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<document id="1170D7CEC222D4BA5554FBEED6156FE2" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCAC52436B0B83FA048FA507EC" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Andrena (Avandrena) collisa WOOD, 2024, spec. nov." docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="12" updateTime="1722787353532" updateUser="ExternalLinkService">
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<mods:title id="0D9C87209DB12BF20B492B4CAD8172B1">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="5F0F2B80C87A4C88C6CC730023A4026E">WOOD, T. J.</mods:namePart>
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<treatment id="612B87FCAC52436B0B83FA048FA507EC" ID-DOI="http://doi.org/10.5281/zenodo.13209437" ID-Zenodo-Dep="13209437" LSID="urn:lsid:plazi:treatment:612B87FCAC52436B0B83FA048FA507EC" httpUri="http://treatment.plazi.org/id/612B87FCAC52436B0B83FA048FA507EC" lastPageId="16" lastPageNumber="17" pageId="11" pageNumber="12">
<subSubSection id="A1986561AC5243700B83FA048C1103AB" box="[151,628,1487,1514]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="E93D36EAAC5243700B83FA048C1103AB" blockId="11.[151,628,1487,1550]" box="[151,628,1487,1514]" pageId="11" pageNumber="12">
<heading id="B2758186AC5243700B83FA048C1103AB" bold="true" box="[151,628,1487,1514]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="DBF6EAF8AC5243700B83FA048C1103AB" bold="true" box="[151,628,1487,1514]" pageId="11" pageNumber="12">
<taxonomicName id="2E824D69AC5243700B83FA048F9003AB" authorityName="WOOD" authorityYear="2024" box="[151,501,1487,1514]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="collisa" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5243700B83FA048F6403AB" bold="true" box="[151,257,1487,1514]" italics="true" pageId="11" pageNumber="12">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5243700A06FA048FF003AB" bold="true" box="[274,405,1487,1514]" italics="true" pageId="11" pageNumber="12">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC5243700AB3FA048F9003AB" bold="true" box="[423,501,1487,1514]" italics="true" pageId="11" pageNumber="12">collisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC5243700AE8FA1B8C1103AB" box="[508,628,1488,1514]" pageId="11" pageNumber="12" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC5243700B83FA3F8C68004F" box="[151,525,1524,1550]" pageId="11" pageNumber="12" type="description">
<paragraph id="E93D36EAAC5243700B83FA3F8C68004F" blockId="11.[151,628,1487,1550]" box="[151,525,1524,1550]" pageId="11" pageNumber="12">
<figureCitation id="71B92A6FAC5243700B83FA3F8F1E004F" box="[151,379,1524,1550]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="11" pageNumber="12">Figures 6A–C; E; G</figureCitation>
;
<figureCitation id="71B92A6FAC5243700A93FA3F8C6C004F" box="[391,521,1524,1550]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="11" pageNumber="12">7A–C; E–F</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC5243700B83F9F28A3000DA" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="E93D36EAAC5243700B83F9F28FCE0039" blockId="11.[151,1436,1593,2037]" pageId="11" pageNumber="12">
<materialsCitation id="59EA3CB7AC5243700B83F9F28FCE0039" collectingDate="2019-03-02" collectorName="M. Mikat &amp; J. Waldhauserova" country="Cyprus" location="Lefkara" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Larnaka" typeStatus="holotype">
<emphasis id="DBF6EAF8AC5243700B83F9F28FA30012" bold="true" box="[151,454,1593,1619]" pageId="11" pageNumber="12">
<typeStatus id="36398848AC5243700B83F9F28F5F0012" box="[151,314,1593,1619]" pageId="11" pageNumber="12" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC5243700A5FF9F28FA40012" box="[331,449,1593,1619]" name="Cyprus" pageId="11" pageNumber="12">CYPRUS</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC5243700AC6F9F18CEC0012" box="[466,649,1593,1620]" country="Cyprus" name="Larnaka" pageId="11" pageNumber="12">Larnaca district</collectingRegion>
,
<location id="EC5D6031AC524370098CF9F28C970012" LSID="urn:lsid:plazi:treatment:612B87FCAC52436B0B83FA048FA507EC:EC5D6031AC524370098CF9F28C970012" box="[664,754,1593,1619]" country="Cyprus" name="Lefkara" pageId="11" pageNumber="12" stateProvince="Larnaka">Lefkara</location>
, 34.8852
<superScript id="1EF79BA2AC5243700849F9F28D190015" attach="right" box="[861,892,1593,1620]" fontSize="6" pageId="11" pageNumber="12">oN</superScript>
, 33.3235
<superScript id="1EF79BA2AC52437008F3F9F28A670015" attach="right" box="[999,1026,1593,1620]" fontSize="6" pageId="11" pageNumber="12">oE</superScript>
,
<date id="9D3C102AAC5243700F07F9F28AE70012" box="[1043,1154,1593,1619]" pageId="11" pageNumber="12" value="2019-03-02">
<collectingDate id="8D78E9C2AC5243700F07F9F28AE70012" box="[1043,1154,1593,1619]" pageId="11" pageNumber="12" value="2019-03-02">2.iii.2019</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC5243700F85F9F18ACD0015" box="[1169,1192,1594,1620]" count="1" pageId="11" pageNumber="12" type="male">♂</specimenCount>
, leg.
