278 lines
40 KiB
XML
278 lines
40 KiB
XML
<document id="556BB39D29E7DEBC7B06CF512FC9EA8E" ID-DOI="10.15298/rusentj.32.2.13" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178585" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722594461677" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2023" docId="527C87DDBB07FFD402CE184663446B9A" docLanguage="en" docName="RussEntomolJ.32.2.221-233.pdf" docOrigin="Russian Entomological Journal 32 (2)" docSource="http://dx.doi.org/10.15298/rusentj.32.2.13" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Empis (Polyblepharis) fandariensis Shamshev 2023, sp.n." docType="treatment" docVersion="2" lastPageNumber="224" masterDocId="AE45FFA5BB06FFD702081C4E61466F6D" masterDocTitle="Dance flies (Diptera: Empididae) in A. P. Fedtschenko’s Collection from Turkestan: Empis subgenus Polyblepharis" masterLastPageNumber="233" masterPageNumber="221" pageNumber="222" updateTime="1722634914522" updateUser="ExternalLinkService">
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<mods:titleInfo id="8CA4928C10FD2237AB5FBB06B2B5FB1A">
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<mods:title id="7940704B282984839B4C99F38523BC4B">Dance flies (Diptera: Empididae) in A. P. Fedtschenko’s Collection from Turkestan: Empis subgenus Polyblepharis</mods:title>
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<mods:name id="B7E2A775143B3F68E198C06454746495" type="personal">
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<mods:roleTerm id="90933CB29652D33DA7ED01ABAF93A40C">Author</mods:roleTerm>
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<mods:namePart id="87541627961CA012187290688CBB8462">Shamshev, I. V.</mods:namePart>
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<mods:typeOfResource id="7CE5FDB6BC57A808908D7F491F65F87B">text</mods:typeOfResource>
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<mods:title id="C198FFC02DEE2F27AC1F9D4EC8A3F72D">Russian Entomological Journal</mods:title>
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</mods:titleInfo>
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<mods:part id="101D633EDD337DA28E43AF21EB2FBDD5">
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<mods:date id="419FC5DAB79A97660C9E68012A5288D3">2023</mods:date>
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<mods:detail id="805199ECFCB44AFA68090F25C06385AD" type="pubDate">
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<mods:number id="C78957B9CC54D706106E1B71EADF0293">2023-06-30</mods:number>
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<mods:detail id="DC83D6B230EB6064BA2E9E19917A2F7B" type="volume">
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<mods:number id="B7220BDF8D10B83EBF3A3BDAAF64BC25">32</mods:number>
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<mods:start id="EDE70E335239955B242F3EC8A5A2FD02">221</mods:start>
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<mods:end id="0D60B1BAAE2C991C9E1A67A856A09660">233</mods:end>
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<mods:url id="3BA49947FA009E18210F7FF06938A66F">http://dx.doi.org/10.15298/rusentj.32.2.13</mods:url>
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<mods:classification id="FC214E6E8845C0F6F34746FE8A94B7FA">journal article</mods:classification>
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<mods:identifier id="E7D9A63B9F92AE377954594D3CEB1129" type="DOI">10.15298/rusentj.32.2.13</mods:identifier>
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<mods:identifier id="5C4E011DAED52CE76DC1949E5802E502" type="ISSN">0132-8069</mods:identifier>
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<mods:identifier id="1B69A004FE622F8EB72211D4198FDFE8" type="Zenodo-Dep">13178585</mods:identifier>
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</mods:mods>
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<treatment id="527C87DDBB07FFD402CE184663446B9A" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A" httpUri="http://treatment.plazi.org/id/527C87DDBB07FFD402CE184663446B9A" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222">
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<subSubSection id="92CF6540BB07FFD602CE184663BB6B72" box="[198,765,1031,1055]" pageId="1" pageNumber="222" type="nomenclature">
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<paragraph id="DA6A36CBBB07FFD602CE184663BB6B72" blockId="1.[189,776,1031,2063]" box="[198,765,1031,1055]" pageId="1" pageNumber="222">
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<heading id="812281A7BB07FFD602CE184663BB6B72" box="[198,765,1031,1055]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="222" reason="8">
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<taxonomicName id="1DD54D48BB07FFD602CE184663FA6B72" ID-CoL="C2Q7T" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[198,700,1031,1055]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="222" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
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<emphasis id="E8A1EAD9BB07FFD602CE1846604F6B72" box="[198,265,1032,1055]" italics="true" pageId="1" pageNumber="222">Empis</emphasis>
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(
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<emphasis id="E8A1EAD9BB07FFD60311184960E96B72" box="[281,431,1031,1055]" italics="true" pageId="1" pageNumber="222">Polyblepharis</emphasis>
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)
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<emphasis id="E8A1EAD9BB07FFD603C8184963026B72" box="[448,580,1031,1055]" italics="true" pageId="1" pageNumber="222">fandariensis</emphasis>
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Shamshev
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</taxonomicName>
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,
