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<document id="DC3A4FE54731F07C25B210C2C204CE1C" ID-CLB-Dataset="27726" ID-DOI="10.11646/zootaxa.4539.1.1" ID-GBIF-Dataset="e7c7a589-9f74-4546-9a7b-c3192b9659e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615911" ID-ZooBank="2C72727B-79C5-407F-BD92-B12F98196800" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553789058718" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2018" docId="193787A0FFDBFFA5F4CBFB27459ECAB8" docLanguage="en" docName="zootaxa.4539.1.1.pdf" docOrigin="Zootaxa 4539 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mediaster ornatus Fisher 1906" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="E50EFFD8FFE2FF9FF45CFFC04464CA17" masterDocTitle="New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean" masterLastPageNumber="116" masterPageNumber="1" pageNumber="58" updateTime="1698715174966" updateUser="ExternalLinkService">
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<mods:title id="F667C6D05EC1BB215A06A264BC162F7B">New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean</mods:title>
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<subSubSection id="D984653DFFDBFFA6F4CBFB2740CBCF74" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="912136B6FFDBFFA6F4CBFB274669CF15" blockId="57.[151,525,1255,1282]" box="[151,525,1255,1282]" pageId="57" pageNumber="58">
<heading id="CA6981DAFFDBFFA6F4CBFB274669CF15" bold="true" box="[151,525,1255,1282]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<taxonomicName id="569E4D35FFDBFFA6F4CBFB274669CF15" ID-CoL="3YNGK" authority="Fisher 1906" authorityName="Fisher" authorityYear="1906" box="[151,525,1255,1282]" class="Asteroidea" family="Goniasteridae" genus="Mediaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="57" pageNumber="58" phylum="Echinodermata" rank="species" species="ornatus">
<emphasis id="A3EAEAA4FFDBFFA6F4CBFB274669CF15" bold="true" box="[151,525,1255,1282]" pageId="57" pageNumber="58">
<emphasis id="A3EAEAA4FFDBFFA6F4CBFB274517CF16" bold="true" box="[151,371,1255,1281]" italics="true" pageId="57" pageNumber="58">Mediaster ornatus</emphasis>
<bibRefCitation id="F50F4B47FFDBFFA6F526FB284669CF15" author="Fisher, W. K." box="[378,525,1256,1282]" pageId="57" pageNumber="58" pagination="987 - 1130" refId="ref70129" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher 1906</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="912136B6FFDBFFA6F4CBFAEE40CBCF74" blockId="57.[151,1436,1326,1379]" pageId="57" pageNumber="58">
<bibRefCitation id="F50F4B47FFDBFFA6F4CBFAEE4577CF53" author="Fisher, W. K." box="[151,275,1326,1348]" pageId="57" pageNumber="58" pagination="987 - 1130" refId="ref70129" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher 1906</bibRefCitation>
: 1046;
<bibRefCitation id="F50F4B47FFDBFFA6F502FAEE4589CF53" author="Koehler, R." box="[350,493,1326,1348]" pageId="57" pageNumber="58" pagination="115 - 131" refId="ref71489" refString="Koehler, R. (1909) An account of the deep-sea Asteroidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, Part 5, 115 - 131. https: // doi. org / 10.5962 / bhl. title. 11614" type="book chapter" year="1909">Koehler 1909</bibRefCitation>
: 78;
<bibRefCitation id="F50F4B47FFDBFFA6F642FAEE46FFCF53" author="Fisher, W. K." box="[542,667,1326,1348]" pageId="57" pageNumber="58" pagination="1 - 547" refId="ref70312" refString="Fisher, W. K. (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 3 (100), 1 - 547, 156 pls. https: // doi. org / 10.1126 / science. 50.1293.348" type="book chapter" year="1919">Fisher 1919</bibRefCitation>
: 256;