<collectorName id="4477533CAC5243700FFAF9F18B380012" box="[1262,1373,1593,1620]" pageId="11" pageNumber="12">M. Mikát</collectorName>
&amp;
<collectorName id="4477533CAC5243700E9EF9F18F230037" pageId="11" pageNumber="12">J. Waldhauserová</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC5243700BD3F9B58A3000DA" blockId="11.[151,1436,1593,2037]" box="[199,1109,1662,1691]" pageId="11" pageNumber="12">
<materialsCitation id="59EA3CB7AC5243700BD3F9B58A3500DA" box="[199,1104,1662,1691]" collectingDate="1984-03-04" collectorName="H. Weiffenbach" country="Cyprus" location="Limassol" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="Lemesos" typeStatus="paratype">
<emphasis id="DBF6EAF8AC5243700BD3F9B58F8200D9" bold="true" box="[199,487,1662,1688]" pageId="11" pageNumber="12">
<typeStatus id="36398848AC5243700BD3F9B58F0500D9" box="[199,352,1662,1688]" pageId="11" pageNumber="12" type="paratype">PARATYPE</typeStatus>
:
<collectingCountry id="9195767AAC5243700A78F9B58F8700D9" box="[364,482,1662,1688]" name="Cyprus" pageId="11" pageNumber="12">CYPRUS</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC5243700AFAF9B58C3D00D9" box="[494,600,1662,1688]" country="Cyprus" name="Lemesos" pageId="11" pageNumber="12">Limassol</collectingRegion>
,
<date id="9D3C102AAC5243700976F9B58CB500D9" box="[610,720,1662,1688]" pageId="11" pageNumber="12" value="1984-03-04">
<collectingDate id="8D78E9C2AC5243700976F9B58CB500D9" box="[610,720,1662,1688]" pageId="11" pageNumber="12" value="1984-03-04">4.iii.1984</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC52437009C8F9B48C9A00D8" box="[732,767,1663,1689]" count="1" pageId="11" pageNumber="12" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC5243700829F9B48D9500D9" box="[829,1008,1662,1689]" pageId="11" pageNumber="12">H. Weiffenbach</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC5243760BD3F96A8F770048" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="E93D36EAAC5243700BD3F96A8FA301B4" blockId="11.[151,1436,1593,2037]" pageId="11" pageNumber="12">
<emphasis id="DBF6EAF8AC5243700BD3F96A8FA600FA" bold="true" box="[199,451,1697,1723]" pageId="11" pageNumber="12">Description. Female.</emphasis>
Body length:
<quantity id="2E7A9B0FAC5243700966F9698CDD00FD" box="[626,696,1698,1724]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
(
<figureCitation id="71B92A6FAC52437009DFF96A8D2D00FA" box="[715,840,1697,1723]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="11" pageNumber="12">Figure 6A</figureCitation>
).
<emphasis id="DBF6EAF8AC5243700875F96A8DC300FA" box="[865,934,1697,1723]" italics="true" pageId="11" pageNumber="12">Head:</emphasis>
Dark, 1.45 times wider than long (
<figureCitation id="71B92A6FAC5243700E45F96A8EDE009F" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="11" pageNumber="12">Figure 6C</figureCitation>
). Clypeus weakly domed, moderately flattened medially, weakly punctate, punctures slightly latitudinally compressed, separated by 1 puncture diameter; underlying surface with granular shagreen, weakly shining, punctures disappearing into underlying sculpture. Process of labrum narrow, slightly wider than long, with deep narrow emargination medially. Gena equalling width of compound eye; ocelloccipital distance equalling diameter of lateral ocellus. Foveae dorsally occupying ½ space between compound eye and lateral ocellus, ventrally slightly narrowing, reaching upper level of antennal insertions; foveae filled with dark brown hairs. Head predominantly covered with long black hairs, with some pale hairs intermixed on frons and vertex; longest hairs equalling length of scape. Antennae basally dark, A6–12 ventrally lightened dark orange, ventral surface covered with isolated silvery scales; A3 equalling A4+5.