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<emphasis id="E8A1EAD9BB07FFD600C0184963BB6B72" bold="true" box="[712,765,1031,1055]" pageId="1" pageNumber="222">
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<taxonomicNameLabel id="F39257A2BB07FFD600C0184963BB6B72" box="[712,765,1031,1055]" pageId="1" pageNumber="222" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="92CF6540BB07FFD603A7186A63526B51" box="[431,532,1060,1084]" pageId="1" pageNumber="222" type="description">
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<paragraph id="DA6A36CBBB07FFD603A7186A63526B51" blockId="1.[189,776,1031,2063]" box="[431,532,1060,1084]" pageId="1" pageNumber="222">
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<figureCitation id="42EE2A4EBB07FFD603A7186A63566B51" box="[431,528,1060,1084]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="1" pageNumber="222">Figs 1–5</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="92CF6540BB07FFD602EA180763036BB9" pageId="1" pageNumber="222" type="materials_examined">
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<paragraph id="DA6A36CBBB07FFD602EA180763D86BC9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
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<materialsCitation id="6ABD3C96BB07FFD602EA180763D86BC9" collectingDate="1869-06-19" collectionCode="MATERIAL, ZMMU" collectorName="A. Fedtschenko & Cyrillic & Russian & Fan" country="Tajikistan" location="Fan Darya River" pageId="1" pageNumber="222" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectionCode id="BCC4AE0EBB07FFD602EA180760176B31" box="[226,337,1097,1116]" pageId="1" pageNumber="222">MATERIAL</collectionCode>
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EXAMINED.
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<typeStatus id="056E8869BB07FFD603D21807636B6B31" box="[474,557,1097,1116]" pageId="1" pageNumber="222" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="CCD3FD42BB07FFD600311809630B6B30" box="[569,589,1095,1117]" count="1" pageId="1" pageNumber="222" type="male">♂</specimenCount>
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, labels as in
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<figureCitation id="42EE2A4EBB07FFD600C3180762446B31" box="[715,770,1097,1116]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="1" pageNumber="222">Fig. 4</figureCitation>
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, [
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<emphasis id="E8A1EAD9BB07FFD602CC182F60656B19" bold="true" box="[196,291,1121,1140]" pageId="1" pageNumber="222">
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<collectingCountry id="A2C2765BBB07FFD602CC182F60656B19" box="[196,291,1121,1140]" name="Tajikistan" pageId="1" pageNumber="222">Tajikistan</collectingCountry>
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</emphasis>
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] small bluish green label only with number 19. [=
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<date id="AE6B100BBB07FFD602B51837605D6BE1" box="[189,283,1145,1164]" pageId="1" pageNumber="222" value="1869-06-19">
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<collectingDate id="BE2FE9E3BB07FFD602B51837605D6BE1" box="[189,283,1145,1164]" pageId="1" pageNumber="222" value="1869-06-19">19.vi.1869</collectingDate>
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</date>
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] [
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<collectorName id="7720531DBB07FFD60326183760FA6BE1" box="[302,444,1145,1164]" pageId="1" pageNumber="222">A. Fedtschenko</collectorName>
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] // [printed in
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<collectorName id="7720531DBB07FFD6004A183763C06BE1" box="[578,646,1145,1164]" pageId="1" pageNumber="222">Cyrillic</collectorName>
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,
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<collectorName id="7720531DBB07FFD6009A1837639F6BE1" box="[658,729,1145,1164]" pageId="1" pageNumber="222">Russian</collectorName>
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, pre- 1918 orthography]
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<collectorName id="7720531DBB07FFD6036E18DC60C06BC8" box="[358,390,1170,1189]" pageId="1" pageNumber="222">Fan</collectorName>
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[=
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<location id="DF0A6010BB07FFD603AE18DC637D6BC9" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A:DF0A6010BB07FFD603AE18DC637D6BC9" box="[422,571,1169,1189]" country="Tajikistan" name="Fan Darya River" pageId="1" pageNumber="222">Fan Darya River</location>
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] (
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<collectionCode id="BCC4AE0EBB07FFD6004418DF63D26BC9" box="[588,660,1169,1188]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="222" type="Museum">ZMMU</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="DA6A36CBBB07FFD602EA18E463036BB9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
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<materialsCitation id="6ABD3C96BB07FFD602EA18E463076BB9" collectingDate="1869-06-19" collectionCode="ZMMU, ZISP" collectorName="A. Fedtschenko & Cyrillic & Russian & Fan" country="Tajikistan" location="Fan Darya River" pageId="1" pageNumber="222" specimenCount="7" specimenCount-female="3" specimenCount-male="4" typeStatus="paratype">
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<typeStatus id="056E8869BB07FFD602EA18E4607F6BD0" box="[226,313,1194,1213]" pageId="1" pageNumber="222" type="paratype">Paratypes</typeStatus>
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.