<bibRefCitation id="F50F4B47FFDBFFA6F685FAEE4718CF53" author="Doderlein, L." box="[729,892,1326,1348]" pageId="57" pageNumber="58" pagination="49 - 69" refId="ref69892" refString="Doderlein, L. (1924) Die Asteriden der Siboga-Expedition. II. Pentagonasteridae. Siboga-Expedition, 46 (2), 49 - 69, pls. 14 - 19. https: // doi. org / 10.5962 / bhl. title. 11319" type="journal article" year="1924">Döderlein 1924</bibRefCitation>
: 52;
<bibRefCitation id="F50F4B47FFDBFFA6F7F1FAEF40F5CF53" author="Jangoux, M. &amp; Aziz, A." box="[941,1169,1326,1348]" pageId="57" pageNumber="58" pagination="631 - 650" refId="ref71303" refString="Jangoux, M. &amp; Aziz, A. (1988) Les asterides (Echinodermata) recoltes autour de l'ile de la Reunion par le N. O. Marion- Dufresne en 1982. Bulletin Museum national d'Historie du naturelle, 4 (10), 631 - 650." type="journal article" year="1988">Jangoux &amp; Aziz 1988</bibRefCitation>
: 633, 642;
<bibRefCitation id="F50F4B47FFDBFFA6F15DFAEE4499CF74" author="Chave, E. H. &amp; Jones, A. T." pageId="57" pageNumber="58" pagination="781 - 803" refId="ref68790" refString="Chave, E. H. &amp; Jones, A. T. (1991) Deep-water megafauna of the Kohala and Haleakala slopes, Alenuihana Channel, Hawaii. Deep-Sea Research, 38 (7), 781 - 803. https: // doi. org / 10.1016 / 0198 - 0149 (91) 90019 - C" type="journal article" year="1991">Chave &amp; Jones 1991</bibRefCitation>
: 794, 796;
<bibRefCitation id="F50F4B47FFDBFFA6F531FA8E4679CF74" author="Clark, A. M." box="[365,541,1357,1379]" pageId="57" pageNumber="58" pagination="187 - 366" refId="ref69077" refString="Clark, A. M. (1993) An index of names of recent Asteroidea-Part 2: Valvatida. E chinoderm Studies, 4, 187 - 366." type="journal article" year="1993">A.M. Clark 1993</bibRefCitation>
: 263 (checklist);
<bibRefCitation id="F50F4B47FFDBFFA6F697FA8D47A3CF74" author="Chave, E. H. &amp; Malahoff, A." box="[715,967,1357,1379]" pageId="57" pageNumber="58" refId="ref68850" refString="Chave, E. H. &amp; Malahoff, A. (1998) In Deeper Waters: photographic studies of Hawaiian deep-sea habitats and life forms. University of Hawaii Press, Honolulu, 127 pp." type="book" year="1998">Chave &amp; Malahoff 1998</bibRefCitation>
: 86;
<bibRefCitation id="F50F4B47FFDBFFA6F7ABFA8D401CCF74" author="Paulay, G." box="[1015,1144,1357,1379]" pageId="57" pageNumber="58" pagination="563 - 583" refId="ref73501" refString="Paulay, G. (2003) The Asteroidea, Echinoidea, and Holothuroidea (Echinodermata) of the Mariana Islands. Micronesica, 3536, 563 - 583." type="journal article" year="2003">Paulay 2003</bibRefCitation>
: 572.
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFDBFFA6F4CBFA4F46F4CCB2" pageId="57" pageNumber="58" type="diagnosis">
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<emphasis id="A3EAEAA4FFDBFFA6F4CBFA4F4575CFBF" bold="true" box="[151,273,1423,1448]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
Body stellate (R/r=2.42.5), well developed tabulae, round in shape. Central surface with one to nine small blunt, cylindrical granules in ordered series, peripheral granules 15 to 30, round to cylindrical in shape, widely and evenly spaced. Superomarginals covered by widely distributed bullet-shaped, blunt granules. Actinal plate surface with four to 20, mostly 1015 peripheral angular granules, triangular to quadrate in cross-section, two to eight located on plate central surface, also angular in cross-section, blunt tipped. Furrow spines variable, ranging from seven on the Hawaiian
<typeStatus id="4E258814FFDBFFA6F580F984465ACC4A" box="[476,574,1604,1629]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
(R=
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, but listed as eight spines by Macan) to six to eight at R=
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.