</paragraph>
<caption id="BDFD6662AC5543770B83F85C8D5401B6" ID-DOI="http://doi.org/10.5281/zenodo.13209319" ID-Zenodo-Dep="13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="12" targetType="figure">
<paragraph id="E93D36EAAC5543770B83F85C8D5401B6" blockId="12.[151,1437,1943,2040]" pageId="12" pageNumber="13">
<emphasis id="DBF6EAF8AC5543770B83F85C8F7101EE" bold="true" box="[151,276,1943,1967]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
<taxonomicName id="2E824D69AC5543770A08F85C8C2101EE" authorityName="WOOD" authorityYear="2024" box="[284,580,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="collisa" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5543770A08F85C8F1701EE" box="[284,370,1943,1967]" italics="true" pageId="12" pageNumber="13">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5543770A97F85C8F9501EE" box="[387,496,1943,1967]" italics="true" pageId="12" pageNumber="13">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC5543770914F85C8C2101EE" box="[512,580,1943,1967]" italics="true" pageId="12" pageNumber="13">collisa</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC554377095FF8538CD601F1" bold="true" box="[587,691,1944,1968]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="C0C55783AC554377095FF8538CD601F1" box="[587,691,1944,1968]" pageId="12" pageNumber="13" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view;
<taxonomicName id="2E824D69AC5543770813F8708A8D0195" authority="(Friese, 1923)" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[775,1256,1979,2004]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="canohirta" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5543770813F8708D380192" box="[775,861,1979,2003]" italics="true" pageId="12" pageNumber="13">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC554377087BF8708DB90192" box="[879,988,1979,2003]" italics="true" pageId="12" pageNumber="13">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC55437708F9F8708A350192" box="[1005,1104,1979,2003]" italics="true" pageId="12" pageNumber="13">canohirta</emphasis>
(
<bibRefCitation id="8D134B1BAC5543770F4BF8708A840195" author="Friese, H." box="[1119,1249,1979,2004]" pageId="12" pageNumber="13" pagination="175 - 216" refId="ref88760" refString="Friese, H. (1923) Eine Kriegsausbeute an Apiden (Bienen) aus Makedonien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 46, 175 - 216." type="journal article" year="1923">Friese, 1923</bibRefCitation>
)
</taxonomicName>
female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC5443760BD3FF5C8AA80458" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3FF5C8F2D06F1" box="[199,328,151,176]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="2E824D69AC5443760BD3FF5C8F2706F1" box="[199,322,151,176]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Scutum and scutellum finely shagreened, laterally weakly shining, shagreen becoming weaker medially, here almost shining; surface punctate, punctures separated by 1–2 puncture diameters (
<figureCitation id="71B92A6FAC5443760E01FF708BF50694" box="[1301,1424,187,213]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6E</figureCitation>
). Pronotum laterally with weak humeral angle. Mesepisternum and dorsolateral parts of propodeum with fine granular shagreen, weakly shining. Propodeal triangle, laterally delineated by fine carinae, internal surface with weaky raised fine rugae in basal half (
<figureCitation id="71B92A6FAC5443760AA0FEEC8C4A0700" box="[436,559,295,321]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6G</figureCitation>
). Mesepisternum with long black plumose hairs, longest equalling length of scape. Scutum and scutellum with shorter predominantly white plumose hairs, with scattered intermixed black hairs. Propodeal corbicula incomplete, sparse, dorsal fringe composed of pale plumose hairs, internal surface with abundant simple light brownish hairs. Legs dark brown, pubescence light brown. Posterior face of hind femorae rounded, without spines. Flocculus incomplete, composed of whitish plumose hairs; femoral scopae composed of white simple hairs, tibial scopae composed of simple hairs, ventrally white, dorsally light brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark brown to orange, nervulus antefurcal.
</paragraph>
<paragraph id="E93D36EAAC5443760BD3FDE88B0C04B0" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3FDE88F20047D" box="[199,325,547,572]" italics="true" pageId="13" pageNumber="14">Metasoma:</emphasis>
Tergal discs dark, marginal areas weakly depressed, most strongly on T3–4, lightened hyaline-brown (
<figureCitation id="71B92A6FAC5443760BE5FD8C8F090420" box="[241,364,583,609]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6B</figureCitation>
). Tergal discs polished and shining, weak microsculpture present only narrowly basally; T1 sparsely punctate, punctures separated by 2–3 puncture diameters, T2–4 with punctures separated by 1–2 puncture diameters. Tergal discs with very short sparse white hairs, T1–4 apically with long whitish hairs, forming weak apical hairbands, broadly interrupted on T1–3, complete on T4, not obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate orange-brown. Pygidial plate rounded triangular, surface dull, featureless.