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<emphasis id="E8A1EAD9BB07FFD6034D18E760ED6BD1" bold="true" box="[325,427,1193,1212]" pageId="1" pageNumber="222">
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<collectingCountry id="A2C2765BBB07FFD6034D18E760E16BD1" box="[325,423,1193,1212]" name="Tajikistan" pageId="1" pageNumber="222">Tajikistan</collectingCountry>
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:
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</emphasis>
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same data as holotype (
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<specimenCount id="CCD3FD42BB07FFD6008318E763816BD0" box="[651,711,1191,1213]" count="2" pageId="1" pageNumber="222" type="male">2 ♂♂</specimenCount>
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,
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<specimenCount id="CCD3FD42BB07FFD600DE18E762476BD0" box="[726,769,1191,1213]" count="2" pageId="1" pageNumber="222" type="female">2 ♀♀</specimenCount>
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,
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<collectionCode id="BCC4AE0EBB07FFD602B5188C60446BB8" box="[189,258,1218,1237]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="222" type="Museum">ZMMU</collectionCode>
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;
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<specimenCount id="CCD3FD42BB07FFD60304188F60036BB8" box="[268,325,1215,1237]" count="2" pageId="1" pageNumber="222" type="male">2 ♂♂</specimenCount>
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(both dissected),
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<specimenCount id="CCD3FD42BB07FFD603EA188F60B96BB8" box="[482,511,1215,1237]" count="1" pageId="1" pageNumber="222" type="female">1 ♀</specimenCount>
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,
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<collectionCode id="BCC4AE0EBB07FFD60003188F637D6BB9" box="[523,571,1217,1236]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="222">ZISP</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="92CF6540BB07FFD602EA189760C76AA9" pageId="1" pageNumber="222" type="diagnosis">
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<paragraph id="DA6A36CBBB07FFD602EA189760C76AA9" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
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DIAGNOSIS. Mid-sized species (body about
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<quantity id="1D2D9B2EBB07FFD600B2189763BA6B83" box="[698,764,1241,1263]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="222" unit="mm" value="5.0">5 mm</quantity>
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); male eyes holoptic; proboscis with labrum nearly 2X longer than eye height; palpus mostly brownish in male, yellow in female; mesoscutum with 3 brownish vittae along rows of acrostichal and dorsocentral setae; acrostichal setae irregularly 2–3-serial, presutural dorsocentrals irregularly 3–4- serial, laterotergite with pale setae; abdominal tergites 2–6 extensively shiny (lateral view); female mid and hind femora with flattened setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="92CF6540BB07FFD402EA198763F86BCA" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222" type="description">
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<paragraph id="DA6A36CBBB07FFD602EA198760986802" blockId="1.[189,776,1031,2063]" pageId="1" pageNumber="222">
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DESCRIPTION. Body length
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<quantity id="1D2D9B2EBB07FFD60000198763386AB2" box="[520,638,1481,1503]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.1" metricValueMin="4.8" pageId="1" pageNumber="222" unit="mm" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8–5.1 mm</quantity>
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, wing
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<quantity id="1D2D9B2EBB07FFD600B6198761A36A94" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="1" pageNumber="222" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
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.
