379 lines
57 KiB
XML
379 lines
57 KiB
XML
<document id="B754F494172EA707DF9EBB93F9304A99" ID-DOI="10.5281/zenodo.283181" ID-GBIF-Dataset="1b90d324-193c-4943-a2d4-069e62a48b7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="283181" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564588079" checkinUser="plazi" docAuthor="Han, Taeman, Park, Haechul, Lee, Youngbo, Kim, Namjeong, Ôhira, Hitoo, Platia, Giuseppe & Lee, Seunghwan" docDate="2012" docId="03893739FFABBC118DAB412AF804F926" docLanguage="en" docName="zt03591p066.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cidnopus nigronitidus Han and Lee, sp. nov." docType="treatment" docVersion="8" lastPageNumber="53" masterDocId="FFB04F41FFA8BC178D3C4726FA3AFF96" masterDocTitle="Taxonomic Review of the Subtribe Athouina Candèze, 1859 (Coleoptera, Elateridae, Dendrometrinae, Dendrometrini) in Korea" masterLastPageNumber="66" masterPageNumber="47" pageNumber="50" updateTime="1698320811505" updateUser="plazi">
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<mods:title id="17A291CDCB4D174C6607D10D5F6C7B14">Taxonomic Review of the Subtribe Athouina Candèze, 1859 (Coleoptera, Elateridae, Dendrometrinae, Dendrometrini) in Korea</mods:title>
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<mods:namePart id="4901212257D289F0E6A6E23621F9C014">Han, Taeman</mods:namePart>
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<mods:namePart id="41C82AC6E690C7C2570E0EC944BFD138">Park, Haechul</mods:namePart>
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</mods:name>
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<mods:roleTerm id="96C1E950BA17222A22953A97FA5B71E4">Author</mods:roleTerm>
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<mods:namePart id="A9F8FB7EADFB0A644D1AC7AA950C9E60">Lee, Youngbo</mods:namePart>
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</mods:name>
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<mods:name id="D7E71EED5AE1DD3951A3D5C462C0DCCA" type="personal">
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<mods:roleTerm id="017F6279BBBF2540CFD44F220F85D157">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A8848E6009D6BA0D2F580124F09C75D3">Kim, Namjeong</mods:namePart>
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</mods:name>
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<mods:name id="41BBAFC679A86EC471B3FE4415049732" type="personal">
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<mods:role id="A5D40A6BEC3D11DA1747819BC3F70444">
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<mods:roleTerm id="9F73B7D0F3B948CD6089F824AD6AFABF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="077047E74DDADBEF40C04D1658DDB6BD">Ôhira, Hitoo</mods:namePart>
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</mods:name>
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<mods:name id="1DA1DFC4B984236086D76FEB78CB6277" type="personal">
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</mods:role>
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<mods:namePart id="6CE8A201356159971174F62D7343ECB5">Platia, Giuseppe</mods:namePart>
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</mods:name>
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<mods:roleTerm id="070AAC7CBB601AF44A0CAC9AEDB3E7C7">Author</mods:roleTerm>
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<mods:namePart id="295D49267011A14DD4749C0FF689D43D">Lee, Seunghwan</mods:namePart>
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<mods:typeOfResource id="4BD734193BA6B52C44A5798F46527AFB">text</mods:typeOfResource>
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<mods:title id="1A992A67CB851F7B1A5AD09184FFC843">Zootaxa</mods:title>
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<mods:part id="5AC8485A0BE56E0789282A65D9F1314D">
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<mods:date id="28EA15F5B8F6A43F7C3976D382EF7726">2012</mods:date>
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<mods:detail id="0B65D6D9F01374B61229A52D4E94C3BB" type="volume">
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<mods:number id="23E2F28079F8110B711635E973D418A8">3591</mods:number>
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<mods:classification id="BB0B7DE9F5EC2ECA68A59DDBFF5EC9BB">journal article</mods:classification>
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<mods:identifier id="AA28AF891F768DF186DBBAA6BE31FF64" type="DOI">10.5281/zenodo.283181</mods:identifier>