</paragraph>
<paragraph id="E93D36EAAC5443760BD3FD308F910274" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3FD308F6E0554" bold="true" box="[199,267,763,789]" pageId="13" pageNumber="14">Male.</emphasis>
Body length:
<quantity id="2E7A9B0FAC5443760AA0FD308F9C0554" box="[436,505,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="mm" value="6.0">6 mm</quantity>
(
<figureCitation id="71B92A6FAC544376091EFD308CE00554" box="[522,645,763,789]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7A</figureCitation>
).
<emphasis id="DBF6EAF8AC5443760989FD308C860554" box="[669,739,763,789]" italics="true" pageId="13" pageNumber="14">Head:</emphasis>
Dark, 1.5 times wider than long (
<figureCitation id="71B92A6FAC5443760F60FD308A880554" box="[1140,1261,763,789]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7B</figureCitation>
). Clypeus very weakly domed, broadly flattened over majority of area with exception of anterior ⅓, this slightly protruding; surface punctate, punctures separated by 0.5–1 puncture diameter with exception of weak longitudinal impunctate mid-line, underlying surface shagreened, weakly shining. Apical margin of clypeus with narrow yellow marking (
<figureCitation id="71B92A6FAC5443760E0EFCAC8BF505C0" box="[1306,1424,871,897]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7B</figureCitation>
). Process of labrum rounded trapezoidal, 2 times broader than long, anterior margin with shallow broad emargination. Gena slightly exceeding width of compound eye; ocelloccipital distance 2 times diameter of lateral ocellus (
<figureCitation id="71B92A6FAC5443760E45FC648EDE05AC" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7C</figureCitation>
). Face and scape covered with long black hairs, gena and vertex predominantly with white hairs, with occasional intermixed black hairs.Antennae basally dark, A6–12 ventrally lightened by presence of orange scales;A3 exceeding A4, slightly shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAAC5443760BD3FBF48C590284" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3FBF48F2D0219" box="[199,328,1087,1112]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="2E824D69AC5443760BD3FBF48F270219" box="[199,322,1087,1112]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
<taxonomicName id="2E824D69AC5443760A5AFBF48FA20218" box="[334,455,1087,1113]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
structurally as in female. Mesepisternum with long black plumose hairs, hairs exceeding length of scape. Scutum, scutellum, and propodeum with mixture of black and white plumose hairs. Legs dark, pubescence whitish to light brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark brown to orange, nervulus antefurcal.
</paragraph>
<paragraph id="E93D36EAAC5443760BD3FB048F770048" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3FB048F2002A9" box="[199,325,1231,1256]" italics="true" pageId="13" pageNumber="14">Metasoma:</emphasis>
Tergal discs dark, apical rims narrowly lightened hyaline-brown (
<figureCitation id="71B92A6FAC5443760F3DFB048AF802A8" box="[1065,1181,1231,1257]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7E</figureCitation>
). Tergal discs polished and shining, weak microsculpture present only narrowly basally; T1–4 punctate, punctures separated by 1–2 puncture diameters. Tergal discs with very short sparse white hairs, not forming apical hairbands. T6–7 with long dark brown hairs overlying pygidial plate of T7. S8 columnar, more or less parallel-sided until short broadened apical process; ventral surface medially with two lateral tufts of dense brown hair. Genital capsule slightly elongate, gonocoxae apically produced into weakly rounded teeth, gonostyli with apexes broadly produced into flattened triangular wedges with inner margins raised, stem part of gonostyli with raised longitudinal ridge along inner margins (
<figureCitation id="71B92A6FAC5443760E45FA6C8EDD03A4" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="13" pageNumber="14">Figure 7F</figureCitation>
). Penis valves moderately developed, occupying ½ space between gonostyli, medially broadened, apically narrowing.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC5443750BD3F9D98FEB07EC" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="E93D36EAAC5443760BD3F9D98F2900A0" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="14">
