866,1017,1949,1974]" italics="true" pageId="5" pageNumber="214">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94753B71FBCDF816FBC5A3C8" box="[1026,1079,1950,1973]" pageId="5" pageNumber="214" rank="species">sp.n.</taxonomicNameLabel>
(i.e., head and thorax flattened ventrally, paracercus reduced, and swimming setae lost). Frons varies from narrow and keeled to wide and flat (
<figureCitation id="B49D2A3094763B72FF50FF1FFED2A4CD" box="[159,288,151,176]" captionStart="FIGURES 40 – 45" captionStartId="10.[151,264,1358,1381]" captionTargetBox="[151,1436,198,1334]" captionTargetId="figure@10.[151,1436,195,1335]" captionTargetPageId="10" captionText="FIGURES 40 – 45. Indobaetis spp., larvae. 40, I. starmuehlneri, head; 41 – 44, I. costai: 41, head; 42, claw of left leg, ventral view; 43, exuviae of right half of pronotum and mesonotum; 44, exuviae of frons. 45, I. microfolius sp. n., the same." httpUri="https://zenodo.org/record/250189/files/figure.png" pageId="6" pageNumber="215">Figs 40, 41</figureCitation>
, 
<figureCitation id="B49D2A3094763B72FEE3FF1FFEB9A4CD" box="[300,331,151,176]" captionStart="FIGURES 53 – 59" captionStartId="12.[151,264,1810,1833]" captionTargetBox="[151,1435,195,1772]" captionTargetId="figure@12.[151,1436,193,1789]" captionTargetPageId="12" captionText="FIGURES 53 – 59. Indobaetis microfolius sp. n. 53, margins of larval abdominal segments IV – VI with tergalii; 54, genitals of male subimago (sterno-styligeral muscle and styliger-gonostylar muscles shown by interrupted lines); 55, larval head; 56, wing venation; 57, head of male subimago; 58, larval claw, posterior view; 59, exuviae of right half of larval pronotum and mesonotum (53, 54, 56 – 58, holotype). Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sterno-styligeral muscle; us, unistyliger." httpUri="https://zenodo.org/record/250191/files/figure.png" pageId="6" pageNumber="215">55</figureCitation>
). Maxillary palp, being 2-segmented (as in all Baetungulata), has 2nd segment either incised (in 
<taxonomicName id="EBA64D3694763B72FA43FF11FF28A4A9" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FA43FF11FF28A4A9" italics="true" pageId="6" pageNumber="215">I. costai</emphasis>
</taxonomicName>
), or not (in other species—Fig. 51). Transverse row of fine setae on glossa is either present (in 
<taxonomicName id="EBA64D3694763B72FB33FF35FAA4A4A9" box="[1276,1366,189,212]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FB33FF35FAA4A4A9" box="[1276,1366,189,212]" italics="true" pageId="6" pageNumber="215">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694763B72FA43FF35FEC6A485" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794763B72FA43FF35FEC6A485" italics="true" pageId="6" pageNumber="215">I. starmuehlneri</emphasis>
</taxonomicName>
), or absent (in 
<taxonomicName id="EBA64D3694763B72FE11FF69FD81A485" box="[478,627,223,248]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794763B72FE11FF69FD81A485" box="[478,627,223,248]" italics="true" pageId="6" pageNumber="215">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FDB4FF57FD59A485" box="[635,683,223,248]" pageId="6" pageNumber="215" rank="species">sp.n.</taxonomicNameLabel>
— 
<figureCitation id="B49D2A3094763B72FD04FF57FCEDA485" box="[715,799,223,248]" captionStart="FIGURES 46 – 52" captionStartId="11.[151,264,1945,1968]" captionTargetBox="[167,1424,193,1919]" captionTargetId="figure@11.[167,1424,187,1929]" captionTargetPageId="11" captionText="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." httpUri="https://zenodo.org/record/250190/files/figure.png" pageId="6" pageNumber="215">Fig. 50</figureCitation>