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<emphasis id="E8A1EAD9BB07FFD602FC19AA606E6A94" bold="true" box="[244,296,1508,1529]" pageId="1" pageNumber="222">Male</emphasis>
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(
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<figureCitation id="42EE2A4EBB07FFD6033319AA603C6A94" box="[315,378,1508,1530]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
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). Head capsule regions black, mostly densely greyish pruinose, face along lower margin and clypeus shiny. Eyes holoptic, upper ommatidia enlarged. Frons represented by small subtriangular space just below ocellar triangle and larger space above antennae, bare. Face broad, bare. Ocellar setae black, long, fine. Occiput covered with numerous, long, fine, black setae (including postoculars); postgena with pale, long, hair-like setae. Antenna with scape and pedicel brownish, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with short setulae; postpedicel nearly 3X as long as basal width; stylus nearly as long as postpedicel basal width. Proboscis with labrum brownish, paler apically; nearly 2X longer than eye height; palpus short, mostly brown, rather yellowish brown at tip; bearing long, black setae.
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</paragraph>
|
||
<paragraph id="DA6A36CBBB07FFD602EA1B3A641B6DC1" blockId="1.[189,776,1031,2063]" lastBlockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">Thorax black in ground-colour, mostly densely greyish pruinose; mesoscutum with 3 brownish, indistinct vittae along rows of acrostichal and dorsocentral setae. Prosternum bare. Proepisternum with tuft of numerous pale, fine setae on lower part, bare on upper part. Antepronotum with 4–5 black, short setae dorsally and numerous pale, long, fine setae laterally. Postpronotal lobe with 1 long, strong, black and numerous short, fine, black and pale setae. Mesonotal setae black (except noted): acrostichals arranged in 2–3 irregular rows (often entirely biserial), rather long, fine, absent on prescutellar depression; presutural dorsocentral setae arranged in 3–4 irregular rows, similar to acrostichals; postsutural dorsocentrals sparser,becoming uniserial,longer and stronger towards scutellum, 1 pair of prescutellars longest; 1 rather fine presutural supra-alar; 3 notopleurals, 2 postsutural supra-alars (anterior seta shorter and finer), 1 long and 1 minute postalars, 4 scutellars (setae of lateral pair slightly shorter); in addition, supra-alar face with numerous black and pale setulae, notopleuron with similar pale setulae anteriorly. Laterotergite with numerous pale setae. Anterior and posterior spiracles pale.</paragraph>
|
||
<paragraph id="DA6A36CBBB07FFD6015A1EFF62CC6901" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">Legs with coxae densely greyish pruinose, remaining podomeres subshiny. Legs colour: almost entirely brown, usually only hind femur at apex and hind tibia near base narrowly brownish yellow (sometimes hind femur entirely brownish). Legs mostly black setose. Fore coxa covered with pale, dense, hair-like setae (2–3 stronger, black, subapical setae); hind trochanter with numerous pale, short, fine setae, 4–5 brownish to black, long, strong, subapical setae ventrally and 1–2 similar setae dorsally. Fore and mid femora whitish pubescent ventrally; hind femur with indistinct ventral pubescence only closer to apex. Fore femur with rows of fine anteroventral and posteroventral setae, which are pale on about basal half, short closer to apex and becoming longer towards base (longest setae nearly as long as femur width); dense, long, fine setae posteriorly (pale closer to base). Fore tibia with 4–6 short anterodorsal setae and subequal to them in length, fine setae posterodorsally (2–3 setae on apical half usually stronger). Fore basitarsus slender; with 2 short anterodorsal setae; anteroventral and posteroventral spine-like setae. Mid femur with rows of subequally long anteroventral and posteroventral setae; additional fine setae near extreme base and spinule-like ventral setulae on basal half; 1 anterior subapical seta. Mid tibia with 4–5 anterodorsal and 4–5 posterodorsal long setae; stronger setulae closer to apex ventrally. Mid tarsus with anteroventral and posteroventral, spine-like setae. Hind femur only slightly broader than mid femur; with 2 anterodorsal setae near apex, 1 anterior subapical seta; complete row of moderately long strong anteroventral setae (longest setae at most half as long as femur width); row of short posteroventral setae intermixed with additional fine setae; dense ventral spinule-like setulae. Hind tibia slightly, gently thickened towards apex; bearing rows of 4–5 moderately long anterodorsal and posterodorsal setae; simple setulae ventrally; no seta in posteroapical comb. Hind tarsomeres rather slightly thickened; basitarsus with 2–3 short anterodorsal and posterodorsal setae; spine-like setae ventrally.</paragraph>
