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<document id="3CB6883BC70FD6F5C41328BCEF1AE7D1" ID-DOI="10.5281/zenodo.181357" ID-GBIF-Dataset="7b6b3cdb-b35e-4c7a-a7a8-8e1925b61fec" ID-ISSN="1175-5326" ID-Zenodo-Dep="181357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832480416" checkinUser="plazi" docAuthor="Liang, Ai-Ping &amp; Wang, Rong-Rong" docDate="2008" docId="03E0878CFF9E3D4DFF5EF9BDBDAC632B" docLanguage="en" docName="zt01732p044.pdf" docOrigin="Zootaxa 1732" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Sogana clara Liang &amp; Wang, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FFD9FFF4FF9A3D4BFFC9FF8FB951676B" masterDocTitle="A review of the Oriental planthopper genus Sogana Matsumura 1914 (Hemiptera: Fulgoromorpha: Tropiduchidae) with descriptions of three new species" masterLastPageNumber="44" masterPageNumber="29" pageNumber="33" updateTime="1698223042257" updateUser="plazi">
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<mods:title id="61CEB5377199ECA1B745C31E8290776B">A review of the Oriental planthopper genus Sogana Matsumura 1914 (Hemiptera: Fulgoromorpha: Tropiduchidae) with descriptions of three new species</mods:title>
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<mods:namePart id="CF74729AC8A66818C90E7A1FF77B76D6">Liang, Ai-Ping</mods:namePart>
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<mods:namePart id="461BBD66BC70DBD79B28FF368C6B86A8">Wang, Rong-Rong</mods:namePart>
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<mods:date id="68924B4E018D5BB83FEE10BDD7F372C2">2008</mods:date>
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<treatment id="03E0878CFF9E3D4DFF5EF9BDBDAC632B" ID-DOI="http://doi.org/10.5281/zenodo.6231108" ID-GBIF-Taxon="119358132" ID-Zenodo-Dep="6231108" LSID="urn:lsid:plazi:treatment:03E0878CFF9E3D4DFF5EF9BDBDAC632B" httpUri="http://treatment.plazi.org/id/03E0878CFF9E3D4DFF5EF9BDBDAC632B" lastPageId="6" lastPageNumber="35" pageId="4" pageNumber="33">
<subSubSection id="C3536511FF9E3D4FFF5EF9BDBB6D6104" pageId="4" pageNumber="33" type="nomenclature">
<paragraph id="8BF6369AFF9E3D4FFF5EF9BDB8CD6127" blockId="4.[151,572,1586,1647]" box="[151,412,1586,1612]" pageId="4" pageNumber="33">
<heading id="D0BE81F6FF9E3D4FFF5EF9BDB8CD6127" bold="true" box="[151,412,1586,1612]" fontSize="11" level="1" pageId="4" pageNumber="33" reason="1">
<emphasis id="B93DEA88FF9E3D4FFF5EF9BDB8CD6127" bold="true" box="[151,412,1586,1612]" pageId="4" pageNumber="33">
<taxonomicName id="4C494D19FF9E3D4FFF5EF9BDB8666127" ID-CoL="4XYXS" box="[151,311,1586,1612]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
<emphasis id="B93DEA88FF9E3D4FFF5EF9BDB8666127" bold="true" box="[151,311,1586,1612]" italics="true" pageId="4" pageNumber="33">Sogana clara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57F3FF9E3D4FFEF7F9BDB8CD6127" box="[318,412,1586,1612]" pageId="4" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF6369AFF9E3D4FFF5EF9DABB6D6104" blockId="4.[151,572,1586,1647]" box="[151,572,1621,1647]" pageId="4" pageNumber="33">
(