<emphasis id="DBF6EAF8AC5443760BD3F9D98F27006D" bold="true" box="[199,322,1554,1580]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC5443760A5DF9D88F9E006C" authorityName="WOOD" authorityYear="2024" box="[329,507,1555,1581]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="collisa">
<emphasis id="DBF6EAF8AC5443760A5DF9D88F9E006C" box="[329,507,1555,1581]" italics="true" pageId="13" pageNumber="14">Andrena collisa</emphasis>
</taxonomicName>
can be recognised as an
<taxonomicName id="2E824D69AC5443760802F9D88DEB006C" authorityName="Warncke" authorityYear="1968" box="[790,910,1555,1581]" pageId="13" pageNumber="9" rank="subGenus" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5443760802F9D88DEB006C" box="[790,910,1555,1581]" italics="true" pageId="13" pageNumber="14">Avandrena</emphasis>
</taxonomicName>
due to its short and broad head with short and broad foveae; recognition of the spineless
<taxonomicName id="2E824D69AC5443760969F9FC8C900010" authorityName="Warncke" authorityYear="1968" box="[637,757,1591,1617]" pageId="13" pageNumber="9" rank="subGenus" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5443760969F9FC8C900010" box="[637,757,1591,1617]" italics="true" pageId="13" pageNumber="14">Avandrena</emphasis>
</taxonomicName>
is always more challenging, since the subgenus is partially defined on the presence of this character, and
<taxonomicName id="2E824D69AC54437609B4F9908D6E0034" authorityName="WOOD" authorityYear="2024" box="[672,779,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="collisa">
<emphasis id="DBF6EAF8AC54437609B4F9908D6E0034" box="[672,779,1627,1653]" italics="true" pageId="13" pageNumber="14">A. collisa</emphasis>
</taxonomicName>
lacks spines on the hind face of the hind femorae. Due to their extensive black pubescence, particularly on the mesepisternum which is entirely black-haired, they can be compared to
<taxonomicName id="2E824D69AC5443760A24F9688C0C00FC" authority="(Friese, 1923)" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[304,617,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5443760A24F9688FDA00FC" box="[304,447,1699,1725]" italics="true" pageId="13" pageNumber="14">A. canohirta</emphasis>
(
<bibRefCitation id="8D134B1BAC5443760AC4F9688C0400FC" author="Friese, H." box="[464,609,1699,1725]" pageId="13" pageNumber="14" pagination="175 - 216" refId="ref88760" refString="Friese, H. (1923) Eine Kriegsausbeute an Apiden (Bienen) aus Makedonien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 46, 175 - 216." type="journal article" year="1923">Friese, 1923</bibRefCitation>
)
</taxonomicName>
(North
<collectingCountry id="9195767AAC54437609DCF9688D2200FC" box="[712,839,1699,1725]" name="Macedonia" pageId="13" pageNumber="14">Macedonia</collectingCountry>
,
<collectingCountry id="9195767AAC5443760841F9688DDD00FC" box="[853,952,1699,1725]" name="Bulgaria" pageId="13" pageNumber="14">Bulgaria</collectingCountry>
,
<collectingCountry id="9195767AAC54437608D3F9688A7C00FC" box="[967,1049,1699,1725]" name="Greece" pageId="13" pageNumber="14">Greece</collectingCountry>
,
<collectingCountry id="9195767AAC5443760F33F9688A1E00FC" box="[1063,1147,1699,1725]" name="Turkey" pageId="13" pageNumber="14">Turkey</collectingCountry>
; =
<taxonomicName id="2E824D69AC5443760F8CF9688EB700A0" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DBF6EAF8AC5443760F8CF9688B4D00FC" box="[1176,1320,1699,1725]" italics="true" pageId="13" pageNumber="14">A. balcanica</emphasis>
<bibRefCitation id="8D134B1BAC5443760E26F9688EB700A0" author="Warncke, K." pageId="13" pageNumber="14" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
;
<figureCitation id="71B92A6FAC5443760BCAF90C8F5B00A0" box="[222,318,1735,1761]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-16@15.[213,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Andrena (Avandrena) canohirta (Friese, 1923) holotype female.A. Label details; B. Habitus, lateral view; Andrena (Avandrena) balcanica Warncke, 1965 holotype female. C. Label details; D. Habitus, lateral view; E. Face, frontal view; F. Scutum, dorsal view; G. Mesepisternum, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209323" httpUri="https://zenodo.org/record/13209323/files/figure.png" pageId="13" pageNumber="14">Figure 8</figureCitation>
).