). Labial palp either has structure usual for 
<taxonomicName id="EBA64D3694763B72FB32FF57FAADA485" box="[1277,1375,223,248]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
(in 
<taxonomicName id="EBA64D3694763B72FA43FF69FEC6A560" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794763B72FA43FF69FEC6A560" italics="true" pageId="6" pageNumber="215">I. starmuehlneri</emphasis>
</taxonomicName>
), or has 2nd segment unusually narrowed (in 
<taxonomicName id="EBA64D3694763B72FC81FE8DFC5EA561" box="[846,940,261,284]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FC81FE8DFC5EA561" box="[846,940,261,284]" italics="true" pageId="6" pageNumber="215">I. costai</emphasis>
</taxonomicName>
), or has 2nd and 3rd segments completely fused together and lacking muscle (in 
<taxonomicName id="EBA64D3694763B72FD88FEA1FD2FA53D" box="[583,733,295,320]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794763B72FD88FEA1FD2FA53D" box="[583,733,295,320]" italics="true" pageId="6" pageNumber="215">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FD2AFEAFFCE4A53D" box="[741,790,295,320]" pageId="6" pageNumber="215" rank="species">sp.n.</taxonomicNameLabel>
— 
<figureCitation id="B49D2A3094763B72FCF9FEAFFC79A53D" box="[822,907,295,320]" captionStart="FIGURES 46 – 52" captionStartId="11.[151,264,1945,1968]" captionTargetBox="[167,1424,193,1919]" captionTargetId="figure@11.[167,1424,187,1929]" captionTargetPageId="11" captionText="FIGURES 46 – 52. Indobaetis microfolius sp. n., larval mouth parts. 46, left mandible; 47, 48, right mandibles; 49, labrum; 50, glossa (dorsal view); 51, maxilla (dorsal view); 52, half of labium (ventral view). Abbreviations: ds 1 and ds 2, first and second dentisetae (52, holotype)." httpUri="https://zenodo.org/record/250190/files/figure.png" pageId="6" pageNumber="215">Fig. 52</figureCitation>
). Patella-tibial suture, being equally developed on all legs (that is characteristic for Anteropatellata), either has normal shape (in 
<taxonomicName id="EBA64D3694763B72FB1DFEC5FAC8A519" box="[1234,1338,333,356]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FB1DFEC5FAC8A519" box="[1234,1338,333,356]" italics="true" pageId="6" pageNumber="215">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694763B72FA43FEC5FEC6A5F5" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794763B72FA43FEC5FEC6A5F5" italics="true" pageId="6" pageNumber="215">I. starmuehlneri</emphasis>
</taxonomicName>
— 
<figureCitation id="B49D2A3094763B72FE9DFEE7FE5BA5F5" box="[338,425,367,392]" captionStart="FIGURES 18 – 22" captionStartId="4.[151,264,1610,1633]" captionTargetBox="[152,1436,197,1589]" captionTargetId="figure@4.[151,1436,197,1590]" captionTargetPageId="4" captionText="FIGURES 18 – 22. Indobaetis costai. 18, subimaginal exuviae of left half of mesonotum; 19, subimaginal exuviae of left subalar sclerite, postsubalar sclerite and lateropostnotal crest; 20, abdominal segments I – VI of male imago, spread on slide; 21, genitals of male imago (intersegmental ventral muscles taken away and unpaired sterno-styligeral muscle visible though integument); 22, part of larval fore leg." httpUri="https://zenodo.org/record/250185/files/figure.png" pageId="6" pageNumber="215">Fig. 22</figureCitation>
), or is smoothed and stretches along tibia (in 
<taxonomicName id="EBA64D3694763B72FC2CFEF9FB8DA5F5" box="[995,1151,367,392]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794763B72FC2CFEF9FB8DA5F5" box="[995,1151,367,392]" italics="true" pageId="6" pageNumber="215">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FB41FEE7FB4CA5F5" box="[1166,1214,367,392]" pageId="6" pageNumber="215" rank="species">sp.n.</taxonomicNameLabel>
