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<document id="9E7D658DDF3B895BD54A3965417DE328" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCACD143F10B83FE118D6F05E4" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Andrena perforata WOOD, 2024, spec. nov." docType="treatment" docVersion="2" lastPageNumber="139" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="137" updateTime="1722787353532" updateUser="ExternalLinkService">
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<mods:title id="D2775056381D77A8C1F54B9702364E6A">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="371E17CD68CAB88BDC7B3BA4CE975AEF">WOOD, T. J.</mods:namePart>
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<mods:title id="03429D5C1F7774B2FEE5B9D4DA2B2009">Zootaxa</mods:title>
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<mods:date id="525CBD73E884673187C5980B1C2ED903">2024</mods:date>
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<mods:number id="A7174B8BB87195D4D357C7814DB58755">2024-07-22</mods:number>
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<mods:classification id="4640E7CB7D6622CEA2D08A17DBA79489">journal article</mods:classification>
<mods:identifier id="7FE93A38974DB06129F804C6FA0D545D" type="DOI">10.11646/zootaxa.5483.1.1</mods:identifier>
<mods:identifier id="370111C38472679198BA753771A1C00B" type="ISSN">1175-5334</mods:identifier>
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<treatment id="612B87FCACD143F10B83FE118D6F05E4" ID-DOI="http://doi.org/10.5281/zenodo.13209523" ID-Zenodo-Dep="13209523" LSID="urn:lsid:plazi:treatment:612B87FCACD143F10B83FE118D6F05E4" httpUri="http://treatment.plazi.org/id/612B87FCACD143F10B83FE118D6F05E4" lastPageId="138" lastPageNumber="139" pageId="136" pageNumber="137">
<subSubSection id="A1986561ACD143F30B83FE118CD307B4" box="[151,694,474,501]" pageId="136" pageNumber="137" type="nomenclature">
<paragraph id="E93D36EAACD143F30B83FE118CD307B4" blockId="136.[151,694,474,537]" box="[151,694,474,501]" pageId="136" pageNumber="137">
<heading id="B2758186ACD143F30B83FE118CD307B4" bold="true" box="[151,694,474,501]" fontSize="11" level="2" pageId="136" pageNumber="137" reason="4">
<emphasis id="DBF6EAF8ACD143F30B83FE118CD307B4" bold="true" box="[151,694,474,501]" pageId="136" pageNumber="137">
<taxonomicName id="2E824D69ACD143F30B83FE118C5207B5" authorityName="WOOD" authorityYear="2024" box="[151,567,474,501]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="perforata" status="sp. nov.">
<emphasis id="DBF6EAF8ACD143F30B83FE118F6407B4" bold="true" box="[151,257,474,501]" italics="true" pageId="136" pageNumber="137">Andrena</emphasis>
(
<emphasis id="DBF6EAF8ACD143F30A06FE108FD307B4" bold="true" box="[274,438,474,501]" italics="true" pageId="136" pageNumber="137">incertae sedis</emphasis>
)
<emphasis id="DBF6EAF8ACD143F30AD3FE118C5207B5" bold="true" box="[455,567,474,500]" italics="true" pageId="136" pageNumber="137">perforata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783ACD143F3092BFE108CD307B4" box="[575,694,475,501]" pageId="136" pageNumber="137" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD143F30B83FE348FD50458" box="[151,432,511,537]" pageId="136" pageNumber="137" type="description">
<paragraph id="E93D36EAACD143F30B83FE348FD50458" blockId="136.[151,694,474,537]" box="[151,432,511,537]" pageId="136" pageNumber="137">
<figureCitation id="71B92A6FACD143F30B83FE348FCE0458" box="[151,427,511,537]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figures 82A–C; E; G–H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD143F30B83FD8D8D870422" box="[151,994,582,611]" pageId="136" pageNumber="137" type="materials_examined">
<paragraph id="E93D36EAACD143F30B83FD8D8D870422" blockId="136.[151,1437,582,2013]" box="[151,994,582,611]" pageId="136" pageNumber="137">
<materialsCitation id="59EA3CB7ACD143F30B83FD8D8D870422" box="[151,994,582,611]" collectingDate="1971-06-01" collectorName="K. Warncke" country="Turkey" location="Ankara" pageId="136" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="Ankara" typeStatus="holotype">
<emphasis id="DBF6EAF8ACD143F30B83FD8D8FA10420" bold="true" box="[151,452,582,609]" pageId="136" pageNumber="137">
<typeStatus id="36398848ACD143F30B83FD8D8F5F0421" box="[151,314,582,608]" pageId="136" pageNumber="137" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AACD143F30A52FD8C8FA50420" box="[326,448,583,609]" name="Turkey" pageId="136" pageNumber="137">TURKEY</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808ACD143F30ADEFD8C8C7A0420" box="[458,543,583,609]" country="Turkey" name="Ankara" pageId="136" pageNumber="137">Ankara</collectingRegion>
,
<date id="9D3C102AACD143F3093EFD8C8CF30420" box="[554,662,583,609]" pageId="136" pageNumber="137" value="1971-06-01">
<collectingDate id="8D78E9C2ACD143F3093EFD8C8CF30420" box="[554,662,583,609]" pageId="136" pageNumber="137" value="1971-06-01">1.vi.1971</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACD143F309B6FD8C8CD20420" box="[674,695,583,609]" count="1" pageId="136" pageNumber="137" type="female">♀</specimenCount>
, leg.