</paragraph>
<paragraph id="E93D36EAAC5443750BD3F9208FEB07EC" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1437,151,429]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
In the female sex, the two species are very close, but
<taxonomicName id="2E824D69AC5443760856F9208DD50144" authorityName="WOOD" authorityYear="2024" box="[834,944,1771,1797]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="collisa">
<emphasis id="DBF6EAF8AC5443760856F9208DD50144" box="[834,944,1771,1797]" italics="true" pageId="13" pageNumber="14">A. collisa</emphasis>
</taxonomicName>
can be separated by the punctation of the clypeus which is obscurely punctate with latitudinally compressed punctures that disappear into the underlying clypeal sculpture (
<figureCitation id="71B92A6FAC5443760A73F8F88FBB010C" box="[359,478,1843,1869]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6C</figureCitation>
; in
<taxonomicName id="2E824D69AC544376091DF8F88CF3010C" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[521,662,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC544376091DF8F88CF3010C" box="[521,662,1843,1869]" italics="true" pageId="13" pageNumber="14">A. canohirta</emphasis>
</taxonomicName>
with the clypeus covered with shallow but large, round, and clearly visible punctures that do not disappear into the underlying sculpture,
<figureCitation id="71B92A6FAC54437608A6F89C8A4F0130" box="[946,1066,1879,1905]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6D</figureCitation>
), by the sculpture of the scutum which is laterally shagreened becoming polished medially (
<figureCitation id="71B92A6FAC544376082DF8B08DCB01D4" box="[825,942,1915,1941]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6E</figureCitation>
; in
<taxonomicName id="2E824D69AC54437608CDF8B08A0301D4" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[985,1126,1915,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC54437608CDF8B08A0301D4" box="[985,1126,1915,1941]" italics="true" pageId="13" pageNumber="14">A. canohirta</emphasis>
</taxonomicName>
with the scutum uniformly shagreened,
<figureCitation id="71B92A6FAC5443760A32F8548FFF01F8" box="[294,410,1951,1977]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="13" pageNumber="14">Figure 6F</figureCitation>
), by the ocelloccipital distance which is subequal to the diameter of a lateral ocellus (in
<taxonomicName id="2E824D69AC5443760B83F8088F40019C" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[151,293,1987,2013]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5443760B83F8088F40019C" box="[151,293,1987,2013]" italics="true" pageId="13" pageNumber="14">A. canohirta</emphasis>
</taxonomicName>
with the ocelloccipital distance 1.2 times the diameter of a lateral ocellus), by the finely delineated propodeal triangle with finely raised internal rugae (
<figureCitation id="71B92A6FAC57437509FFFF5C8D0106F0" box="[747,868,151,177]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="14" pageNumber="15">Figure 6G</figureCitation>
; in
<taxonomicName id="2E824D69AC5743750884FF5C8A7B06F0" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[912,1054,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5743750884FF5C8A7B06F0" box="[912,1054,151,177]" italics="true" pageId="14" pageNumber="15">A. canohirta</emphasis>
</taxonomicName>
with the propodeal triangle more coarsely delineated, internal surface slightly depressed and covered in more strongly raised internal rugae,
<figureCitation id="71B92A6FAC5743750E47FF708ED806B8" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="14" pageNumber="15">Figure 6H</figureCitation>
), by the golden-brown terminal fringe (
<figureCitation id="71B92A6FAC574375096FFF148C9406B8" box="[635,753,223,249]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="14" pageNumber="15">Figure 6B</figureCitation>
; in
<taxonomicName id="2E824D69AC5743750808FF148DCC06B8" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[796,937,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5743750808FF148DCC06B8" box="[796,937,223,249]" italics="true" pageId="14" pageNumber="15">A. canohirta</emphasis>
</taxonomicName>
with the terminal fringe dark brown,
<figureCitation id="71B92A6FAC5743750E47FF148ED8075C" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-16@15.[213,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Andrena (Avandrena) canohirta (Friese, 1923) holotype female.A. Label details; B. Habitus, lateral view; Andrena (Avandrena) balcanica Warncke, 1965 holotype female. C. Label details; D. Habitus, lateral view; E. Face, frontal view; F. Scutum, dorsal view; G. Mesepisternum, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209323" httpUri="https://zenodo.org/record/13209323/files/figure.png" pageId="14" pageNumber="15">Figure 8H</figureCitation>
), and by the slightly smaller body length of
<quantity id="2E7A9B0FAC57437509B1FEC88C83075C" box="[677,742,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="mm" value="7.0">7 mm</quantity>
(
<taxonomicName id="2E824D69AC57437509E1FEC88DE5075C" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[757,896,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC57437509E1FEC88DE5075C" box="[757,896,259,285]" italics="true" pageId="14" pageNumber="15">A. canohirta</emphasis>
</taxonomicName>
<quantity id="2E7A9B0FAC5743750893FEC88D86075C" box="[903,995,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="14" pageNumber="15" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