— 
<figureCitation id="B49D2A3094763B72FB11FEE7FACBA5F5" box="[1246,1337,367,392]" captionStart="FIGURES 66 – 70" captionStartId="14.[151,264,1648,1671]" captionTargetBox="[151,1436,447,1623]" captionTargetId="figure@14.[151,1436,447,1624]" captionTargetPageId="14" captionText="FIGURES 66 – 70. Indobaetis microfolius sp. n. (holotype). 66 – 69, details of larval middle leg; 70, abdominal sterna and terga of male subimago, spread on slide." httpUri="https://zenodo.org/record/250193/files/figure.png" pageId="6" pageNumber="215">Fig. 67</figureCitation>
). Larval paraproct either has tongue-like denticulate prolongation of inner posterior edge (in 
<taxonomicName id="EBA64D3694763B72FB1BFE1DFAC9A5D1" box="[1236,1339,405,428]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FB1BFE1DFAC9A5D1" box="[1236,1339,405,428]" italics="true" pageId="6" pageNumber="215">I. costai</emphasis>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694763B72FA43FE1DFEC6A5AD" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794763B72FA43FE1DFEC6A5AD" italics="true" pageId="6" pageNumber="215">I. starmuehlneri</emphasis>
</taxonomicName>
— 
<figureCitation id="B49D2A3094763B72FE9EFE3FFE22A5AD" box="[337,464,439,464]" captionStart="FIGURES 36 – 39" captionStartId="9.[151,264,1558,1581]" captionTargetBox="[151,1436,195,1508]" captionTargetId="figure@9.[151,1436,195,1538]" captionTargetPageId="9" captionText="FIGURES 36 – 39. Posterior view on 10 th abdominal segment (cerci and paracercus taken away). 36, Nigrobaetis (Takobia) maxillaris; 37, undescribed species of Papuanatula from Baliem Valley (Papua province of Indonesia); 38, 39, two specimens of Indobaetis costai." httpUri="https://zenodo.org/record/250188/files/figure.png" pageId="6" pageNumber="215">Figs 38, 39</figureCitation>
), or with posterior margin non-expressed (in 
<taxonomicName id="EBA64D3694763B72FC04FE31FB92A5AD" box="[971,1120,439,464]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794763B72FC04FE31FB92A5AD" box="[971,1120,439,464]" italics="true" pageId="6" pageNumber="215">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FBA8FE3FFB6AA5AD" box="[1127,1176,439,464]" pageId="6" pageNumber="215" rank="species">sp.n.</taxonomicNameLabel>
— 
<figureCitation id="B49D2A3094763B72FB77FE3FFAFEA5AD" box="[1208,1292,439,464]" captionStart="FIGURES 71 – 75" captionStartId="16.[151,264,1726,1749]" captionTargetBox="[151,1436,195,1701]" captionTargetId="figure@16.[151,1436,195,1703]" captionTargetPageId="16" captionText="FIGURES 71 – 75. Indobaetis microfolius sp. n. 71, hypopharynx and superlingua; 72 – 75, abdominal segment X with paracercus and bases of cerci: 72, 74, 75, ventral view; 73, dorsal view." httpUri="https://zenodo.org/record/250194/files/figure.png" pageId="6" pageNumber="215">Fig. 72</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="64BC653E94763B72FF08FE54FCD5A641" pageId="6" pageNumber="215" type="distribution">
<paragraph id="2C1936B594763B72FF08FE54FD4AA588" blockId="6.[151,1437,151,824]" box="[199,696,476,501]" pageId="6" pageNumber="215">
<emphasis id="1ED2EAA794763B72FF08FE54FE92A588" bold="true" box="[199,352,476,501]" pageId="6" pageNumber="215">Distribution.</emphasis>
Southern 
<collectingCountry id="54B1762594763B72FE1CFE54FDFEA588" box="[467,524,476,501]" name="India" pageId="6" pageNumber="215">India</collectingCountry>
and 
<collectingCountry id="54B1762594763B72FD8CFE54FD47A588" box="[579,693,476,501]" name="Sri Lanka" pageId="6" pageNumber="215">Sri Lanka</collectingCountry>
.