<figureCitation id="13722A1FFF9E3D4FFF56F9DAB9A06104" box="[159,241,1621,1647]" captionStart="FIGURES 1 6" captionStartId="3.[151,269,1913,1937]" captionTargetBox="[193,1391,437,1869]" captionTargetId="figure@3.[151,1436,403,1875]" captionTargetPageId="3" captionText="FIGURES 1 6. Dorsal habitus of Sogana species. 1. S. clara sp. nov. (♂, Laos, BPBM). 2. S. hopponis Matsumura (♂, Taiwan, BPBM). 3. Same (♂, Taiwan, BPBM). 4. S. longiceps Fennah (Ψ, Guangxi, IZCAS). 5. S. pseudohopponis sp. nov. (♂, Laos, BPBM). 6. S. robustocarina sp. nov. (♂, Borneo, BPBM)." httpUri="https://zenodo.org/record/181358/files/figure.png" pageId="4" pageNumber="33">Figs. 1</figureCitation>
,
<figureCitation id="13722A1FFF9E3D4FFF36F9DAB87F6104" box="[255,302,1621,1647]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="4" pageNumber="33">79</figureCitation>
,
<figureCitation id="13722A1FFF9E3D4FFEF3F9DAB8086104" box="[314,345,1621,1647]" captionStart="FIGURES 16 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 23. Sogana species. 16 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="4" pageNumber="33">19</figureCitation>
,
<figureCitation id="13722A1FFF9E3D4FFEACF9DAB88C6104" box="[357,477,1621,1647]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="4" pageNumber="33">24, 2831</figureCitation>
,
<figureCitation id="13722A1FFF9E3D4FFE20F9DABB586104" box="[489,521,1621,1647]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="4" pageNumber="33">44</figureCitation>
, 45)
</paragraph>
</subSubSection>
<subSubSection id="C3536511FF9E3D4DFF5EF92ABA2D650E" lastPageId="6" lastPageNumber="35" pageId="4" pageNumber="33" type="description">
<paragraph id="8BF6369AFF9E3D4FFF5EF92ABDE761D4" blockId="4.[151,1436,1700,2007]" box="[151,1206,1700,1727]" pageId="4" pageNumber="33">
<emphasis id="B93DEA88FF9E3D4FFF5EF92AB87E61D4" bold="true" box="[151,303,1701,1727]" pageId="4" pageNumber="33">Description.</emphasis>
Length (from apex of vertex to tip of fore wings): ď 8.38.5 mm, Ψ 9.3 mm.
</paragraph>
<paragraph id="8BF6369AFF9E3D4FFF0CF942B82560BC" blockId="4.[151,1436,1700,2007]" pageId="4" pageNumber="33">
General color light yellowish. Vertex (
<figureCitation id="13722A1FFF9E3D4FFD4FF942BB9D618C" box="[646,716,1741,1767]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="4" pageNumber="33">Fig. 7</figureCitation>
) with three black spots at apex, the median one roundish; lateral areas suffused with fuscous, central disc with a pair of brownish spots near posterior margin; median carina yellowish-brown, sublateral and lateral carinae orange, lateral margins brownish. Frons (
<figureCitation id="13722A1FFF9E3D4FFB3BF892BC6A605C" box="[1266,1339,1821,1847]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="4" pageNumber="33">Fig. 8</figureCitation>
) with 5 reddish transverse bands. Gena (
<figureCitation id="13722A1FFF9E3D4FFDEFF8CABB206034" box="[550,625,1861,1887]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="4" pageNumber="33">Fig. 9</figureCitation>
) with one brown patch. Carinae on frons, clypeus, pronotum and mesonotum orange to yellowish-brown. Fore wings (
<figureCitation id="13722A1FFF9E3D4FFCD2F8E2BA2260EC" box="[795,883,1901,1927]" captionStart="FIGURES 16 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 23. Sogana species. 16 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="4" pageNumber="33">Fig. 19</figureCitation>
) pale yellowish; veins yellowish, apical cells with blackish suffusion. Legs pale yellowish; fore femora, long stripes on hind femora, and apices and bases of tibiae brownish.