<bibRefCitation id="F50F4B47FFDBFFA6F4CBF9A8455CCC97" author="Macan, T. T." box="[151,312,1640,1664]" pageId="57" pageNumber="58" pagination="324 - 432" refId="ref71993" refString="Macan, T. T. (1938) Asteroidea. British Museum of Natural History, John Murray Expedition 1933 - 34, Scientific Reports, IV (9), 324 - 432." type="journal article" year="1938">Macan (1938)</bibRefCitation>
summarizes variation ranging from five to seven in the Philippine specimens to those listed by
<bibRefCitation id="F50F4B47FFDBFFA6F4CBF94C453CCCB3" author="Doderlein, L." box="[151,344,1676,1701]" pageId="57" pageNumber="58" pagination="49 - 69" refId="ref69892" refString="Doderlein, L. (1924) Die Asteriden der Siboga-Expedition. II. Pentagonasteridae. Siboga-Expedition, 46 (2), 49 - 69, pls. 14 - 19. https: // doi. org / 10.5962 / bhl. title. 11319" type="journal article" year="1924">Döderlein (1924)</bibRefCitation>
as showing eight to eleven.
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFDBFFA6F49BF96F454ACDFF" pageId="57" pageNumber="58" type="discussion">
<paragraph id="912136B6FFDBFFA6F49BF96F4029CDB7" blockId="57.[151,1437,1423,2025]" pageId="57" pageNumber="58">
<emphasis id="A3EAEAA4FFDBFFA6F49BF96F4529CCDF" bold="true" box="[199,333,1711,1736]" pageId="57" pageNumber="58">Comments.</emphasis>
Although no new records of this species are included herein, this species is included for comparisons with the other
<taxonomicName id="569E4D35FFDBFFA6F560F91445CFCCFA" authorityName="Stimpson" authorityYear="1857" box="[316,427,1748,1773]" class="Asteroidea" family="Goniasteridae" genus="Mediaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="57" pageNumber="58" phylum="Echinodermata" rank="genus">
<emphasis id="A3EAEAA4FFDBFFA6F560F91445CFCCFA" box="[316,427,1748,1773]" italics="true" pageId="57" pageNumber="58">Mediaster</emphasis>
</taxonomicName>
species. This is the most widely occurring species of
<taxonomicName id="569E4D35FFDBFFA6F04CF914401ACCFA" authorityName="Stimpson" authorityYear="1857" box="[1040,1150,1748,1773]" class="Asteroidea" family="Goniasteridae" genus="Mediaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="57" pageNumber="58" phylum="Echinodermata" rank="genus">
<emphasis id="A3EAEAA4FFDBFFA6F04CF914401ACCFA" box="[1040,1150,1748,1773]" italics="true" pageId="57" pageNumber="58">Mediaster</emphasis>
</taxonomicName>
, and includes individuals identified from deep-sea settings in the Pacific and Indian Oceans (
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).
<bibRefCitation id="F50F4B47FFDBFFA6F053F93840CDCD07" author="Macan, T. T." box="[1039,1193,1784,1808]" pageId="57" pageNumber="58" pagination="324 - 432" refId="ref71993" refString="Macan, T. T. (1938) Asteroidea. British Museum of Natural History, John Murray Expedition 1933 - 34, Scientific Reports, IV (9), 324 - 432." type="journal article" year="1938">Macan (1938)</bibRefCitation>
summarized variation from different reported localities including Hawaii, the
<collectingCountry id="E9897626FFDBFFA6F759F8DC47E4CD22" box="[773,896,1820,1845]" name="Philippines" pageId="57" pageNumber="58">Philippines</collectingCountry>
and
<collectingCountry id="E9897626FFDBFFA6F7E5F8DC4043CD22" box="[953,1063,1820,1845]" name="Indonesia" pageId="57" pageNumber="58">Indonesia</collectingCountry>
. Later reports include those from Reunion Island (
<bibRefCitation id="F50F4B47FFDBFFA6F50EF880462ACD4F" author="Jangoux, M. &amp; Aziz, A." box="[338,590,1855,1880]" pageId="57" pageNumber="58" pagination="631 - 650" refId="ref71303" refString="Jangoux, M. &amp; Aziz, A. (1988) Les asterides (Echinodermata) recoltes autour de l'ile de la Reunion par le N. O. Marion- Dufresne en 1982. Bulletin Museum national d'Historie du naturelle, 4 (10), 631 - 650." type="journal article" year="1988">Jangoux &amp; Aziz 1988</bibRefCitation>
) and the
<collectingCountry id="E9897626FFDBFFA6F6E2F8FF4714CD4F" box="[702,880,1855,1880]" name="Northern Mariana Islands" pageId="57" pageNumber="58">Mariana Islands</collectingCountry>
(
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).