|
||
<paragraph id="DA6A36CBBB07FFD6015A1A3F65FE681A" blockId="1.[812,1399,315,2071]" pageId="1" pageNumber="222">
|
||
Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins mostly brownish, well-sclerotized (except noted). Veins R
|
||
<subScript id="4651348EBB07FFD606491AFB650E69AC" attach="left" box="[1089,1096,1717,1729]" fontSize="5" pageId="1" pageNumber="222">5</subScript>
|
||
and M
|
||
<subScript id="4651348EBB07FFD606821AFB65D769AC" attach="left" box="[1162,1169,1717,1729]" fontSize="5" pageId="1" pageNumber="222">1</subScript>
|
||
divergent towards wing margin; radial fork acute; R
|
||
<subScript id="4651348EBB07FFD6063D1A9E657A69B1" attach="left" box="[1077,1084,1744,1756]" fontSize="5" pageId="1" pageNumber="222">5</subScript>
|
||
meeting costa before wing apex; CuA+CuP (anal vein) complete but weakened beyond middle. Cell dm rather short, with elongate apex; apical portion of M
|
||
<subScript id="4651348EBB07FFD601531B6E62246841" attach="left" box="[859,866,1824,1836]" fontSize="5" pageId="1" pageNumber="222">4</subScript>
|
||
nearly 3.5X longer than its middle portion. Pterostigma absent. Basal costal seta present, short. Anal lobe well-developed; axillary incision acute but closer to 90°. Squama yellow, pale fringed. Halter pale yellow.
|
||
</paragraph>
|
||
<paragraph id="DA6A36CBBB07FFD5015A1B3560B96DFC" blockId="1.[812,1399,315,2071]" lastBlockId="2.[189,775,315,791]" lastPageId="2" lastPageNumber="223" pageId="1" pageNumber="222">Abdomen entirely dark-brown. Tergite 1 entirely greyish pruinose; tergites 2–6 mostly shiny, narrowly brown pruinose antero-dorsally (dorsal view); tergite 7 with brownish, slightly iridescent pruinosity dorsally and greyish pruinose laterally; tergite 8 invisible; tergite 1 with pale setae, tergites 2–6 with black setulae dorsally and numerous long pale setae laterally, slightly stronger lateral posteromarginal setae pale yellow; tergite 7 with scattered pale setulae laterally. Sternites 1–7 densely greyish pruinose, with pale setae, sternite 1 bare; sternite 8 contrastingly subshiny, mostly with black setae. Pregenital segments: segment 6 unmodified; tergite 7 with straight posterior margin, slightly humped mid-dorsally (lateral view); sternite 7 simple; pleuron 7 unmodified; segment 8 with separated tergite and sternite; tergite 8 represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 scoop-shaped, unmodified anteriorly, with 2 digitiform, short, lateral projections on each side closer to upper margin anteriorly (posterior projection slightly longer).</paragraph>
|
||
<paragraph id="DA6A36CBBB04FFD502EA1ED864316DAA" blockId="2.[189,775,315,791]" lastBlockId="2.[812,1399,315,791]" pageId="2" pageNumber="223">
|
||
Hypopygium (
|
||
<figureCitation id="42EE2A4EBB04FFD503661ED860E06DC6" box="[366,422,662,684]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="2" pageNumber="223">Fig. 2</figureCitation>
|
||
) moderately large; epandrium black; cerci pale brown to yellowish brown; hypandrium invisible; phallus brownish yellow. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella subtriangular (lateral view), with broadly rounded apex; bearing dark fine setae more numerous along lower margin and dense at apex. Hypandrium separated from epandrium; mostly membranous, narrowly sclerotized along margin, with truncate apex, narrowly separated on apex; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus egg-shaped, with posterior margin extending beyond apex of epandrial lamella, undivided, with smoothed inner margin; covered with dark setulae. Phallus (
|
||
<figureCitation id="42EE2A4EBB04FFD506E01DB864636D66" box="[1256,1317,502,524]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="2" pageNumber="223">Fig. 3</figureCitation>
|
||
) almost entirely hidden, only medial curvature and, often, short apical portion visible; zigzag bent; its basal portion almost horizontal, mostly thick, with mid-dorsal tubercle; apical portion long, almost vertical and straight, slightly curved on apex (with its tip pointing almost to rear), becoming gently slenderer towards apex. Ejaculatory apodeme very large, extended far beyond basal curvature of phallus, with lateral wings.