</paragraph>
<paragraph id="2C1936B594763B72FF08FE77FCD5A641" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="215">
<emphasis id="1ED2EAA794763B72FF08FE77FE37A665" bold="true" box="[199,453,511,536]" pageId="6" pageNumber="215">Species composition.</emphasis>
3 species: 
<taxonomicName id="EBA64D3694763B72FD99FE77FB3FA665" authority="Muller-Liebenau &amp; Morihara 1982" authorityName="Muller-Liebenau &amp; Morihara" authorityYear="1982" box="[598,1229,511,536]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="costai">
<emphasis id="1ED2EAA794763B72FD99FE77FCECA665" box="[598,798,511,536]" italics="true" pageId="6" pageNumber="215">Indobaetis costai</emphasis>
<bibRefCitation id="48374B4494763B72FCE3FE77FB3FA665" author="Muller-Liebenau" box="[812,1229,511,536]" pageId="6" pageNumber="215" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="EBA64D3694763B72FB2FFD89FDEDA641" authority="Muller-Liebenau &amp; Morihara 1982" authorityName="Muller-Liebenau &amp; Morihara" authorityYear="1982" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="starmuehlneri">
<emphasis id="1ED2EAA794763B72FB2FFD89FA6EA665" box="[1248,1436,511,536]" italics="true" pageId="6" pageNumber="215">I. starmuehlneri</emphasis>
<bibRefCitation id="48374B4494763B72FF58FDACFDEDA641" author="Muller-Liebenau" box="[151,543,548,573]" pageId="6" pageNumber="215" refString="Muller-Liebenau, I. &amp; Morihara, D. K. (1982) Indobaetis: a new genus of Baetidae from Sri Lanka (Insecta: Ephemeroptera) with two new species. Gewasser und Abwasser, 68 / 69, 26 - 34." type="journal article" year="1982">Müller-Liebenau &amp; Morihara 1982</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="EBA64D3694763B72FD9AFDADFD18A640" box="[597,746,548,573]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="microfolius" status="sp. nov.">
<emphasis id="1ED2EAA794763B72FD9AFDADFD18A640" box="[597,746,548,573]" italics="true" pageId="6" pageNumber="215">I. microfolius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FD3DFDADFCD5A641" box="[754,807,549,572]" pageId="6" pageNumber="215" rank="species">sp.n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="64BC653E94763B72FF08FDCFFEB7A745" pageId="6" pageNumber="215" type="discussion">
<paragraph id="2C1936B594763B72FF08FDCFFEB7A745" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="215">
<emphasis id="1ED2EAA794763B72FF08FDCFFEBDA61D" bold="true" box="[199,335,583,608]" pageId="6" pageNumber="215">Comments.</emphasis>
Selva-kumar 
<emphasis id="1ED2EAA794763B72FE22FDC1FDD2A61D" box="[493,544,583,608]" italics="true" pageId="6" pageNumber="215">et al</emphasis>
. (2012) described 
<taxonomicName id="EBA64D3694763B72FD3BFDCFFBCCA61D" box="[756,1086,583,608]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="michaelohubbardi">
<emphasis id="1ED2EAA794763B72FD3BFDCFFBCCA61D" box="[756,1086,583,608]" italics="true" pageId="6" pageNumber="215">Indobaetis michaelohubbardi</emphasis>
</taxonomicName>
based on larvae from southern 
<collectingCountry id="54B1762594763B72FF58FDE4FF26A6F8" box="[151,212,620,645]" name="India" pageId="6" pageNumber="215">India</collectingCountry>
. Subsequent reexamination of the 
<typeStatus id="F31D881794763B72FDA5FDE5FD68A6F8" box="[618,666,621,645]" pageId="6" pageNumber="215">type</typeStatus>
specimens reveals that this species has hind protoptera and other characters of 
<taxonomicName id="EBA64D3694763B72FEF9FD19FE8EA6D5" box="[310,380,657,680]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="1ED2EAA794763B72FEF9FD19FE8EA6D5" box="[310,380,657,680]" italics="true" pageId="6" pageNumber="215">Baetis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FE47FD18FE4EA6DA" box="[392,444,656,679]" pageId="6" pageNumber="215" sensu="stricto">s.str.</taxonomicNameLabel>
(K.G. Sivaramakrishnan, personal communication). In the photo (Selva-kumar 
<emphasis id="1ED2EAA794763B72FA90FD19FA67A6D5" box="[1375,1429,655,680]" italics="true" pageId="6" pageNumber="215">et al</emphasis>
. 2012: 
<figureCitation id="B49D2A3094763B72FF12FD3CFEDFA6B1" box="[221,301,692,717]" captionStart="FIGURES 30 – 35" captionStartId="8.[151,264,1685,1708]" captionTargetBox="[151,1436,195,1661]" captionTargetId="figure@8.[151,1436,195,1662]" captionTargetPageId="8" captionText="FIGURES 30 – 35. Indobaetis costai, eggs and their enlarged micropilar areas." httpUri="https://zenodo.org/record/250187/files/figure.png" pageId="6" pageNumber="215">Fig. 34</figureCitation>