</paragraph>
<caption id="DF366612FF9F3D4EFF5EF94CBB6D604B" httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="5" pageNumber="34" targetBox="[194,1421,185,1713]" targetPageId="5">
<paragraph id="8BF6369AFF9F3D4EFF5EF94CBB6D604B" blockId="5.[151,1436,1731,1825]" pageId="5" pageNumber="34">
<emphasis id="B93DEA88FF9F3D4EFF5EF94CB80361B0" bold="true" box="[151,338,1731,1755]" pageId="5" pageNumber="34">FIGURES 715.</emphasis>
<taxonomicName id="4C494D19FF9F3D4EFE93F94BB8FB61B0" box="[346,426,1732,1755]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA88FF9F3D4EFE93F94BB8FB61B0" box="[346,426,1732,1755]" italics="true" pageId="5" pageNumber="34">Sogana</emphasis>
</taxonomicName>
species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 79.
<taxonomicName id="4C494D19FF9F3D4EFDE7F968BBD46195" box="[558,645,1767,1790]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
<emphasis id="B93DEA88FF9F3D4EFDE7F968BBD46195" box="[558,645,1767,1790]" italics="true" pageId="5" pageNumber="34">S. clara</emphasis>
</taxonomicName>
<emphasis id="B93DEA88FF9F3D4EFD58F969BBBA6195" bold="true" box="[657,747,1766,1790]" pageId="5" pageNumber="34">
<taxonomicNameLabel id="A20E57F3FF9F3D4EFD58F969BBBA6195" box="[657,747,1766,1790]" pageId="5" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂, holotype). 1012.
<taxonomicName id="4C494D19FF9F3D4EFC27F968BDA26196" authority="Matsumura" authorityName="Matsumura" box="[1006,1267,1766,1790]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="hopponis">
<emphasis id="B93DEA88FF9F3D4EFC27F968BD3E6195" box="[1006,1135,1767,1790]" italics="true" pageId="5" pageNumber="34">S. hopponis</emphasis>
Matsumura
</taxonomicName>
(♂). 1315.
<taxonomicName id="4C494D19FF9F3D4EFA41F968B814604A" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="pseudohopponis" status="sp. nov.">
<emphasis id="B93DEA88FF9F3D4EFA41F968B814604A" italics="true" pageId="5" pageNumber="34">S. pseudohopponis</emphasis>
</taxonomicName>
<emphasis id="B93DEA88FF9F3D4EFE82F886B8F1604A" bold="true" box="[331,416,1801,1825]" pageId="5" pageNumber="34">
<taxonomicNameLabel id="A20E57F3FF9F3D4EFE82F886B8F1604A" box="[331,416,1801,1825]" pageId="5" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂, holotype).
</paragraph>
</caption>
<paragraph id="8BF6369AFF9F3D4EFF0CF8DBBB4C608D" blockId="5.[151,1436,1876,2022]" pageId="5" pageNumber="34">
Vertex with anterior margin projected at an obtuse angle in dorsal view, lateral areas distinctly depressed; median carina nearly percurrent. Frons with median carina thick and wide. Fore wings (
<figureCitation id="13722A1FFF9F3D4EFB5AF8F3BDB660FD" box="[1171,1255,1916,1942]" captionStart="FIGURES 16 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 23. Sogana species. 16 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="5" pageNumber="34">Fig. 19</figureCitation>
) with 14 apical cells and 7 subapical cells, two middle anteapical cells equally long. Spines of hind legs: hind tibia laterally 3, distally 7, basitarsus distally 89.
</paragraph>
<paragraph id="8BF6369AFF9C3D4DFF0CFF17B8B26643" blockId="6.[151,1437,152,1092]" pageId="6" pageNumber="35">
<emphasis id="B93DEA88FF9C3D4DFF0CFF17B80B67DA" box="[197,346,152,177]" italics="true" pageId="6" pageNumber="35">Proportions.</emphasis>
Vertex with ratio of median length to the widest breadth 0.82:1, frons with ratio of median length to the widest breadth 1.49:1, pronotum with ratio of width to median length 4.93:1, pronotum and mesonotum together medially about 2 times as long as median length of vertex, fore wings with ratio of length to maximum breadth 3.49:1.
</paragraph>
<paragraph id="8BF6369AFF9C3D4DFF0CFEB9BA2D650E" blockId="6.[151,1437,152,1092]" pageId="6" pageNumber="35">
Male genitalia (
<figureCitation id="13722A1FFF9C3D4DFEB7FEB9BB68663B" box="[382,569,310,336]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Figs. 24, 2831</figureCitation>
,
<figureCitation id="13722A1FFF9C3D4DFD8CFEB9BBC1663B" box="[581,656,310,336]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="6" pageNumber="35">44, 45</figureCitation>
) with pygofer relatively broad and short, posterior margin broadly caudad expanded subapically in lateral view (
<figureCitation id="13722A1FFF9C3D4DFD0AFED1BA4F6613" box="[707,798,350,376]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Fig. 24</figureCitation>
). Anal tube (
<figureCitation id="13722A1FFF9C3D4DFC08FED1BD1B6613" box="[961,1098,350,376]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Fig. 24, 28</figureCitation>
) with apical ventral margin slightly bent ventrad in lateral view. Gonostylus (
<figureCitation id="13722A1FFF9C3D4DFD17FE0ABAC866F4" box="[734,921,389,415]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Figs. 24, 30, 31</figureCitation>
) in lateral view about 3.54 times as long as broad. Aedeagus (
<figureCitation id="13722A1FFF9C3D4DFEB9FE22B88466AC" box="[368,469,429,455]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Figs. 29</figureCitation>
,
<figureCitation id="13722A1FFF9C3D4DFE2AFE22BB6166AC" box="[483,560,429,455]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="6" pageNumber="35">44, 45</figureCitation>
) with three processes: two anteriorly directed lamellate processes arising from right side with the longer one spiral-like and produced at apical fifth and the shorter one produced at apical third; and the third posteriorly directed process slender, originating from ventral side; endosomal process digitiform; phallotrema exposed subapically on left side (
<figureCitation id="13722A1FFF9C3D4DFCF0FDABBAC06555" box="[825,913,548,574]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="6" pageNumber="35">Fig. 44</figureCitation>
); periandrium (
<figureCitation id="13722A1FFF9C3D4DFB8FFDABBDF86555" box="[1094,1193,548,574]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Figs. 24</figureCitation>
,
<figureCitation id="13722A1FFF9C3D4DFB7CFDABBC516555" box="[1205,1280,548,574]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="6" pageNumber="35">44, 45</figureCitation>
) with a long, tongue-shaped process protruding backward from ventral apex.
</paragraph>
</subSubSection>
<subSubSection id="C3536511FF9C3D4DFF0FFDFDB9AB646F" pageId="6" pageNumber="35" type="materials_examined">
<paragraph id="8BF6369AFF9C3D4DFF0FFDFDB9AB646F" blockId="6.[151,1437,152,1092]" pageId="6" pageNumber="35">
<emphasis id="B93DEA88FF9C3D4DFF0FFDFDB8ED65E7" bold="true" box="[198,444,626,652]" pageId="6" pageNumber="35">Material examined.</emphasis>
<typeStatus id="54F28838FF9C3D4DFE0DFDFCBB6065E6" box="[452,561,627,653]" pageId="6" pageNumber="35" type="holotype">Holotype</typeStatus>
ď,
<collectingCountry id="F35E760AFF9C3D4DFD94FDFCBBFF65E6" box="[605,686,627,653]" name="Laos" pageId="6" pageNumber="35">LAOS</collectingCountry>
: Vientiane Prov., Ban Van Eue,
<date id="FFF7105AFF9C3D4DFBE3FDFCBD9265E6" box="[1066,1219,627,653]" pageId="6" pageNumber="35" value="1967-08-15">15.VIII.1967</date>
“native collector” (
<collectionCode id="ED58AE5FFF9C3D4DFF56FD15B9A665DF" LSID="urn:lsid:biocol.org:col:34697" box="[159,247,666,692]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="35">BPBM</collectionCode>
).
<typeStatus id="54F28838FF9C3D4DFEC5FD15B8D065DF" box="[268,385,666,692]" pageId="6" pageNumber="35" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F35E760AFF9C3D4DFE44FD15B88F65DF" box="[397,478,666,692]" name="Laos" pageId="6" pageNumber="35">LAOS</collectingCountry>
: 1ď, Vientiane Prov., Ban Van Eue,
<date id="FFF7105AFF9C3D4DFC42FD15BD4065DF" box="[907,1041,666,692]" pageId="6" pageNumber="35" value="1967-09-30">30.IX.1967</date>
“native collector”; 1Ψ, Vientiane Prov., Ban Van Eue, malaise trap,
<date id="FFF7105AFF9C3D4DFDF2FD4DBBE465B7" box="[571,693,706,732]" pageId="6" pageNumber="35" value="1966-02-15">15.II.1966</date>
“native collector”, Rondon-Bishop Mus. Collection) (both in
<collectionCode id="ED58AE5FFF9C3D4DFF5EFD65B9BF646F" LSID="urn:lsid:biocol.org:col:34697" box="[151,238,746,772]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="35">BPBM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536511FF9C3D4DFF0FFC9EBA856440" box="[198,980,785,811]" pageId="6" pageNumber="35" type="etymology">
<paragraph id="8BF6369AFF9C3D4DFF0FFC9EBA856440" blockId="6.[151,1437,152,1092]" box="[198,980,785,811]" pageId="6" pageNumber="35">
<emphasis id="B93DEA88FF9C3D4DFF0FFC9EB8026440" bold="true" box="[198,339,785,811]" pageId="6" pageNumber="35">Etymology.</emphasis>
This new species is named for its general bright color.
</paragraph>
</subSubSection>
<subSubSection id="C3536511FF9C3D4DFF0FFCB7BB6B6438" box="[198,570,824,851]" pageId="6" pageNumber="35" type="distribution">
<paragraph id="8BF6369AFF9C3D4DFF0FFCB7BB6B6438" blockId="6.[151,1437,152,1092]" box="[198,570,824,851]" pageId="6" pageNumber="35">
<emphasis id="B93DEA88FF9C3D4DFF0FFCB7B8366439" bold="true" box="[198,359,824,850]" pageId="6" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="F35E760AFF9C3D4DFEA7FCB6B8F66438" box="[366,423,825,851]" name="Laos" pageId="6" pageNumber="35">Laos</collectingCountry>
(Vientiane).
</paragraph>
</subSubSection>
<subSubSection id="C3536511FF9C3D4DFF0FFCEFBDAC632B" pageId="6" pageNumber="35" type="discussion">
<paragraph id="8BF6369AFF9C3D4DFF0FFCEFBDAC632B" blockId="6.[151,1437,152,1092]" pageId="6" pageNumber="35">
<emphasis id="B93DEA88FF9C3D4DFF0FFCEFB86E6411" bold="true" box="[198,319,864,890]" pageId="6" pageNumber="35">Remarks.</emphasis>
This species can be distinguished from other known species in the genus by its general bright color and the male genitalic structure (
<figureCitation id="13722A1FFF9C3D4DFD94FC07BA4864C9" box="[605,793,904,930]" captionStart="FIGURES 24 35" captionStartId="10.[151,269,1838,1862]" captionTargetBox="[237,1320,193,1812]" captionTargetId="figure@10.[201,1370,156,1825]" captionTargetPageId="10" captionText="FIGURES 24 35. Sogana species. 24 27. male genitalia (lateral view). 24. S. clara sp. nov. (holotype). 25. S. hopponis Matsumura. 26. S. pseudohopponis sp. nov. (holotype). 27. S. robustocarina sp. nov. (holotype). 28 35. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 28 31. S. clara sp. nov. (holotype). 32 35. S. hopponis Matsumura. Abbreviations: ap = anal apodeme, At = anal tube, CC = corpus connective, Epr = epiproct, G = gonostylus, P = pygofer, Ppr = paraproct, STC = structure tectiform of connective." httpUri="https://zenodo.org/record/181361/files/figure.png" pageId="6" pageNumber="35">Figs. 24, 2831</figureCitation>
,
<figureCitation id="13722A1FFF9C3D4DFCEDFC07BA2364C9" box="[804,882,904,930]" captionStart="FIGURES 36 47" captionStartId="13.[151,269,1904,1928]" captionTargetBox="[227,1370,203,1879]" captionTargetId="figure@13.[196,1390,194,1900]" captionTargetPageId="13" captionText="FIGURES 36 47. Sogana species. 36 43. anal segment and pygofer (dorsal view); aedeagus (ventral view); gonostylus (dorsal view); gonostylus and pygofer (ventral view). 36 39. S. pseudohopponis sp. nov. (holotype). 40 43. S. robustocarina sp. nov. (holotype). 44 47. aedeagus (left view); same (right view). 44 45. S. clara sp. nov. (holotype). 46 47. S. hopponis Matsumura." httpUri="https://zenodo.org/record/181362/files/figure.png" pageId="6" pageNumber="35">44, 45</figureCitation>
). In external appearance, this species is similar to
<taxonomicName id="4C494D19FF9C3D4DFF7EFC3FB8A064A1" authority="Melichar, 1914" authorityName="Melichar" authorityYear="1914" box="[183,497,944,970]" class="Insecta" family="Tropiduchidae" genus="Sogana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="extrema">
<emphasis id="B93DEA88FF9C3D4DFF7EFC3FB86264A2" box="[183,307,944,969]" italics="true" pageId="6" pageNumber="35">S. extrema</emphasis>
<bibRefCitation id="EFD84B6BFF9C3D4DFEF5FC3FB8A064A1" author="Melichar" box="[316,497,944,970]" pageId="6" pageNumber="44" refString="Melichar, L. (1914) Monographie der Tropiduchinen (Homoptera). Verhandlungen des Naturforschenden Vereins in Brunn, 53, 1 - 145." type="journal article" year="1914">Melichar, 1914</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F35E760AFF9C3D4DFDF2FC3FBBDA64A1" box="[571,651,944,970]" name="Myanmar" pageId="6" pageNumber="35">Burma</collectingCountry>
(Tenasserim) and
<collectingCountry id="F35E760AFF9C3D4DFCA3FC3FBAE164A1" box="[874,944,944,970]" name="China" pageId="6" pageNumber="35">China</collectingCountry>
(Chekiang), but differs from the latter in the vertex (
<figureCitation id="13722A1FFF9C3D4DFED2FC58B831649A" box="[283,352,983,1009]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="6" pageNumber="35">Fig. 7</figureCitation>
) with anterior margin projected at an obtuse angle in dorsal view and the median black spots at apex roundish, the frons (
<figureCitation id="13722A1FFF9C3D4DFE2DFC70BB7A6372" box="[484,555,1023,1049]" captionStart="FIGURES 7 15" captionStartId="5.[151,269,1731,1755]" captionTargetBox="[194,1421,185,1713]" captionTargetId="figure@5.[182,1426,170,1728]" captionTargetPageId="5" captionText="FIGURES 7 15. Sogana species: head, pronotum and mesonotum (dorsal view); head (ventral view); head, pronotum and mesonotum (lateral view). 7 9. S. clara sp. nov. (♂, holotype). 10 12. S. hopponis Matsumura (♂). 13 15. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181359/files/figure.png" pageId="6" pageNumber="35">Fig. 8</figureCitation>
) with median carina thick and the reddish transverse bands wider, and the fore wings with Cu1 forked more basally and with 14 apical cells and 7 subapical cells (
<figureCitation id="13722A1FFF9C3D4DFB50FBA9BDBE632B" box="[1177,1263,1062,1088]" captionStart="FIGURES 16 23" captionStartId="9.[151,269,1870,1894]" captionTargetBox="[199,1384,220,1835]" captionTargetId="figure@9.[151,1436,194,1846]" captionTargetPageId="9" captionText="FIGURES 16 23. Sogana species. 16 18. S. robustocarina sp. nov. (♂, holotype). 16. head, pronotum and mesonotum (dorsal view); 17. head (ventral view); 18. head, pronotum and mesonotum (lateral view). 19 22. fore wing. 19. S. clara sp. nov. (♂, holotype). 20. S. hopponis Matsumura (♂). 21. S. pseudohopponis sp. nov. (♂, holotype). 22. S. robustocarina sp. nov. (♂, holotype). 23. hind wing. S. pseudohopponis sp. nov. (♂, holotype)." httpUri="https://zenodo.org/record/181360/files/figure.png" pageId="6" pageNumber="35">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>