|
||
</paragraph>
|
||
<paragraph id="DA6A36CBBB04FFD4015A1E8563406C5C" blockId="2.[812,1399,315,791]" lastBlockId="3.[189,776,315,1271]" lastPageId="3" lastPageNumber="224" pageId="2" pageNumber="223">
|
||
<emphasis id="E8A1EAD9BB04FFD5015A1E8562DB6D8D" bold="true" box="[850,925,715,736]" pageId="2" pageNumber="223">Female</emphasis>
|
||
(
|
||
<figureCitation id="42EE2A4EBB04FFD501B91E8562B56D8C" box="[945,1011,715,737]" captionStart="Figs 1–5" captionStartId="2.[226,263,1918,1937]" captionTargetBox="[188,1393,818,1902]" captionTargetId="figure-436@2.[188,1398,809,1902]" captionTargetPageId="2" captionText="Figs 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm. Рис. 1–5. Empis (Polyblepharis) fandariensis Shamshev, sp.n.: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм." figureDoi="http://doi.org/10.5281/zenodo.13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="2" pageNumber="223">Fig. 5</figureCitation>
|
||
). Similar to male except as follows. Dichoptic, ommatidia equally small.Frons very broad, broader than distance between outer margins of posterior ocelli; with marginal setulae. Occiput with sparser and stronger setae. Palpus yellow, with scattered short setae. Mesoscutum with shorter acrostichal and dorsocentral setae; presutural supra-alar space only with black fine setae. Legs colour: mid and hind femora narrowly yellow at apex; mid and hind tibiae rather brownish yellow (the former generally slightly darker). Hind trochanter only with pale fine setae ventrally. Fore leg with simple setae (similar pattern but somewhat shorter than in male). Mid femur with short, fine anteroventral setae longer closer to base; moderately long, flattened setae on about apical 2/3 posteroventrally. Mid tibia with 2–3 anteroventral and 1–2 posteroventral setae. Hind femur with fridge of short, dense, slightly flattened setae posteroventrally (except extreme base). Hind tibia with 1–2 anteroventral setae. Abdomen with shorter setae, only pale setose; tergites mostly shiny, narrowly greyish pruinose along lateral margin (in addition, tergite 2 narrowly pollinose anteriorly; tergite 1 entirely pruinose); sternite 8 subshiny. Cercus very short, covered with dense, erect setulae.
|
||
</paragraph>
|
||
<caption id="8EAA6643BB04FFD502EA1B3063AC6760" ID-DOI="http://doi.org/10.5281/zenodo.13178587" ID-Zenodo-Dep="13178587" httpUri="https://zenodo.org/record/13178587/files/figure.png" pageId="2" pageNumber="223" startId="2.[226,263,1918,1937]" targetBox="[188,1393,818,1902]" targetPageId="2" targetType="figure">
|
||
<paragraph id="DA6A36CBBB04FFD502EA1B30625468AC" blockId="2.[189,1398,1917,2061]" pageId="2" pageNumber="223">
|
||
Figs 1–5.
|
||
<taxonomicName id="1DD54D48BB04FFD503351B30639068FC" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[317,726,1918,1937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
|
||
<emphasis id="E8A1EAD9BB04FFD503351B30603468FD" box="[317,370,1918,1936]" italics="true" pageId="2" pageNumber="223">Empis</emphasis>
|
||
(
|
||
<emphasis id="E8A1EAD9BB04FFD5038A1B3060BD68FC" box="[386,507,1918,1937]" italics="true" pageId="2" pageNumber="223">Polyblepharis</emphasis>
|
||
)
|
||
<emphasis id="E8A1EAD9BB04FFD500021B30633268FC" box="[522,628,1918,1937]" italics="true" pageId="2" pageNumber="223">fandariensis</emphasis>
|
||
Shamshev
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E8A1EAD9BB04FFD500EA1B30624B68FC" bold="true" box="[738,781,1918,1937]" pageId="2" pageNumber="223">
|
||
<taxonomicNameLabel id="F39257A2BB04FFD500EA1B30624B68FC" box="[738,781,1918,1937]" pageId="2" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 1 — male, holotype, habitus; 2 — hypopygium; 3 — phallus; 4 — holotype labels; 5 — female, habitus; 1–3, 5 — lateral view. Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; hypd — hypandrium; ph — phallus. Scale bar is 1 mm.
|
||
</paragraph>
|
||
<paragraph id="DA6A36CBBB04FFD502EA1B8463AC6760" blockId="2.[189,1398,1917,2061]" pageId="2" pageNumber="223">
|
||
Рис. 1–5.