<collectorName id="4477533CACD143F309E1FD8C8D180420" box="[757,893,583,609]" pageId="136" pageNumber="137">K. Warncke</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD143F30BD3FDA18FEF006C" pageId="136" pageNumber="137" type="description">
<paragraph id="E93D36EAACD143F30BD3FDA18C5305AC" blockId="136.[151,1437,582,2013]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3FDA18FA304C4" bold="true" box="[199,454,618,645]" pageId="136" pageNumber="137">Description: Female:</emphasis>
Body length:
<quantity id="2E7A9B0FACD143F30965FDA08CD304C4" box="[625,694,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="136" pageNumber="137" unit="mm" value="6.0">6 mm</quantity>
(
<figureCitation id="71B92A6FACD143F309DCFDA08D3604C4" box="[712,851,619,645]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82A</figureCitation>
).
<emphasis id="DBF6EAF8ACD143F3087DFDA08DCA04C4" box="[873,943,619,645]" italics="true" pageId="136" pageNumber="137">Head:</emphasis>
Dark, 1.25 times wider than long (
<figureCitation id="71B92A6FACD143F30E45FDA08EAC04E8" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82B</figureCitation>
). Clypeus domed, irregularly punctate, punctures separated by 0.5–1.5 puncture diameters, punctures deep, interspaces unevenly raised, poorly defined impunctate longitudinal mid-line present; underlying surface weakly shagreened and more or less shining. Process of labrum narrowly trapezoidal, slightly longer than wide. Gena subequal to width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Frons polished and shining, deeply and regularly punctate, punctures separated by 0.5–1 puncture diameters (
<figureCitation id="71B92A6FACD143F30FA7FCD48B5D0578" box="[1203,1336,799,825]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82C</figureCitation>
). Foveae dorsally narrow, dorsally occupying ⅓ of space between compound eye and lateral ocellus, equalling width of flagellum; foveae narrowing ventrally, at level of antennal insertions separated from inner margin of compound eye by own diameter, this space shining; foveae filled with light brown hairs. Face, gena, vertex, and scape with short whitish-brown hairs, none equalling length of scape. Antennae dark basally, A4–12 ventrally lightened by presence of orange scales, A3 equalling A4+5.