<taxonomicName id="569E4D35FFDBFFA6F075F8814015CD4F" box="[1065,1137,1857,1880]" class="Asteroidea" family="Goniasteridae" genus="Iconaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="57" pageNumber="58" phylum="Echinodermata" rank="species" species="situ">
<emphasis id="A3EAEAA4FFDBFFA6F075F8814015CD4F" box="[1065,1137,1857,1880]" italics="true" pageId="57" pageNumber="58">In situ</emphasis>
</taxonomicName>
observations made by the Hawaiian Undersea Research Laboratory (
<bibRefCitation id="F50F4B47FFDBFFA6F637F8A444BCCDB7" author="Chave, E. H. &amp; Malahoff, A." pageId="57" pageNumber="58" refId="ref68850" refString="Chave, E. H. &amp; Malahoff, A. (1998) In Deeper Waters: photographic studies of Hawaiian deep-sea habitats and life forms. University of Hawaii Press, Honolulu, 127 pp." type="book" year="1998">Chave &amp; Malahoff, 1998; Mah, personal observation, HURL unpublished video</bibRefCitation>
), show this species is orange in color and was present on a coarse, sandy bottom.
</paragraph>
<paragraph id="912136B6FFDBFFA6F49BF86C454ACDFF" blockId="57.[151,1437,1423,2025]" pageId="57" pageNumber="58">
<emphasis id="A3EAEAA4FFDBFFA6F49BF86C4538CDD2" bold="true" box="[199,348,1964,1989]" pageId="57" pageNumber="58">Occurrence.</emphasis>
Hawaiian Islands,
<collectingCountry id="E9897626FFDBFFA6F615F86C46F6CDD3" box="[585,658,1964,1988]" name="Guam" pageId="57" pageNumber="58">Guam</collectingCountry>
,
<collectingCountry id="E9897626FFDBFFA6F6FFF86C4743CDD2" box="[675,807,1964,1989]" name="Philippines" pageId="57" pageNumber="58">Philippines</collectingCountry>
,
<collectingCountry id="E9897626FFDBFFA6F764F86C47CFCDD2" box="[824,939,1964,1989]" name="Indonesia" pageId="57" pageNumber="58">Indonesia</collectingCountry>
,
<collectingCountry id="E9897626FFDBFFA6F7E0F86C404ECDD2" box="[956,1066,1964,1989]" name="Maldives" pageId="57" pageNumber="58">Maldives</collectingCountry>
, Arabian Sea, Reunion Island.
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.
</paragraph>
</subSubSection>
<subSubSection id="D984653DFFD8FFA5F49BFF57459ECAB8" box="[199,506,151,176]" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="912136B6FFD8FFA5F49BFF57459ECAB8" blockId="58.[199,506,151,176]" box="[199,506,151,176]" pageId="58" pageNumber="59">
<emphasis id="A3EAEAA4FFD8FFA5F49BFF5745D2CAA7" bold="true" box="[199,438,151,176]" pageId="58" pageNumber="59">Material Examined.</emphasis>
None
</paragraph>
</subSubSection>
</treatment>
</document>