|
||
<taxonomicName id="1DD54D48BB04FFD503351B84639368B1" authority="Shamshev" authorityName="Shamshev" authorityYear="2023" box="[317,725,1993,2012]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov." subGenus="Polyblepharis">
|
||
<emphasis id="E8A1EAD9BB04FFD503351B84603468B1" box="[317,370,1994,2012]" italics="true" pageId="2" pageNumber="223">Empis</emphasis>
|
||
(
|
||
<emphasis id="E8A1EAD9BB04FFD503891B8760BC68B1" box="[385,506,1993,2012]" italics="true" pageId="2" pageNumber="223">Polyblepharis</emphasis>
|
||
)
|
||
<emphasis id="E8A1EAD9BB04FFD500011B87633568B1" box="[521,627,1993,2012]" italics="true" pageId="2" pageNumber="223">fandariensis</emphasis>
|
||
Shamshev
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E8A1EAD9BB04FFD500D71B87624C68B1" bold="true" box="[735,778,1993,2012]" pageId="2" pageNumber="223">
|
||
<taxonomicNameLabel id="F39257A2BB04FFD500D71B87624C68B1" box="[735,778,1993,2012]" pageId="2" pageNumber="223" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 1 — самец, голотип, габитус; 2 — гипопигий; 3 — фаллус; 4 — Этикетки голотипа; 5 — самка, габитус; 1–3, 5 — сбоку. СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; hypd — гипандрий; ph — фаллус. Шкала = 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA6A36CBBB05FFD402EA1F7863F86BCA" blockId="3.[189,776,315,1271]" pageId="3" pageNumber="224">
|
||
DIFFERENTIAL DIAGNOSIS. The new species is very similar to
|
||
<taxonomicName id="1DD54D48BB05FFD403141F1F631E6C0B" authority="Chvala, 1999" authorityName="Chvala" authorityYear="1999" box="[284,600,849,871]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
|
||
<emphasis id="E8A1EAD9BB05FFD403141F1F60976C0B" box="[284,465,849,870]" italics="true" pageId="3" pageNumber="224">Empis tajikistanica</emphasis>
|
||
<bibRefCitation id="BE444B3ABB05FFD403DF1F1F631E6C0B" author="Chvala M." box="[471,600,849,871]" pageId="3" pageNumber="224" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="1DD54D48BB05FFD4006A1F1F61906CEC" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="fandariensis" status="sp. nov.">
|
||
<emphasis id="E8A1EAD9BB05FFD4006A1F1F61906CEC" italics="true" pageId="3" pageNumber="224">Empis fandariensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8A1EAD9BB05FFD402E91F2560566CED" bold="true" box="[225,272,875,896]" pageId="3" pageNumber="224">
|
||
<taxonomicNameLabel id="F39257A2BB05FFD402E91F2560566CED" box="[225,272,875,896]" pageId="3" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="1DD54D48BB05FFD403951F2263766CED" authorityName="Chvala" authorityYear="1999" box="[413,560,875,897]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
|
||
<emphasis id="E8A1EAD9BB05FFD403951F2260EA6CEC" box="[413,428,876,897]" italics="true" pageId="3" pageNumber="224">E</emphasis>
|
||
.
|
||
<emphasis id="E8A1EAD9BB05FFD403B41F2563766CED" box="[444,560,875,896]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
|
||
</taxonomicName>
|
||
by shorter proboscis (labrum nearly 2X longer than head height) and extensively shiny abdominal tergites 2–6 (lateral view). In addition, the male of the new species has sparser and finer subapical, ventral setae on hind trochanter and rounded cerci. In
|
||
<taxonomicName id="1DD54D48BB05FFD4007F1F9962406C86" authorityName="Chvala" authorityYear="1999" box="[631,774,982,1004]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
|
||
<emphasis id="E8A1EAD9BB05FFD4007F1F9963C36C81" box="[631,645,983,1004]" italics="true" pageId="3" pageNumber="224">E</emphasis>
|
||
.
|
||
<emphasis id="E8A1EAD9BB05FFD4009A1F9862406C86" box="[658,774,982,1003]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
|
||
</taxonomicName>
|
||
the labrum of proboscis is nearly 3X longer than head height and abdominal tergites 2–6 extensively greyish pruinose (lateral view). The male of
|
||
<taxonomicName id="1DD54D48BB05FFD403AE1869637E6B56" authorityName="Chvala" authorityYear="1999" box="[422,568,1062,1084]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
|
||
<emphasis id="E8A1EAD9BB05FFD403AE186960F36B51" box="[422,437,1063,1084]" italics="true" pageId="3" pageNumber="224">E</emphasis>
|
||
.