) three maxillary dentisetae are visible, that allows to exclude this species from 
<taxonomicName id="EBA64D3694763B72FB6DFD3CFAEAA6B0" box="[1186,1304,692,717]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794763B72FB6DFD3CFAEAA6B0" box="[1186,1304,692,717]" italics="true" pageId="6" pageNumber="215">Indobaetis</emphasis>
</taxonomicName>
. According to the photos (Selva-kumar 
<emphasis id="1ED2EAA794763B72FE01FD51FE0CA68D" box="[462,510,727,752]" italics="true" pageId="6" pageNumber="215">et al</emphasis>
. 2012: 
<figureCitation id="B49D2A3094763B72FD9AFD5FFD21A68D" box="[597,723,727,752]" captionStart-0="FIGURES 23 – 29" captionStart-1="FIGURES 30 – 35" captionStart-2="FIGURES 36 – 39" captionStartId-0="7.[151,264,1729,1752]" captionStartId-1="8.[151,264,1685,1708]" captionStartId-2="9.[151,264,1558,1581]" captionTargetBox-0="[158,1424,204,1705]" captionTargetBox-1="[151,1436,195,1661]" captionTargetBox-2="[151,1436,195,1508]" captionTargetId-0="figure@7.[151,1436,195,1705]" captionTargetId-1="figure@8.[151,1436,195,1662]" captionTargetId-2="figure@9.[151,1436,195,1538]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 23 – 29. Indobaetis costai. 23 – 25, male imago: 23, wing; 24, genitals (ventral view, muscles of middle and left parts shown by interrupted lines); 25, head and dorsal outline of pronotum and anteronotal protuberance, lateral view. 26, head of female imago, dorsal view. 27, subimaginal exuviae of right unistyliger and gonostylus; 28, subimaginal genitals developing under larval cuticle; 29, the same at earlier phase. Abbreviations: gs 1 – gs 3, 1 st – 3 rd segments of gonostylus; m. s, median sternostyligeral muscle; m. IX – X, intersegmental ventral muscles going from base of 9 th abdominal segment to base of 10 th abdominal segment; us, unistyliger." captionText-1="FIGURES 30 – 35. Indobaetis costai, eggs and their enlarged micropilar areas." captionText-2="FIGURES 36 – 39. Posterior view on 10 th abdominal segment (cerci and paracercus taken away). 36, Nigrobaetis (Takobia) maxillaris; 37, undescribed species of Papuanatula from Baliem Valley (Papua province of Indonesia); 38, 39, two specimens of Indobaetis costai." httpUri-0="https://zenodo.org/record/250186/files/figure.png" httpUri-1="https://zenodo.org/record/250187/files/figure.png" httpUri-2="https://zenodo.org/record/250188/files/figure.png" pageId="6" pageNumber="215">Figs 29–38</figureCitation>
) this species belongs to subgenus 
<taxonomicName id="EBA64D3694763B72FB9BFD5FFA84A68D" authority="Kang &amp; Yang" authorityName="Kang &amp; Yang" box="[1108,1398,727,752]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="1ED2EAA794763B72FB9BFD5FFB23A68D" box="[1108,1233,727,752]" italics="true" pageId="6" pageNumber="215">Tenuibaetis</emphasis>
Kang &amp; Yang
</taxonomicName>
(in 
<bibRefCitation id="48374B4494763B72FF58FD75FEACA769" author="Kang" box="[151,350,764,789]" pageId="6" pageNumber="215" refString="Kang, S. C., Chang, H. C. &amp; Yang, C. T. (1994) A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum, 47 (2), 9 - 44." type="journal article" year="1994">
Kang 
<emphasis id="1ED2EAA794763B72FF13FD75FEFCA768" box="[220,270,764,789]" italics="true" pageId="6" pageNumber="215">et al</emphasis>
.) 1994
</bibRefCitation>
of the genus 
<taxonomicName id="EBA64D3694763B72FE35FD75FDB2A769" box="[506,576,765,788]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="1ED2EAA794763B72FE35FD75FDB2A769" box="[506,576,765,788]" italics="true" pageId="6" pageNumber="215">Baetis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05E157DC94763B72FD87FD75FD89A769" box="[584,635,765,788]" pageId="6" pageNumber="215" sensu="stricto">s.str.</taxonomicNameLabel>
So, the valid binomen is 
<taxonomicName id="EBA64D3694763B72FC6FFD75FB48A768" box="[928,1210,764,789]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="michaelohubbardi">
<emphasis id="1ED2EAA794763B72FC6FFD75FB48A768" box="[928,1210,764,789]" italics="true" pageId="6" pageNumber="215">Baetis michaelohubbardi</emphasis>
</taxonomicName>
(Selva-kumar 
<emphasis id="1ED2EAA794763B72FAACFD75FA66A768" box="[1379,1428,764,789]" italics="true" pageId="6" pageNumber="215">et al</emphasis>
. 2012), 
<emphasis id="1ED2EAA794763B72FF29FC97FEB7A745" bold="true" box="[230,325,799,824]" pageId="6" pageNumber="215">comb.n.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>