in length). Separation of males is much easier, as the ocelloccipital distance is 2 times the diameter of a lateral ocellus (
<figureCitation id="71B92A6FAC5743750F1CFEEC8A1B0700" box="[1032,1150,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="14" pageNumber="15">Figure 7A</figureCitation>
), whereas in
<taxonomicName id="2E824D69AC5743750E05FEEC8BF90700" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[1297,1436,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5743750E05FEEC8BF90700" box="[1297,1436,295,321]" italics="true" pageId="14" pageNumber="15">A. canohirta</emphasis>
</taxonomicName>
it is 3.5 times this diameter (
<figureCitation id="71B92A6FAC5743750AC9FE808C330724" box="[477,598,331,357]" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="14" pageNumber="15">Figure 7D</figureCitation>
). The clypeus is also apically marked with a small area of yellow (
<figureCitation id="71B92A6FAC5743750E45FE808EDF07C8" captionStart="FIGURE 7" captionStartId="14.[151,250,1917,1941]" captionTargetBox="[151,1436,453,1893]" captionTargetId="figure-163@14.[151,1436,453,1893]" captionTargetPageId="14" captionText="FIGURE 7. Andrena (Avandrena) collisa spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) male. D. Vertex, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="14" pageNumber="15">Figure 7B</figureCitation>
), whereas
<taxonomicName id="2E824D69AC5743750A25FEA48FD907C8" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[305,444,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC5743750A25FEA48FD907C8" box="[305,444,367,393]" italics="true" pageId="14" pageNumber="15">A. canohirta</emphasis>
</taxonomicName>
always has the clypeus black; additional material is necessary to establish whether this is a consistent character.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC5743750B83F8B68F77019C" ID-DOI="http://doi.org/10.5281/zenodo.13209321" ID-Zenodo-Dep="13209321" httpUri="https://zenodo.org/record/13209321/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1917,1941]" targetBox="[151,1436,453,1893]" targetPageId="14" targetType="figure">
<paragraph id="E93D36EAAC5743750B83F8B68F77019C" blockId="14.[151,1437,1917,2013]" pageId="14" pageNumber="15">
<emphasis id="DBF6EAF8AC5743750B83F8B68F7001D4" bold="true" box="[151,277,1917,1941]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
<taxonomicName id="2E824D69AC5743750A09F8B68C2001D4" authorityName="WOOD" authorityYear="2024" box="[285,581,1917,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="collisa" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5743750A09F8B68F1601D4" box="[285,371,1917,1941]" italics="true" pageId="14" pageNumber="15">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5743750A90F8B68F9401D4" box="[388,497,1917,1941]" italics="true" pageId="14" pageNumber="15">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC5743750915F8B68C2001D4" box="[513,581,1917,1941]" italics="true" pageId="14" pageNumber="15">collisa</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC5743750959F8B68CD301D4" bold="true" box="[589,694,1917,1941]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="C0C55783AC5743750959F8B68CD301D4" box="[589,694,1917,1941]" pageId="14" pageNumber="15" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC574375081BF86A8A8901F8" authority="(Friese, 1923)" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[783,1260,1953,1977]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="canohirta" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC574375081BF86A8D0001F8" box="[783,869,1953,1977]" italics="true" pageId="14" pageNumber="15">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5743750861F86A8D8701F8" box="[885,994,1953,1977]" italics="true" pageId="14" pageNumber="15">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC57437508E6F86A8A3001F8" box="[1010,1109,1953,1977]" italics="true" pageId="14" pageNumber="15">canohirta</emphasis>
(
<bibRefCitation id="8D134B1BAC5743750F70F86A8A8001F8" author="Friese, H." box="[1124,1253,1953,1977]" pageId="14" pageNumber="15" pagination="175 - 216" refId="ref88760" refString="Friese, H. (1923) Eine Kriegsausbeute an Apiden (Bienen) aus Makedonien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 46, 175 - 216." type="journal article" year="1923">Friese, 1923</bibRefCitation>
)
</taxonomicName>
male. D. Vertex, dorsal view.
</paragraph>
</caption>
<caption id="BDFD6662AC5643740B83F85C8DFA01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209323" ID-Zenodo-Dep="13209323" httpUri="https://zenodo.org/record/13209323/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1943,1967]" targetBox="[213,1374,181,1919]" targetPageId="15" targetType="figure">
<paragraph id="E93D36EAAC5643740B83F85C8DFA01B6" blockId="15.[151,1437,1943,2040]" pageId="15" pageNumber="16">
<emphasis id="DBF6EAF8AC5643740B83F85C8F7701EE" bold="true" box="[151,274,1943,1967]" pageId="15" pageNumber="16">FIGURE 8.</emphasis>
<taxonomicName id="2E824D69AC5643740A03F85C8C8F01F1" authority="(Friese, 1923)" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[279,746,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="canohirta" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5643740A03F85C8F0801EE" box="[279,365,1943,1967]" italics="true" pageId="15" pageNumber="16">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5643740A6FF85C8F8D01EE" box="[379,488,1943,1967]" italics="true" pageId="15" pageNumber="16">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC5643740AE2F85C8C3C01EE" box="[502,601,1943,1967]" italics="true" pageId="15" pageNumber="16">canohirta</emphasis>