|
||
<emphasis id="E8A1EAD9BB05FFD403CC1868637E6B56" box="[452,568,1062,1083]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
|
||
</taxonomicName>
|
||
has a tuft of strong, black, ventral setae on the hind trochanter and somewhat pointed cerci [
|
||
<bibRefCitation id="BE444B3ABB05FFD4034F181560B96B1D" author="Chvala M." box="[327,511,1115,1137]" pageId="3" pageNumber="224" pagination="113 - 225" refId="ref12779" refString="Chvala M. 1999. Revision of Palaearctic species of the Empis subgenus Polyblepharis (Diptera, Empididae), with descriptions of fourteen new species // Acta Universitatis Carolinae Biologica. Vol. 42. P. 113 - 225." type="journal article" year="1999">Chvála, 1999: 210</bibRefCitation>
|
||
, fig. 28]. The female of
|
||
<taxonomicName id="1DD54D48BB05FFD400FA181260746BE6" authorityName="Shamshev" authorityYear="2023" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="fandariensis" status="sp.n.">
|
||
<emphasis id="E8A1EAD9BB05FFD400FA181262466B1C" box="[754,768,1116,1137]" italics="true" pageId="3" pageNumber="224">E</emphasis>
|
||
.
|
||
<emphasis id="E8A1EAD9BB05FFD402B5183860746BE6" box="[189,306,1142,1163]" italics="true" pageId="3" pageNumber="224">fandariensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8A1EAD9BB05FFD403311838602F6BE6" bold="true" box="[313,361,1142,1163]" pageId="3" pageNumber="224">
|
||
<taxonomicNameLabel id="F39257A2BB05FFD403311838602F6BE6" box="[313,361,1142,1163]" pageId="3" pageNumber="224" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the female of
|
||
<taxonomicName id="1DD54D48BB05FFD4007D183962406BE6" authorityName="Chvala" authorityYear="1999" box="[629,774,1142,1164]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="tajikistanica">
|
||
<emphasis id="E8A1EAD9BB05FFD4007D183963C26BE1" box="[629,644,1143,1164]" italics="true" pageId="3" pageNumber="224">E</emphasis>
|
||
.
|
||
<emphasis id="E8A1EAD9BB05FFD4009A183862406BE6" box="[658,774,1142,1163]" italics="true" pageId="3" pageNumber="224">tajikistanica</emphasis>
|
||
</taxonomicName>
|
||
primarily by somewhat pennate mid and hind femora.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92CF6540BB05FFD402EA18E562466BB1" pageId="3" pageNumber="224" type="etymology">
|
||
<paragraph id="DA6A36CBBB05FFD402EA18E562466BB1" blockId="3.[189,776,315,1271]" pageId="3" pageNumber="224">
|
||
<materialsCitation id="6ABD3C96BB05FFD402EA18E563BB6BB1" country="Tajikistan" location="Fan Darya River" pageId="3" pageNumber="224" specimenCount="1">
|
||
ETYMOLOGY. The epithet refers to the
|
||
<typeStatus id="056E8869BB05FFD4007C18E263D86BAC" box="[628,670,1196,1217]" pageId="3" pageNumber="224">type</typeStatus>
|
||
locality of the new species, an area near
|
||
<location id="DF0A6010BB05FFD403D7188863C46BB1" LSID="urn:lsid:plazi:treatment:527C87DDBB07FFD402CE184663446B9A:DF0A6010BB05FFD403D7188863C46BB1" box="[479,642,1222,1244]" country="Tajikistan" name="Fan Darya River" pageId="3" pageNumber="224">Fan Darya River</location>
|
||
(
|
||
<collectingCountry id="A2C2765BBB05FFD40087188863B06BB1" box="[655,758,1222,1244]" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92CF6540BB05FFD402EA18AF63446B9A" box="[226,514,1249,1271]" pageId="3" pageNumber="224" type="distribution">
|
||
<paragraph id="DA6A36CBBB05FFD402EA18AF63446B9A" blockId="3.[189,776,315,1271]" box="[226,514,1249,1271]" pageId="3" pageNumber="224">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="A2C2765BBB05FFD4039218AF60B86B9A" box="[410,510,1249,1271]" name="Tajikistan" pageId="3" pageNumber="224">Tajikistan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |