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<document ID-DOI="10.1080/00222933.2022.2088311" ID-ISSN="1464-5262" ID-Zenodo-Dep="7049416" approvalRequired="8" approvalRequired_for_textStreams="8" checkinTime="1662359225307" checkinUser="plazi" docAuthor="Binoy, C., van Achterberg, Cornelis, Polaszek, Andrew, Kumar, P. Girish &amp; Santhosh, S." docDate="2022" docId="03E07D1EFFFF923CFF0D322EC9F5344A" docLanguage="en" docName="JNATHIST.56.21-24.1153-1185.pdf" docOrigin="Journal of Natural History 56 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2022.2088311" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="1161" masterDocId="FFD90566FFFB9234FFD83168CA783032" masterDocTitle="A review of Taeniogonalos (Hymenoptera: Trigonalyidae) from India with the description of two new species" masterLastPageNumber="1185" masterPageNumber="1153" pageNumber="1157" updateTime="1662360113089" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Taeniogonalos (Hymenoptera: Trigonalyidae) from India with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Binoy, C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Achterberg, Cornelis</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Polaszek, Andrew</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kumar, P. Girish</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santhosh, S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>21 - 24</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1153</mods:start>
<mods:end>1185</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1080/00222933.2022.2088311</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2022.2088311</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">7049416</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03E07D1EFFFF923CFF0D322EC9F5344A" httpUri="http://treatment.plazi.org/id/03E07D1EFFFF923CFF0D322EC9F5344A" lastPageId="8" lastPageNumber="1161" pageId="4" pageNumber="1157">
<subSubSection box="[213,1102,838,864]" pageId="4" pageNumber="1157" type="nomenclature">
<paragraph blockId="4.[213,1102,838,864]" box="[213,1102,838,864]" pageId="4" pageNumber="1157">
<heading bold="true" box="[213,1102,838,864]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="1157" reason="3">
<emphasis bold="true" box="[213,1102,838,864]" pageId="4" pageNumber="1157">
<taxonomicName authority="Binoy, van Achterberg and Girish Kumar" authorityName="Binoy, van Achterberg and Girish Kumar" authorityYear="2022" box="[213,996,838,864]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="1157" phylum="Arthropoda" rank="species" species="ayyari" status="sp. nov.">
<emphasis bold="true" box="[213,478,838,864]" italics="true" pageId="4" pageNumber="1157">Taeniogonalos ayyari</emphasis>
Binoy, van Achterberg and Girish Kumar
</taxonomicName>
<taxonomicNameLabel box="[1004,1102,838,864]" pageId="4" pageNumber="1157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[516,799,883,909]" pageId="4" pageNumber="1157" type="description">
<paragraph blockId="4.[516,799,883,909]" box="[516,799,883,909]" pageId="4" pageNumber="1157">
(
<figureCitation box="[524,673,883,909]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="4" pageNumber="1157">Figures 1ae</figureCitation>
and
<figureCitation box="[734,791,883,909]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="4" pageNumber="1157">2ah</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1157" type="materials_examined">
<paragraph blockId="4.[160,1156,971,1101]" box="[160,324,971,997]" pageId="4" pageNumber="1157">
<emphasis bold="true" box="[160,324,971,997]" italics="true" pageId="4" pageNumber="1157">Type material</emphasis>
</paragraph>
<paragraph blockId="4.[160,1156,971,1101]" pageId="4" pageNumber="1157">
<materialsCitation collectingDate="2019-08-23" collectionCode="ZSIK, WGRC" collectorName="P. Girish Kumar &amp; Regd. No." country="India" elevation="264" latitude="8.682" location="Ambasamudram" longLatPrecision="77" longitude="77.548" municipality="Thirunelveli district" pageId="4" pageNumber="1157" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus box="[160,264,1007,1032]" pageId="4" pageNumber="1157">Holotype</typeStatus>
<specimenCount box="[276,296,1007,1030]" pageId="4" pageNumber="1157" type="female"></specimenCount>
,
<collectingCountry box="[314,373,1007,1032]" name="India" pageId="4" pageNumber="1157">India</collectingCountry>
:
<collectingRegion box="[387,523,1007,1032]" country="India" name="Tamil Nadu" pageId="4" pageNumber="1157">Tamil Nadu</collectingRegion>
,
<collectingMunicipality box="[539,763,1007,1032]" pageId="4" pageNumber="1157">Thirunelveli district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03E07D1EFFFF923CFF0D322EC9F5344A:8E969AD3FFFF9230FCD23287C9BE343A" box="[778,966,1007,1032]" country="India" latitude="8.682" longLatPrecision="77" longitude="77.548" municipality="Thirunelveli district" name="Ambasamudram" pageId="4" pageNumber="1157" stateProvince="Tamil Nadu">Ambasamudram</location>
range (
<geoCoordinate box="[1061,1152,1007,1032]" degrees="8.682" direction="north" orientation="latitude" pageId="4" pageNumber="1157" precision="55" value="8.682">8.682°N</geoCoordinate>
,
<geoCoordinate box="[160,259,1041,1066]" degrees="77.548" direction="east" orientation="longitude" pageId="4" pageNumber="1157" precision="55" value="77.548">77.548°E</geoCoordinate>
; alt.
<quantity box="[311,385,1041,1066]" metricMagnitude="2" metricUnit="m" metricValue="2.6399999999999997" pageId="4" pageNumber="1157" unit="m" value="264.0">
<elevation box="[311,385,1041,1066]" metricMagnitude="2" metricUnit="m" metricValue="2.6399999999999997" pageId="4" pageNumber="1157" unit="m" value="264.0">264 m</elevation>
</quantity>
),
<date box="[404,585,1041,1066]" pageId="4" pageNumber="1157" value="2019-08-23">
<collectingDate box="[404,585,1041,1066]" pageId="4" pageNumber="1157" value="2019-08-23">23 August 2019</collectingDate>
</date>
, coll.
<collectorName box="[650,826,1041,1066]" pageId="4" pageNumber="1157">P. Girish Kumar</collectorName>
,
<emphasis bold="true" box="[837,908,1041,1066]" pageId="4" pageNumber="1157">
(
<collectionCode box="[843,900,1041,1066]" pageId="4" pageNumber="1157">ZSIK</collectionCode>
)
</emphasis>
<collectorName box="[916,1025,1041,1066]" pageId="4" pageNumber="1157">Regd. No.</collectorName>
ZSI/
<collectionCode box="[1075,1148,1041,1066]" pageId="4" pageNumber="1157">WGRC</collectionCode>
/ IR/INV.17487.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1157" type="diagnosis">
<paragraph blockId="4.[160,1155,1145,1414]" box="[160,276,1145,1171]" pageId="4" pageNumber="1157">
<emphasis bold="true" box="[160,276,1145,1171]" italics="true" pageId="4" pageNumber="1157">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="4.[160,1155,1145,1414]" pageId="4" pageNumber="1157">
Scutellum black medially and pale yellow laterally; middle lobe of mesoscutum tricoloured (
<figureCitation box="[167,275,1216,1241]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="4" pageNumber="1157">Figure 2b</figureCitation>
); body markedly setose; outer side of supra-antennal elevations oblique and elevations 0.3× as long as scape; additional pair of oblique elevations at inner margin of eye; occipital carina distinct; head rugose-punctate, area adjoining ocelli smooth, devoid of punctures; interspaces smooth and shiny, punctures sparse on occiput and ventrally on temple; temple dorsally largely smooth except for some punctures; head distinctly wide in dorsal view; posterior margin of second metasomal sternite straight, armature lacking.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="1161" pageId="4" pageNumber="1157" type="description">
<paragraph blockId="4.[160,844,1458,1519]" box="[160,294,1458,1484]" pageId="4" pageNumber="1157">
<emphasis bold="true" box="[160,294,1458,1484]" italics="true" pageId="4" pageNumber="1157">Description</emphasis>
</paragraph>
<paragraph blockId="4.[160,844,1458,1519]" box="[160,844,1494,1519]" pageId="4" pageNumber="1157">
<typeStatus box="[160,265,1494,1519]" pageId="4" pageNumber="1157">Holotype</typeStatus>
, female, length of body
<quantity box="[540,627,1494,1519]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="1157" unit="mm" value="5.6">5.6 mm</quantity>
; fore wing
<quantity box="[753,840,1494,1519]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="1157" unit="mm" value="4.6">4.6 mm</quantity>
.
</paragraph>
<paragraph blockId="4.[160,1156,1564,1728]" lastBlockId="7.[160,1156,142,237]" lastPageId="7" lastPageNumber="1160" pageId="4" pageNumber="1157">
<emphasis bold="true" box="[160,229,1564,1590]" italics="true" pageId="4" pageNumber="1157">Head.</emphasis>
Head in dorsal view 1.6× as wide as long; antenna with 24 antennomeres; frons coarsely rugose punctate, with narrow shiny interspaces; vertex with similar sculpture, moderately setose with medium-sized setae; OOD 2.4× POD, OOD 1.2× OD; temple largely smooth except some sparse setigerous punctation (
<figureCitation box="[765,873,1669,1694]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="4" pageNumber="1157">Figure 1c</figureCitation>
); interocular distance at middle of face 3.5× distance between antennal toruli (
<figureCitation box="[771,877,1703,1728]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="4" pageNumber="1157">Figure 1e</figureCitation>
). Eye in dorsal view 1.1× as long as temple; occipital carina distinct; supra-antennal elevation medium-sized, about 0.3× as long as scape, outer margin oblique; a pair of oblique projection on frons at inner eye margin (
<figureCitation box="[299,405,212,237]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="7" pageNumber="1160">Figure 1d</figureCitation>
); clypeus markedly concave medio-apically, setose; mandible large.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7049418" ID-Zenodo-Dep="7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="5" pageNumber="1158" startId="5.[160,235,1579,1602]" targetBox="[164,1150,143,1546]" targetPageId="5">
<paragraph blockId="5.[160,1155,1579,1661]" pageId="5" pageNumber="1158">
<emphasis bold="true" box="[160,311,1579,1602]" pageId="5" pageNumber="1158">Figures 1(ae).</emphasis>
<taxonomicName authority="Binoy, van Achterberg &amp; Girish Kumar" authorityName="Binoy, van Achterberg &amp; Girish Kumar" authorityYear="2022" box="[317,930,1579,1602]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="1158" phylum="Arthropoda" rank="species" species="ayyari" status="sp. nov.">
<emphasis box="[317,527,1579,1602]" italics="true" pageId="5" pageNumber="1158">Taeniogonalos ayyari</emphasis>
Binoy, van Achterberg &amp; Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel box="[942,1023,1579,1602]" pageId="5" pageNumber="1158" rank="species">sp. nov.</taxonomicNameLabel>
Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7049420" ID-Zenodo-Dep="7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="6" pageNumber="1159" startId="6.[160,235,1579,1602]" targetBox="[164,1150,142,1546]" targetPageId="6">
<paragraph blockId="6.[160,1156,1579,1690]" pageId="6" pageNumber="1159">
<emphasis bold="true" box="[160,313,1579,1602]" pageId="6" pageNumber="1159">Figures 2(ah).</emphasis>
<taxonomicName authority="Binoy, van Achterberg and Girish Kumar" authorityName="Binoy, van Achterberg and Girish Kumar" authorityYear="2022" box="[320,962,1579,1602]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="1159" phylum="Arthropoda" rank="species" species="ayyari" status="sp. nov.">
<emphasis box="[320,530,1579,1602]" italics="true" pageId="6" pageNumber="1159">Taeniogonalos ayyari</emphasis>
Binoy, van Achterberg and Girish Kumar
</taxonomicName>
,
<taxonomicNameLabel box="[974,1056,1579,1602]" pageId="6" pageNumber="1159" rank="species">sp. nov.</taxonomicNameLabel>
Holotype ♀. a.
<taxonomicName box="[226,336,1608,1631]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="1179" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, lateral view; b.
<taxonomicName box="[520,630,1608,1631]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="1179" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view.
</paragraph>
</caption>
<paragraph blockId="7.[160,1156,289,592]" pageId="7" pageNumber="1160">
<emphasis bold="true" box="[160,294,289,315]" italics="true" pageId="7" pageNumber="1160">
<taxonomicName box="[160,289,289,315]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1160" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
.
</emphasis>
<taxonomicName box="[309,429,290,315]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1179" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
1.2× as long as high (
<figureCitation box="[688,794,290,315]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2a</figureCitation>
); transverse mesopleural sulcus indistinct; mesopleuron irregularly rugose anteriorly, indistinct below setation; posteriorly smooth, shiny; notauli deep, moderately narrow and largely crenulate; middle lobe of mesoscutum coarsely transversely rugose-punctate, lateral lobes with similar sculpture, short longitudinal parapsidal line indicated on lateral lobes; scutellar sulcus complete, medium-sized, finely crenulate; scutellum similarly sculptured as mesoscutum, convex, above level of mesoscutum, metanotum medially slightly convex, not protruding, faintly rugose (
<figureCitation box="[254,363,532,557]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2b</figureCitation>
); propodeum largely longitudinally or obliquely rugose or nearly so; posterior propodeal carina arched (
<figureCitation box="[558,662,567,592]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2c</figureCitation>
).
</paragraph>
<paragraph blockId="7.[160,1155,645,705]" pageId="7" pageNumber="1160">
<emphasis bold="true" box="[160,239,645,671]" italics="true" pageId="7" pageNumber="1160">Wings.</emphasis>
Fore wing: length of vein 1-M 0.9× as long as vein 1-SR, straight; subdiscal and discal cells less setose than remainder of wing (
<figureCitation box="[695,802,680,705]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2d</figureCitation>
).
</paragraph>
<paragraph blockId="7.[160,1156,757,922]" pageId="7" pageNumber="1160">
<emphasis bold="true" box="[160,295,757,783]" italics="true" pageId="7" pageNumber="1160">Metasoma.</emphasis>
First metasomal tergite 2.5× as long as apically wide, smooth (
<figureCitation box="[168,273,793,818]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">
Figure
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</figureCitation>
); second metasomal tergite rugose-punctate, shiny, moderately setose; remaining tergites conspicuously rugose punctate, strongly setose (
<figureCitation box="[866,976,828,853]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">
Figure
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</figureCitation>
); sternites with similar sculpture as tergites; second sternite with posterior margin straight; third sternite 0.2× as long as second sternite; hypopygium bluntly triangular in ventral view (
<figureCitation box="[1037,1142,897,922]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2h</figureCitation>
).
</paragraph>
<paragraph blockId="7.[160,1156,974,1728]" lastBlockId="8.[160,1155,142,237]" lastPageId="8" lastPageNumber="1161" pageId="7" pageNumber="1160">
<emphasis bold="true" box="[160,247,974,1000]" italics="true" pageId="7" pageNumber="1160">Colour.</emphasis>
Black with the following parts differently coloured: head in dorsal view with a distinct V-shaped red band behind stemmaticum extending along occipital line and forming diverging band from occiput towards eye on either side (
<figureCitation box="[911,1018,1045,1070]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="7" pageNumber="1160">Figure 1d</figureCitation>
), pale spots anterior to median ocellus, faint red patch just beyond middle of post-ocellar line; head in frontal view black with following parts variously coloured: patch arising ventrad to lower ocular line, not extending onto middle of face, not meeting medially, on supra-antennal elevation, non-confluent patch on clypeal shield, medially interrupted by black band; mandible yellow with brown-red to black teeth; head in lateral view with yellow patch on outer orbital margin, meeting with red-brown patch from occiput enclosing black patch (
<figureCitation box="[167,271,1287,1312]" captionStart="Figures 1" captionStartId="5.[160,235,1579,1602]" captionTargetBox="[164,1150,143,1546]" captionTargetId="figure-3@5.[163,1153,142,1547]" captionTargetPageId="5" captionText="Figures 1(ae). Taeniogonalos ayyari Binoy, van Achterberg &amp; Girish Kumar, sp. nov. Holotype ♀. a. Habitus, lateral view; b. Antenna; c. Head, lateral view; d. Head, dorsal view; e. Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7049418" httpUri="https://zenodo.org/record/7049418/files/figure.png" pageId="7" pageNumber="1160">Figure 1e</figureCitation>
); antennomeres testaceous, ventrally pale; mesosoma variably coloured, pronotum black with postero-lateral angle pale yellow; mesoscutum red-brown with the following parts variously coloured: middle lobe of mesoscutum red-brown with antero-medial inverted triangular black patch, lateral corner pale yellow; lateral lobe of mesoscutum redbrown with anterior part darker; axilla red-brown; scutellum black flanked by pale yellow patch on each side, anterior margin red-brown, lateral margin beyond yellow patch redbrown; metanotum pale yellow with median lobe dark (
<figureCitation box="[779,883,1495,1520]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2b</figureCitation>
); propodeum black with two pale yellow patches sub-medially (
<figureCitation box="[594,696,1530,1555]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="7" pageNumber="1160">Figure 2c</figureCitation>
); fore and mid coxae brown with ventral area dark; fore and mid femora dark brown with ventral margin pale brown; fore and mid tibiae brown with apex dark brown; hind coxa pale yellow with ventral margin black; all trochanters pale yellow; hind trochanter pale yellow with ventral margin brown; hind femur brown with apico-ventral area pale; all tarsi brown; fore wing largely fuscous with subapical dark brown patch; first metasomal tergite with pale yellow patch on antero-medial margin; second tergite with narrow subapical yellow band (
<figureCitation box="[847,955,142,167]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="8" pageNumber="1161">
Figure
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</figureCitation>
); fourth and fifth metasomal tergites with non-confluent yellow band/macula on apical margin; sixth tergite yellow (
<figureCitation box="[244,347,212,237]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="8" pageNumber="1161">
Figure
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</figureCitation>
); firstsecond metasomal sternites with subapical yellow band (
<figureCitation box="[1040,1142,212,237]" captionStart="Figures 2" captionStartId="6.[160,235,1579,1602]" captionTargetBox="[164,1150,142,1546]" captionTargetId="figure-3@6.[163,1153,142,1547]" captionTargetPageId="6" captionText="Figures 2(ah). Taeniogonalos ayyari Binoy, van Achterberg and Girish Kumar, sp. nov. Holotype ♀. a. Mesosoma, lateral view; b. Mesosoma, dorsal view; c. Propodeum, dorsal view; d. Fore wing; e. Metasoma, lateral view; f. Terminal tergite and sternite, lateral view; g. Metasomal tergites, dorsal view; h. Metasomal sternites, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7049420" httpUri="https://zenodo.org/record/7049420/files/figure.png" pageId="8" pageNumber="1161">Figure 2h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[160,275,290,351]" pageId="8" pageNumber="1161" type="biology_ecology">
<paragraph blockId="8.[160,275,290,351]" box="[160,248,290,316]" pageId="8" pageNumber="1161">
<emphasis bold="true" box="[160,248,290,316]" italics="true" pageId="8" pageNumber="1161">Biology</emphasis>
</paragraph>
<paragraph blockId="8.[160,275,290,351]" box="[160,275,326,351]" pageId="8" pageNumber="1161">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="1161" type="etymology">
<paragraph blockId="8.[160,1155,404,534]" box="[160,285,404,430]" pageId="8" pageNumber="1161">
<emphasis bold="true" box="[160,285,404,430]" italics="true" pageId="8" pageNumber="1161">Etymology</emphasis>
</paragraph>
<paragraph blockId="8.[160,1155,404,534]" pageId="8" pageNumber="1161">
The species name is a genitive noun from the patronym Ayyar, after the Indian entomologist and pioneer T.V. Ramakrishna Ayyar who described many species including
<taxonomicName box="[160,344,509,534]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="fulvoscutellata">
<emphasis box="[160,344,509,534]" italics="true" pageId="8" pageNumber="1161">T. fulvoscutellata</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Ayyar" authorityYear="1919" baseAuthorityName="Ayyar" box="[401,494,509,534]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="kerala">
<emphasis box="[401,494,509,534]" italics="true" pageId="8" pageNumber="1161">T. kerala</emphasis>
</taxonomicName>
from southern
<collectingCountry box="[670,728,509,534]" name="India" pageId="8" pageNumber="1161">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection box="[160,373,588,649]" pageId="8" pageNumber="1161" type="distribution">
<paragraph blockId="8.[160,373,588,649]" box="[160,300,588,614]" pageId="8" pageNumber="1161">
<emphasis bold="true" box="[160,300,588,614]" italics="true" pageId="8" pageNumber="1161">Distribution</emphasis>
</paragraph>
<paragraph blockId="8.[160,373,588,649]" box="[160,373,624,649]" pageId="8" pageNumber="1161">
<collectingCountry box="[160,215,624,649]" name="India" pageId="8" pageNumber="1161">India</collectingCountry>
(
<collectingRegion box="[229,363,624,649]" country="India" name="Tamil Nadu" pageId="8" pageNumber="1161">Tamil Nadu</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="1161" type="discussion">
<paragraph blockId="8.[160,1156,702,1144]" box="[160,262,702,728]" pageId="8" pageNumber="1161">
<emphasis bold="true" box="[160,262,702,728]" italics="true" pageId="8" pageNumber="1161">Remarks</emphasis>
</paragraph>
<paragraph blockId="8.[160,1156,702,1144]" pageId="8" pageNumber="1161">
<taxonomicName authority="Binoy &amp; van Achterberg &amp; Polaszek &amp; Kumar &amp; Santhosh, 2022" authorityName="Binoy &amp; van Achterberg &amp; Polaszek &amp; Kumar &amp; Santhosh" authorityYear="2022" box="[160,392,738,763]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="ayyari" status="sp. nov.">
<emphasis box="[160,392,738,763]" italics="true" pageId="8" pageNumber="1161">Taeniogonalos ayyari</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[399,485,738,763]" pageId="8" pageNumber="1161" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName authority="(Ayyar)" authorityName="Ayyar" authorityYear="1919" baseAuthorityName="Ayyar" box="[612,789,738,763]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="kerala">
<emphasis box="[612,704,738,763]" italics="true" pageId="8" pageNumber="1161">T. kerala</emphasis>
(Ayyar)
</taxonomicName>
in having the head with rugosepunctate sculpture; supra-antennal tubercles short and obtuse, apico-medial margin of second metasomal sternite of female straight, and metasomal armature lacking.
<taxonomicName authority="Binoy &amp; van Achterberg &amp; Polaszek &amp; Kumar &amp; Santhosh, 2022" authorityName="Binoy &amp; van Achterberg &amp; Polaszek &amp; Kumar &amp; Santhosh" authorityYear="2022" box="[160,393,842,867]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="ayyari" status="sp. nov.">
<emphasis box="[160,393,842,867]" italics="true" pageId="8" pageNumber="1161">Taeniogonalos ayyari</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[400,488,842,867]" pageId="8" pageNumber="1161" rank="species">sp. nov.</taxonomicNameLabel>
however differs from
<taxonomicName authorityName="Ayyar" authorityYear="1919" baseAuthorityName="Ayyar" box="[737,830,842,867]" class="Insecta" family="Trigonalidae" genus="Taeniogonalos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="1161" phylum="Arthropoda" rank="species" species="kerala">
<emphasis box="[737,830,842,867]" italics="true" pageId="8" pageNumber="1161">T. kerala</emphasis>
</taxonomicName>
in having the head in dorsal view 1.6× as wide as long (vs 1.4× as wide as long); antenna 24-segmented (vs antenna 22- segmented); mesosoma red-brown (vs brown); second metasomal tergite without any yellow patch on anterior margin (vs second tergite with yellow patch at anterior margin); yellow patch on fourth tergite large (vs yellow patch on fourth tergite comparatively small); hind tibia dark brown without pale basal patch (vs hind tibia brown with pale basal patch); metasoma black with yellow marking (vs metasoma red-brown with yellow marking); yellow band on second metasomal sternite small (vs yellow band on second sternite wider than in alternative); different colour pattern of mesonotum).
</paragraph>
</subSubSection>
</treatment>
</document>