(
<bibRefCitation id="8D134B1BAC5643740971F85C8C8101F1" author="Friese, H." box="[613,740,1943,1968]" pageId="15" pageNumber="16" pagination="175 - 216" refId="ref88760" refString="Friese, H. (1923) Eine Kriegsausbeute an Apiden (Bienen) aus Makedonien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 46, 175 - 216." type="journal article" year="1923">Friese, 1923</bibRefCitation>
)
</taxonomicName>
<emphasis id="DBF6EAF8AC56437409E4F8538D2801F1" bold="true" box="[752,845,1944,1968]" pageId="15" pageNumber="16">holotype</emphasis>
female.A. Label details; B. Habitus, lateral view;
<taxonomicName id="2E824D69AC5643740E52F85C8C410192" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="balcanica" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC5643740E52F85C8BF901EE" box="[1350,1436,1943,1967]" italics="true" pageId="15" pageNumber="16">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC5643740BB4F8708F680192" box="[160,269,1979,2003]" italics="true" pageId="15" pageNumber="16">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC5643740A09F8708FE40192" box="[285,385,1979,2003]" italics="true" pageId="15" pageNumber="16">balcanica</emphasis>
<bibRefCitation id="8D134B1BAC5643740A9DF8708C410192" author="Warncke, K." box="[393,548,1979,2003]" pageId="15" pageNumber="16" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC5643740939F8778CEF0195" bold="true" box="[557,650,1980,2004]" pageId="15" pageNumber="16">holotype</emphasis>
female. C. Label details; D. Habitus, lateral view; E. Face, frontal view; F. Scutum, dorsal view; G. Mesepisternum, lateral view; H. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC49436B0BD3FF5C8D970700" pageId="16" pageNumber="17" type="discussion">
<paragraph id="E93D36EAAC49436B0BD3FF5C8D970700" blockId="16.[151,1437,151,429]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3FF5C8F5E06F0" bold="true" box="[199,315,151,177]" pageId="16" pageNumber="17">Remarks.</emphasis>
<bibRefCitation id="8D134B1BAC49436B0A53FF5C8F8706F0" author="Friese, H." box="[327,482,151,177]" pageId="16" pageNumber="17" pagination="175 - 216" refId="ref88760" refString="Friese, H. (1923) Eine Kriegsausbeute an Apiden (Bienen) aus Makedonien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 46, 175 - 216." type="journal article" year="1923">Friese (1923)</bibRefCitation>
inconsistently referred to his material of
<taxonomicName id="2E824D69AC49436B08C6FF5C8A0606F0" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[978,1123,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC49436B08C6FF5C8A0606F0" box="[978,1123,151,177]" italics="true" pageId="16" pageNumber="17">A. canohirta</emphasis>
</taxonomicName>
, referring to a male on p. 186, then a female in the description of the species on p. 206, and then a male at the end of the description. The description is clearly of a female specimen, since Friese mentions the scopae. The only specimen in the ZMHB is a female (
<figureCitation id="71B92A6FAC49436B0A12FEC88FCF075C" box="[262,426,259,285]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-16@15.[213,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Andrena (Avandrena) canohirta (Friese, 1923) holotype female.A. Label details; B. Habitus, lateral view; Andrena (Avandrena) balcanica Warncke, 1965 holotype female. C. Label details; D. Habitus, lateral view; E. Face, frontal view; F. Scutum, dorsal view; G. Mesepisternum, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209323" httpUri="https://zenodo.org/record/13209323/files/figure.png" pageId="16" pageNumber="17">Figures 8A–B</figureCitation>
), and it matches the collecting details given by Friese; it is considered to automatically be the
<typeStatus id="36398848AC49436B0BD4FEEC8F400700" box="[192,293,295,321]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
, and references to males are simply lapsus on the part of Friese.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC49436B0BD3FE808A5507C8" pageId="16" pageNumber="17" type="etymology">
<paragraph id="E93D36EAAC49436B0BD3FE808A5507C8" blockId="16.[151,1437,151,429]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3FE808F2B0724" bold="true" box="[199,334,331,357]" pageId="16" pageNumber="17">Etymology.</emphasis>
Feminine singular nominative form of the Latin adjective
<emphasis id="DBF6EAF8AC49436B0F18FE808A070724" box="[1036,1122,331,357]" italics="true" pageId="16" pageNumber="17">collisus</emphasis>
meaning clashed together, conflicted, contended, in reference to the flattened or somewhat compressed head.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC49436B0BD3FE598FA507EC" box="[199,448,402,429]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="E93D36EAAC49436B0BD3FE598FA507EC" blockId="16.[151,1437,151,429]" box="[199,448,402,429]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3FE598F0407ED" bold="true" box="[199,353,402,428]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="9195767AAC49436B0A73FE588FD907EC" box="[359,444,403,429]" name="Cyprus" pageId="16" pageNumber="17">Cyprus</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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