</paragraph>
<paragraph id="E93D36EAACD143F30BD3FC3C8A620314" blockId="136.[151,1437,582,2013]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3FC3C8F2D0251" box="[199,328,1015,1040]" italics="true" pageId="136" pageNumber="137">
<taxonomicName id="2E824D69ACD143F30BD3FC3C8F270251" box="[199,322,1015,1040]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Scutum and scutellum regularly but shallowly punctate, punctures separated by 1–2 puncture diameters, underlying surface evenly shagreened, weakly shining (
<figureCitation id="71B92A6FACD143F30868FBD08D9A0274" box="[892,1023,1051,1077]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82E</figureCitation>
). Pronotum rounded. Mesepisternum with coarse granular shagreen, anteriorly overlain by weakly raised rugosity. Dorsolateral parts of propodeum with extremely fine shagreen, contrasting propodeal triangle which is covered with coarser granular shagreen, basally with obscure, short, and weakly raised rugae, surface weakly shining (
<figureCitation id="71B92A6FACD143F30F03FB4C8AFA02E0" box="[1047,1183,1159,1185]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82G</figureCitation>
). Mesepisternum with thickly plumose white hairs, hairs becoming shorter on scutum and scutellum, sub-squamous. Propodeal corbicula incomplete, dorsal fringe composed of whitish plumose hairs, internal surface with scattered long whitish hairs. Legs dark, apical tarsal segments lightened brownish, pubescence light brown. Flocculus incomplete, composed of white plumose hairs; femoral and tibial scopae composed of white simple hairs. Hind tarsal claws without inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal.
</paragraph>
<paragraph id="E93D36EAACD143F30BD3FA948A240048" blockId="136.[151,1437,582,2013]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3FA948F200339" box="[199,325,1375,1400]" italics="true" pageId="136" pageNumber="137">Metasoma:</emphasis>
Tergal discs dark, marginal areas broadly lightened hyaline yellow to obscure reddish brown (
<figureCitation id="71B92A6FACD143F30E45FA948EAE03DC" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="136" pageNumber="137">Figure 82H</figureCitation>
). Tergal discs strongly microreticulate, dull more or less impunctate, sculpture becoming weaker on margins, weakly shining. T2–4 laterally with weak, widely separated apical hairbands of whitish hairs. Apical fringe of T5 composed of long whitish hairs, overlying orange hairs flanking pygidial plate. Pygidial plate subtriangular, apically rounded, surface flat and featureless, surface finely shagreened and weakly shining.
</paragraph>
<paragraph id="E93D36EAACD143F30BD3F9D88FEF006C" blockId="136.[151,1437,582,2013]" box="[199,394,1555,1581]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3F9D88F68006C" bold="true" box="[199,269,1555,1581]" pageId="136" pageNumber="137">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD143F10BD3F9FD8B320578" lastPageId="138" lastPageNumber="139" pageId="136" pageNumber="137" type="diagnosis">
<paragraph id="E93D36EAACD143F30BD3F9FD8B3B019C" blockId="136.[151,1437,582,2013]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3F9FD8F5A0011" bold="true" box="[199,319,1590,1616]" pageId="136" pageNumber="137">Diagnosis.</emphasis>
<taxonomicName id="2E824D69ACD143F30A52F9FC8C770010" authorityName="WOOD" authorityYear="2024" box="[326,530,1591,1617]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD143F30A52F9FC8C770010" box="[326,530,1591,1617]" italics="true" pageId="136" pageNumber="137">Andrena perforata</emphasis>
</taxonomicName>
is difficult to place in a subgenus. It is morphologically closest to
<taxonomicName id="2E824D69ACD143F30FF8F9FC8FFB0034" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="konyella" subGenus="Aciandrena">
<emphasis id="DBF6EAF8ACD143F30FF8F9FC8B610010" box="[1260,1284,1591,1617]" italics="true" pageId="136" pageNumber="137">A.</emphasis>
(
<emphasis id="DBF6EAF8ACD143F30E00F9FC8BF60010" box="[1300,1427,1591,1617]" italics="true" pageId="136" pageNumber="137">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8ACD143F30B83F9908E910034" box="[151,244,1627,1653]" italics="true" pageId="136" pageNumber="137">konyella</emphasis>
Warncke, 1975
</taxonomicName>
(
<collectingCountry id="9195767AACD143F30ABFF9908F9E0034" box="[427,507,1627,1653]" name="Turkey" pageId="136" pageNumber="137">Turkey</collectingCountry>
only) due to the small body size (
<quantity id="2E7A9B0FACD143F3087DF9918DAD0034" box="[873,968,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="136" pageNumber="137" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
), black integument, ventrally narrow facial foveae, shagreened impunctate terga, propodeal triangle which is covered with fine to moderately coarse granular shagreen, with minimal raised rugae basally, weakly antefurcal nervulus, and long ocelloccipital distance (