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<mods:identifier id="74401712D0859ECC584E9AD51E3DB196" type="GBIF-Dataset">1b90d324-193c-4943-a2d4-069e62a48b7a</mods:identifier>
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<mods:identifier id="79EEB35AACA4433562C992C5C8A908C8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0A95B9ED02D8B1CA7CCDA9F26C693CBF" type="Zenodo-Dep">283181</mods:identifier>
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<treatment id="03893739FFABBC118DAB412AF804F926" ID-DOI="http://doi.org/10.5281/zenodo.6173528" ID-GBIF-Taxon="119557417" ID-Zenodo-Dep="6173528" LSID="urn:lsid:plazi:treatment:03893739FFABBC118DAB412AF804F926" httpUri="http://treatment.plazi.org/id/03893739FFABBC118DAB412AF804F926" lastPageId="6" lastPageNumber="53" pageId="3" pageNumber="50">
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<subSubSection id="C33AD5A4FFABBC148DAB412AFBD5F96C" pageId="3" pageNumber="50" type="nomenclature">
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<paragraph id="8B9F862FFFABBC148DAB412AF889F9B3" blockId="3.[151,691,1547,1607]" box="[151,691,1547,1574]" pageId="3" pageNumber="50">
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<heading id="D0D73143FFABBC148DAB412AF889F9B3" bold="true" box="[151,691,1547,1574]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
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<emphasis id="B9545A3DFFABBC148DAB412AF889F9B3" bold="true" box="[151,691,1547,1574]" pageId="3" pageNumber="50">
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<taxonomicName id="4C20FDACFFABBC148DAB412AF874F9B3" authority="Han and Lee" authorityName="Han and Lee" box="[151,590,1547,1574]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="nigronitidus" status="sp. nov.">
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<emphasis id="B9545A3DFFABBC148DAB412AFB9CF9B0" bold="true" box="[151,422,1548,1574]" italics="true" pageId="3" pageNumber="50">Cidnopus nigronitidus</emphasis>
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Han and Lee
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</taxonomicName>
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<taxonomicNameLabel id="A267E746FFABBC148F6A412DF889F9B3" box="[598,691,1547,1573]" pageId="3" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9F862FFFABBC148DAB4109FBFDF9D1" blockId="3.[151,691,1547,1607]" box="[151,455,1583,1607]" pageId="3" pageNumber="50">
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(
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<figureCitation id="131B9AAAFFABBC148DA34109FB27F9D1" box="[159,285,1583,1607]" captionStart="FIGURE 1. A, B" captionStartId="3.[151,250,1414,1437]" captionTargetBox="[217,1326,242,1325]" captionTargetId="figure@3.[151,1436,199,1395]" captionTargetPageId="3" captionText="FIGURE 1. A, B: Cidnopus nigronitidus sp. nov. from Korea. C, D: Limonius scutellaris (Dolin, 2003) com. nov. E – H: Limoniscus kraazi. A (holotype), C, E, G: male; B (paratype), D, F, H: female. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/283182/files/figure.png" pageId="3" pageNumber="50">Figs.1A–B</figureCitation>
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; 2A–R; 3A–B)
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</paragraph>
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<paragraph id="8B9F862FFFABBC148DAB4150FBD5F96C" blockId="3.[151,1436,1654,2002]" pageId="3" pageNumber="50">
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<emphasis id="B9545A3DFFABBC148DAB4150FBA8F919" bold="true" box="[151,402,1654,1679]" pageId="3" pageNumber="50">Specimens examined</emphasis>
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.
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<typeStatus id="549B388DFFABBC148C984151F82BF919" box="[420,529,1655,1679]" pageId="3" pageNumber="50" type="holotype">Holotype</typeStatus>
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: male, Mt. Geryeong, Gongju, CN,
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<collectingCountry id="F337C6BFFFABBC148E84415EFE3AF919" box="[952,1024,1656,1679]" name="South Korea" pageId="3" pageNumber="50">Korea</collectingCountry>
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.
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<date id="FF9EA0EFFFABBC1489324151FEA3F919" box="[1038,1177,1655,1679]" pageId="3" pageNumber="50" value="1989-05-24">24. V. 1989</date>
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. Y. J. Kwon [NAAS].
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<typeStatus id="549B388DFFABBC148DAB41BDFAC7F925" box="[151,253,1691,1715]" pageId="3" pageNumber="50" type="paratype">Paratype</typeStatus>
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: female, Haean-myeon, Yanggu, GW,
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<collectingCountry id="F337C6BFFFABBC148FF441BDF92AF924" box="[712,784,1691,1714]" name="South Korea" pageId="3" pageNumber="50">Korea</collectingCountry>
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.
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<date id="FF9EA0EFFFABBC148E1C41BCF987F924" box="[800,957,1690,1714]" pageId="3" pageNumber="50" value="1990-06-13">13. VI. 1990</date>
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. J. I. Kim [SWU]; female, Mt. Sobaek, Youngju, GB,
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<collectingCountry id="F337C6BFFFABBC148C0641E6FBB8F941" box="[314,386,1728,1751]" name="South Korea" pageId="3" pageNumber="50">Korea</collectingCountry>
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.
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<date id="FF9EA0EFFFABBC148CB14199F828F941" box="[397,530,1727,1751]" pageId="3" pageNumber="50" value="1989-05-30">30. V. 1989</date>
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. Y. J. Kwon [NAAS]; male, Mt. Odae, Jinbu, Pyeongchang, GW,
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<collectingCountry id="F337C6BFFFABBC1489C541E6FF7BF941" box="[1273,1345,1728,1751]" name="South Korea" pageId="3" pageNumber="50">Korea</collectingCountry>
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.
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<date id="FF9EA0EFFFABBC1488714199FAE8F96C" pageId="3" pageNumber="50" value="1998-06-24">24. VI. 1998</date>
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. T. M. HAN, DNA 3049.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33AD5A4FFABBC138DFB4020FBEDF82C" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="description">
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<paragraph id="8B9F862FFFABBC138DFB4020F8D1F9B9" blockId="3.[151,1436,1654,2002]" lastBlockId="4.[151,1437,151,2015]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
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<emphasis id="B9545A3DFFABBC148DFB4020FB6EF889" bold="true" box="[199,340,1798,1823]" pageId="3" pageNumber="50">Description</emphasis>
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.
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<typeStatus id="549B388DFFABBC148C5A4021FBEAF889" box="[358,464,1799,1823]" pageId="3" pageNumber="50" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="131B9AAAFFABBC148CD84021F87EF889" box="[484,580,1799,1823]" captionStart="FIGURE 1. A, B" captionStartId="3.[151,250,1414,1437]" captionTargetBox="[217,1326,242,1325]" captionTargetId="figure@3.[151,1436,199,1395]" captionTargetPageId="3" captionText="FIGURE 1. A, B: Cidnopus nigronitidus sp. nov. from Korea. C, D: Limonius scutellaris (Dolin, 2003) com. nov. E – H: Limoniscus kraazi. A (holotype), C, E, G: male; B (paratype), D, F, H: female. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/283182/files/figure.png" pageId="3" pageNumber="50">Fig. 1A</figureCitation>
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). Male
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<quantity id="4CD82BCAFFABBC148F9E4021F8C7F889" box="[674,765,1799,1823]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="50" unit="mm" value="5.7">5.7 mm</quantity>
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long,
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<quantity id="4CD82BCAFFABBC148E714021F992F889" box="[845,936,1799,1823]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="50" unit="mm" value="1.7">1.7 mm</quantity>
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wide.
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<emphasis id="B9545A3DFFABBC148EC14021FE06F889" bold="true" box="[1021,1084,1799,1823]" pageId="3" pageNumber="50">Body</emphasis>
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sub-cylindrical, rather broad, convex above, parallel-sided; color black, lustrous, with apex of mandibles, tibiae, tarsus dusky reddish brown; pubescences rather short, subrecumbent, pale yellow.
