209 lines
18 KiB
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209 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="526CABE7441F88169A4B4761AB67F971" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus denticulatus Seeley 1870" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="37" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="37" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044998" LSID="urn:lsid:plazi:treatment:526CABE7441F88169A4B4761AB67F971" httpUri="http://treatment.plazi.org/id/526CABE7441F88169A4B4761AB67F971" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection pageId="36" pageNumber="37" type="nomenclature">
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<paragraph pageId="36" pageNumber="37">
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<taxonomicName LSID="526CABE7-441F-8816-9A4B-4761AB67F971" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus denticulatus" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="denticulatus">
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<pageBreakToken pageId="36" pageNumber="37" start="start">'</pageBreakToken>
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Ornithocheirus ' denticulatus Seeley, 1870
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</taxonomicName>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-D ' Pterodactylus' daviesii, holotype NHMUK PV 43074 (Albian, Gault Clay Formation), anterior part of the mandibular symphysis. A right lateral view B respective line drawing C dorsal view D respective line drawing. E-H ' Ornithocheirus' denticulatus, holotype? CAMSM B 54794 (Albian, Cambridge Greensand), anterior part of the rostrum E ventral view F respective line drawing G right lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. E and G from Seeley 1870. Photos A and C courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22081" pageId="36" pageNumber="37">Fig. 13E-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="reference_group">
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<paragraph pageId="36" pageNumber="37">
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<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="denticulatus">Ornithocheirus denticulatus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 122, pl. XII, fig. 8-9
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</paragraph>
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<paragraph pageId="36" pageNumber="37">
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<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="denticulatus">Ornithocheirus denticulatus</taxonomicName>
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Seeley:Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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<paragraph pageId="36" pageNumber="37">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cuvieri">Anhanguera cuvieri</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="holotype">
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<paragraph pageId="36" pageNumber="37">Holotype.</paragraph>
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<paragraph pageId="36" pageNumber="37">
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Anterior portion of the rostrum (?CAMSM B 54794) (
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-D ' Pterodactylus' daviesii, holotype NHMUK PV 43074 (Albian, Gault Clay Formation), anterior part of the mandibular symphysis. A right lateral view B respective line drawing C dorsal view D respective line drawing. E-H ' Ornithocheirus' denticulatus, holotype? CAMSM B 54794 (Albian, Cambridge Greensand), anterior part of the rostrum E ventral view F respective line drawing G right lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. E and G from Seeley 1870. Photos A and C courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22081" pageId="36" pageNumber="37">Fig. 13E-H</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="type locality">
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<paragraph pageId="36" pageNumber="37">Type locality.</paragraph>
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<paragraph pageId="36" pageNumber="37">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="type horizon">
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<paragraph pageId="36" pageNumber="37">Type horizon.</paragraph>
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<paragraph pageId="36" pageNumber="37">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="diagnosis">
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<paragraph pageId="36" pageNumber="37">Diagnosis.</paragraph>
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<paragraph pageId="36" pageNumber="37">Pterodactyloid pterosaur with the following combination of characters that distinguishes it from other members of the clade: anterior expansion absent; palate dorsally curved; moderate palatal ridge; palatal ridge extending anteriorly up to the third pair of alveoli; spacing between alveoli approximately equal to their diameters; approximately 4.5 alveoli per 3 cm of jaw margin.</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="description">
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<paragraph pageId="36" pageNumber="37">Description.</paragraph>
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<paragraph pageId="36" pageNumber="37">
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In the original description,
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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contrasted this species with
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="polyodon">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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polyodon
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</emphasis>
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</taxonomicName>
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(see below).
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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shares with the latter the absence of an anterior expansion, the dorsally curved palate, and moderate size of the palatal ridge. However, in
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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the palatal ridge extends up to the third pair of alveoli, the spacing between the alveoli is larger, and the alveolar density is lower (4.5 alveoli per 3 cm). Because the dorsal margin of the premaxillae is broken, it cannot be access if
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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had a crest.
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</paragraph>
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<paragraph pageId="36" pageNumber="37">
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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does not share the combination of characters present in
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<taxonomicName authorityName="Seeley" authorityYear="1869" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="37">Ornithocheirus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="37">Lonchodraco</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="37">Cimoliopterus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" genus="Camposipterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="36" pageNumber="37">Camposipterus</emphasis>
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</taxonomicName>
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and thus cannot be referred to any of these genera. In particular,
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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lacks the alveolar pattern present in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cuvieri">
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<emphasis italics="true" pageId="36" pageNumber="37">Cimoliopterus cuvieri</emphasis>
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</taxonomicName>
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.
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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somewhat resembles
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="daviesii">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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in that both lack an anterior expansion of the jaw, sagittal crests, and have alveoli without significant variation in size and equally spaced. They differ in the alveolar density, which is slightly higher in
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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. Unfortunately, the known material of both species is not directly comparable and their taxonomic identity cannot be confirmed.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="remarks">
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<paragraph pageId="36" pageNumber="37">Remarks.</paragraph>
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<paragraph pageId="36" pageNumber="37">
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Collection data provided by the curators at the Sedgwick Museum of Earth Sciences in Cambridge list the specimen CAMSM B 54794 as the holotype of
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<taxonomicName lsidName="" pageId="36" pageNumber="37" rank="species" species="denticulatus">
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<emphasis italics="true" pageId="36" pageNumber="37">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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denticulatus
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</emphasis>
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</taxonomicName>
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. However, the specimen could not be found during review of the collection in October 2009. The specimen was, however, one of the few figured by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870</bibRefCitation>
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: pl. XII, figs 8 and 9).
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</paragraph>
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<paragraph pageId="36" pageNumber="37">In the lack of more complete material, we here refrain from naming a new genus based on the present specimen, and refer to it using the binomen in which it was originally proposed.</paragraph>
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</subSubSection>
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</treatment>
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</document> |