<figureCitation id="71B92A6FACD143F30E5CF9688E8F00A0" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="136" pageNumber="137">Figures 81C–D</figureCitation>
;
<figureCitation id="71B92A6FACD143F30BE2F90D8F5100A0" box="[246,308,1734,1761]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="6.[151,250,1218,1242]" captionStartId-1="7.[151,250,1149,1173]" captionTargetBox-0="[151,1436,237,1194]" captionTargetBox-1="[151,1433,183,1123]" captionTargetId-0="figure-45@6.[151,1436,237,1194]" captionTargetId-1="figure-16@7.[151,1435,181,1125]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." captionText-1="FIGURE 2. Phylogenetic tree (maximum likelihood) of Andrena subgenus Avandrena Warncke, 1968 basedon the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi-0="http://doi.org/10.5281/zenodo.13209309" figureDoi-1="http://doi.org/10.5281/zenodo.13209311" httpUri-0="https://zenodo.org/record/13209309/files/figure.png" httpUri-1="https://zenodo.org/record/13209311/files/figure.png" pageId="136" pageNumber="137">1.5–2</figureCitation>
times the diameter of a lateral ocellus). However,
<taxonomicName id="2E824D69ACD143F30873F90C8D8600A0" authorityName="Warncke" authorityYear="1975" box="[871,995,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD143F30873F90C8D8600A0" box="[871,995,1735,1761]" italics="true" pageId="136" pageNumber="137">A. konyella</emphasis>
</taxonomicName>
is not a typical
<taxonomicName id="2E824D69ACD143F30F82F90C8B7000A0" authorityName="Warncke" authorityYear="1968" box="[1174,1301,1735,1761]" pageId="136" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8ACD143F30F82F90C8B7000A0" box="[1174,1301,1735,1761]" italics="true" pageId="136" pageNumber="137">Aciandrena</emphasis>
</taxonomicName>
. Though no genetic data are currently available, in the female sex the hind tarsal claws of
<taxonomicName id="2E824D69ACD143F308FAF9208A0C0144" authorityName="Warncke" authorityYear="1975" box="[1006,1129,1771,1797]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD143F308FAF9208A0C0144" box="[1006,1129,1771,1797]" italics="true" pageId="136" pageNumber="137">A. konyella</emphasis>
</taxonomicName>
are bidentate, with an inner subapical tooth. In contrast,
<taxonomicName id="2E824D69ACD143F30AC3F8C48C3A0168" authorityName="WOOD" authorityYear="2024" box="[471,607,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD143F30AC3F8C48C3A0168" box="[471,607,1807,1833]" italics="true" pageId="136" pageNumber="137">A. perforata</emphasis>
</taxonomicName>
does not display an inner subapical tooth on the hind tarsal claws, which would be more consistent with
<taxonomicName id="2E824D69ACD143F30914F8F88C1A010C" authorityName="Warncke" authorityYear="1968" box="[512,639,1843,1869]" pageId="136" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8ACD143F30914F8F88C1A010C" box="[512,639,1843,1869]" italics="true" pageId="136" pageNumber="137">Aciandrena</emphasis>
</taxonomicName>
, but the species is best placed as
<emphasis id="DBF6EAF8ACD143F30F1CF8F88AC6010C" box="[1032,1187,1843,1869]" italics="true" pageId="136" pageNumber="137">incertae sedis</emphasis>
for now until genetic data are available, and species-level diagnosis should be made against
<taxonomicName id="2E824D69ACD143F308B0F89C8A450130" authorityName="Warncke" authorityYear="1975" box="[932,1056,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD143F308B0F89C8A450130" box="[932,1056,1879,1905]" italics="true" pageId="136" pageNumber="137">A. konyella</emphasis>
</taxonomicName>
. Overall,
<taxonomicName id="2E824D69ACD143F30F9BF89C8B730130" authorityName="WOOD" authorityYear="2024" box="[1167,1302,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD143F30F9BF89C8B730130" box="[1167,1302,1879,1905]" italics="true" pageId="136" pageNumber="137">A. perforata</emphasis>
</taxonomicName>
only differs meaningfully from a typical
<taxonomicName id="2E824D69ACD143F30AC7F8B08C3701D4" authorityName="Warncke" authorityYear="1968" box="[467,594,1915,1941]" pageId="136" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8ACD143F30AC7F8B08C3701D4" box="[467,594,1915,1941]" italics="true" pageId="136" pageNumber="137">Aciandrena</emphasis>
</taxonomicName>
species by the long ocelloccipital distance (usually &lt;1 times the diameter of a lateral ocellus) and the shiny and clearly punctate frons (usually sculptured, dull, and not noticeably punctate). Due to the confounding effect of the uncertain placement of
<taxonomicName id="2E824D69ACD143F309EEF8088D13019C" authorityName="Warncke" authorityYear="1975" box="[762,886,1987,2013]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD143F309EEF8088D13019C" box="[762,886,1987,2013]" italics="true" pageId="136" pageNumber="137">A. konyella</emphasis>
</taxonomicName>
, further study with genetic tools is required.
</paragraph>
<caption id="BDFD6662ACD043F20B83F85C8D3F01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209491" ID-Zenodo-Dep="13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="137" pageNumber="138" startId="137.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="137" targetType="figure">
<paragraph id="E93D36EAACD043F20B83F85C8D3F01B6" blockId="137.[151,1436,1943,2040]" pageId="137" pageNumber="138">
<emphasis id="DBF6EAF8ACD043F20B83F85C8F4401EE" bold="true" box="[151,289,1943,1967]" pageId="137" pageNumber="138">FIGURE 82.</emphasis>
<taxonomicName id="2E824D69ACD043F20A3DF85C8CEF01F1" authorityName="WOOD" authorityYear="2024" box="[297,650,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="perforata" status="sp. nov.">
<emphasis id="DBF6EAF8ACD043F20A3DF85C8F1A01EE" box="[297,383,1943,1967]" italics="true" pageId="137" pageNumber="138">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8ACD043F2093EF8538CEF01F1" box="[554,650,1944,1968]" italics="true" pageId="137" pageNumber="138">perforata</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8ACD043F20986F8538C9F01F1" bold="true" box="[658,762,1944,1968]" pageId="137" pageNumber="138">
<taxonomicNameLabel id="C0C55783ACD043F20986F8538C9F01F1" box="[658,762,1944,1968]" pageId="137" pageNumber="138" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view.
<taxonomicName id="2E824D69ACD043F20F41F8708F5501B6" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="konyella" subGenus="Aciandrena">
<emphasis id="DBF6EAF8ACD043F20F41F8708ACE0192" box="[1109,1195,1979,2003]" italics="true" pageId="137" pageNumber="138">Andrena</emphasis>
(
<emphasis id="DBF6EAF8ACD043F20FAAF8708B560192" box="[1214,1331,1979,2003]" italics="true" pageId="137" pageNumber="138">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8ACD043F20E52F8708BF90192" box="[1350,1436,1979,2003]" italics="true" pageId="137" pageNumber="138">konyella</emphasis>
Warncke, 1975
</taxonomicName>
female. D. Head, dorsal view; F. Scutum, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAACD343F10BD3FF5C8F4107B4" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
In addition to the presence of an inner tooth on the hind tarsal claws,
<taxonomicName id="2E824D69ACD343F108AAFF5C8A2006F0" authorityName="WOOD" authorityYear="2024" box="[958,1093,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F108AAFF5C8A2006F0" box="[958,1093,151,177]" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2E824D69ACD343F10FC2FF5C8B3606F0" authorityName="Warncke" authorityYear="1975" box="[1238,1363,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD343F10FC2FF5C8B3606F0" box="[1238,1363,151,177]" italics="true" pageId="138" pageNumber="139">A. konyella</emphasis>
</taxonomicName>
by the densely punctate frons, punctures separated by 0.5–1 puncture diameters, the underlying surface shiny and hence punctures are clearly visible (
<figureCitation id="71B92A6FACD343F10AF1FF148C0E06B8" box="[485,619,223,249]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="138" pageNumber="139">Figure 82C</figureCitation>
; in
<taxonomicName id="2E824D69ACD343F10981FF148D7106B8" authorityName="Warncke" authorityYear="1975" box="[661,788,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD343F10981FF148D7106B8" box="[661,788,223,249]" italics="true" pageId="138" pageNumber="139">A. konyella</emphasis>
</taxonomicName>
with punctures separated by 1–2 puncture diameters, the underlying surface shagreened and hence the punctures are less distinct,
<figureCitation id="71B92A6FACD343F108DEFEC88A2B075C" box="[970,1102,259,285]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="138" pageNumber="139">Figure 82D</figureCitation>
), by the shorter ocelloccipital distance of 1.5 times the diameter of a lateral ocellus (in
<taxonomicName id="2E824D69ACD343F1080CFEEC8DF30700" authorityName="Warncke" authorityYear="1975" box="[792,918,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD343F1080CFEEC8DF30700" box="[792,918,295,321]" italics="true" pageId="138" pageNumber="139">A. konyella</emphasis>
</taxonomicName>
with the ocelloccipital distance of 2 times the diameter of a lateral ocellus), by the dorsally narrower facial foveae, dorsally occupying ⅓ of the space between the compound eye and a lateral ocellus (in
<taxonomicName id="2E824D69ACD343F10956FEA48CDB07C8" authorityName="Warncke" authorityYear="1975" box="[578,702,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD343F10956FEA48CDB07C8" box="[578,702,367,393]" italics="true" pageId="138" pageNumber="139">A. konyella</emphasis>
</taxonomicName>
occupying slightly less than ½ of the space between the compound eye and a lateral ocellus), and by the more regularly and densely punctate scutum, with punctures separated by 1–2 puncture diameters (
<figureCitation id="71B92A6FACD343F10A97FE7C8C6F0790" box="[387,522,439,465]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="138" pageNumber="139">Figure 82E</figureCitation>
; in
<taxonomicName id="2E824D69ACD343F1092DFE7C8CDE0790" authorityName="Warncke" authorityYear="1975" box="[569,699,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="konyella">
<emphasis id="DBF6EAF8ACD343F1092DFE7C8CDE0790" box="[569,699,439,465]" italics="true" pageId="138" pageNumber="139">A. konyella</emphasis>
</taxonomicName>
with punctures irregular, separated by 1–4 puncture diameters,
<figureCitation id="71B92A6FACD343F10B83FE108F7D07B4" box="[151,280,475,501]" captionStart="FIGURE 82" captionStartId="137.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@137.[214,1373,181,1919]" captionTargetPageId="137" captionText="FIGURE 82. Andrena (incertae sedis) perforata spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Aciandrena) konyella Warncke, 1975 female. D. Head, dorsal view; F. Scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209491" httpUri="https://zenodo.org/record/13209491/files/figure.png" pageId="138" pageNumber="139">Figure 82F</figureCitation>
).
</paragraph>
<paragraph id="E93D36EAACD343F10BD3FE348B320578" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
Finally, comparison should be made to
<taxonomicName id="2E824D69ACD343F1098EFE348ADB0458" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" box="[666,1214,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="punctifrons" subGenus="Graecandrena">
<emphasis id="DBF6EAF8ACD343F1098EFE348CD70458" box="[666,690,511,537]" italics="true" pageId="138" pageNumber="139">A.</emphasis>
(
<emphasis id="DBF6EAF8ACD343F109D2FE348D0D0458" box="[710,872,511,537]" italics="true" pageId="138" pageNumber="139">Graecandrena</emphasis>
)
<emphasis id="DBF6EAF8ACD343F10869FE348D990458" box="[893,1020,511,537]" italics="true" pageId="138" pageNumber="139">punctifrons</emphasis>
Morawitz, 1876
</taxonomicName>
which is currently known only from a single specimen from
<collectingCountry id="9195767AACD343F10970FDE88C87047C" box="[612,738,547,573]" name="Uzbekistan" pageId="138" pageNumber="139">Uzbekistan</collectingCountry>
(
<typeStatus id="36398848ACD343F109FBFDE88D36047C" box="[751,851,547,573]" pageId="138" pageNumber="139" type="holotype">holotype</typeStatus>
illustrated by
<bibRefCitation id="8D134B1BACD343F108FBFDE88B4A047C" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[1007,1327,546,573]" pageId="138" pageNumber="139" pagination="105 - 176" refId="ref87433" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. Zookeys, 1120, 105 - 176. https: // doi. org / 10.3897 / zookeys. 1120.90206" type="journal article" year="2022">
Astafurova
<emphasis id="DBF6EAF8ACD343F10F65FDE88AC4047C" box="[1137,1185,547,573]" italics="true" pageId="138" pageNumber="139">et al</emphasis>
. 2022b: 152
</bibRefCitation>
). The two species are similar due to the small body size, ventrally narrow foveae, and densely punctate shiny frons. However,
<taxonomicName id="2E824D69ACD343F10B83FDA08F5204C4" authorityName="Morawitz" authorityYear="1876" box="[151,311,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="punctifrons">
<emphasis id="DBF6EAF8ACD343F10B83FDA08F5204C4" box="[151,311,619,645]" italics="true" pageId="138" pageNumber="139">A. punctifrons</emphasis>
</taxonomicName>
differs in many regards including its entirely black bodily pubescence (pale in
<taxonomicName id="2E824D69ACD343F10FD4FDA08B2F04C4" authorityName="WOOD" authorityYear="2024" box="[1216,1354,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F10FD4FDA08B2F04C4" box="[1216,1354,619,645]" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
), more rounded head which is 1.1 times wider than long (1.25 times wider than long in
<taxonomicName id="2E824D69ACD343F10F2EFD448AA004E8" authorityName="WOOD" authorityYear="2024" box="[1082,1221,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F10F2EFD448AA004E8" box="[1082,1221,655,681]" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
), polished clypeus with irregular punctures typically separated by&gt;1 puncture diameter (clypeus shagreened to weakly shining in
<taxonomicName id="2E824D69ACD343F10B83FD1C8F4604B0" authorityName="WOOD" authorityYear="2024" box="[151,291,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F10B83FD1C8F4604B0" box="[151,291,727,753]" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
, with punctures rarely separated by&gt;1 puncture diameter), propodeal triangle with raised rugae in basal half, hence the placement in the subgenus
<taxonomicName id="2E824D69ACD343F109D3FD308D0C0554" authorityName="Warncke" authorityYear="1968" box="[711,873,763,789]" pageId="138" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8ACD343F109D3FD308D0C0554" box="[711,873,763,789]" italics="true" pageId="138" pageNumber="139">Graecandrena</emphasis>
</taxonomicName>
(propodeal triangle with granular shagreen in
<taxonomicName id="2E824D69ACD343F10E90FD308F640578" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F10E90FD308F640578" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
), and scutum and scutellum polished and shining (shagreened and weakly shining in
<taxonomicName id="2E824D69ACD343F10FA9FCD48B220578" authorityName="WOOD" authorityYear="2024" box="[1213,1351,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="DBF6EAF8ACD343F10FA9FCD48B220578" box="[1213,1351,799,825]" italics="true" pageId="138" pageNumber="139">A. perforata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD343F10BD3FC888C9105C0" pageId="138" pageNumber="139" type="etymology">
<paragraph id="E93D36EAACD343F10BD3FC888C9105C0" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
<emphasis id="DBF6EAF8ACD343F10BD3FC888F2B051C" bold="true" box="[199,334,835,861]" pageId="138" pageNumber="139">Etymology.</emphasis>
Nominative feminine singular form of the Latin adjective
<emphasis id="DBF6EAF8ACD343F108D9FC888A27051C" box="[973,1090,835,861]" italics="true" pageId="138" pageNumber="139">perforatus</emphasis>
, meaning pierced or penetrated in reference to the densely punctate frons and scutum.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD343F10BD3FC418D6F05E4" box="[199,778,906,933]" pageId="138" pageNumber="139" type="distribution">
<paragraph id="E93D36EAACD343F10BD3FC418D6F05E4" blockId="138.[151,1437,151,933]" box="[199,778,906,933]" pageId="138" pageNumber="139">
<emphasis id="DBF6EAF8ACD343F10BD3FC418F0405E5" bold="true" box="[199,353,906,932]" pageId="138" pageNumber="139">Distribution.</emphasis>
Central
<collectingCountry id="9195767AACD343F10AD4FC408C7505E4" box="[448,528,907,933]" name="Turkey" pageId="138" pageNumber="139">Turkey</collectingCountry>
(province of
<collectingRegion id="2B46F808ACD343F109B2FC408C9B05E4" box="[678,766,907,933]" country="Turkey" name="Ankara" pageId="138" pageNumber="139">Ankara</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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