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<emphasis id="B9545A3DFFABBC148E134069F94AF8F1" bold="true" box="[815,880,1871,1895]" pageId="3" pageNumber="50">Head</emphasis>
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(
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<figureCitation id="131B9AAAFFABBC148EB44069F9EEF8F1" box="[904,980,1871,1895]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
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D) flat between eyes; frontal margin expanded anteriorly at above antennal sockets, distinctly and thickly carinate but feebly sinuate inwardly, obscure at middle; punctures (
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<figureCitation id="131B9AAAFFABBC148F3B40B1F86BF839" box="[519,593,1943,1967]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
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E) sub-circular, irregular, simple, deep, but becoming larger and subumbilicate, dense, coarse to outsides, intervals between punctures about 0.3–0.5 times narrower than the diameters of punctures; surface rather rugose laterally; eyes moderately prominent, with ocelli weakly convex; frontal groove (
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<figureCitation id="131B9AAAFFACBC138C62479AFB91FF42" box="[350,427,188,212]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
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F) transversely excavated, slightly narrowed at middle in anterior view; labrum semicircular with long setae anteriorly; mandibles bidentate; maxillary palpi (
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<figureCitation id="131B9AAAFFACBC13891447F9FE54FF61" box="[1064,1134,223,247]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
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Q) axe shaped.
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<emphasis id="B9545A3DFFACBC13881A47F9FFA1FF6E" bold="true" box="[1318,1435,223,248]" pageId="4" pageNumber="51">Antennae</emphasis>
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(
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<figureCitation id="131B9AAAFFACBC138DA34622FADFFE8A" box="[159,229,260,284]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
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G) moderate; 1st antennomere robust, cylindrical; 2nd smallest, sub-globular, 1.5 times as long as wide (9:6), width sub-equal to 3rd (sometimes slightly wider than 3rd in females); 3rd sub-triangular, 1.66 times as long as wide (10:6), slightly longer than 2nd; 4th triangular, 1.33 times as long as wide, 0.63 times shorter than combined length of 2nd and 3rd segments (12:19); 4th to 10th bell-shaped, feebly serrated, each slightly longer than wide; 11th cylindrical, parallel-sided, longest, 2.0 times as long as wide (cat. from 2nd to 11th = 9/6,
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<date id="FF9EA0EFFFACBC13885E46B1FFA2FE39" box="[1378,1432,407,431]" pageId="4" pageNumber="51" value="2010-09-06">10/6</date>
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, 12/9, 11.5/10, 12/9.6,
|
||
<date id="FF9EA0EFFFACBC138CA4469CFBE6FE44" box="[408,476,442,466]" pageId="4" pageNumber="51" value="1912-10-10">12/10</date>
|
||
,
|
||
<date id="FF9EA0EFFFACBC138CD5469CF816FE44" box="[489,556,442,466]" pageId="4" pageNumber="51" value="2011-10-10">11/10</date>
|
||
, 12/10, 12/9.3, 18/9).
|
||
<emphasis id="B9545A3DFFACBC138E0C469DF991FE45" bold="true" box="[816,939,443,467]" pageId="4" pageNumber="51">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC138E80469CFE3BFE44" box="[956,1025,442,466]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
A) sub-quadrate, dome-like convex above, sub-equal in length with width (29:28), widest at middle; anterior margin simple; lateral margins entirely and finely carinate, entirely visible in dorsal view; anterior corner simple, arched from the widest and then slightly convergent anteriorly; punctures (
|
||
<figureCitation id="131B9AAAFFACBC138FB84501F8F6FDA9" box="[644,716,551,575]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
B) distinct, deep, sub-elliptical and of regular size (
|
||
<figureCitation id="131B9AAAFFACBC13880C4501FF43FDA9" box="[1328,1401,551,575]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
C); intervals glabrous, about 1.2–2.0 times wider than punctures; hind angles (
|
||
<figureCitation id="131B9AAAFFACBC138ED1456CFE0BFDF4" box="[1005,1073,586,610]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
H) broad, feebly sinuate at base and divergent posteriorly, with a distinct carina extending inwards at base; posterior margin (
|
||
<figureCitation id="131B9AAAFFACBC1389D84549FF11FD11" box="[1252,1323,623,647]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">
|
||
Fig.
|
||
<date id="FF9EA0EFFFACBC1388204549FF11FD11" box="[1308,1323,623,647]" pageId="4" pageNumber="51">2</date>
|
||
</figureCitation>
|
||
I) lacking basal incisures and basal notch in front of scutellum.
|
||
<emphasis id="B9545A3DFFACBC138E2545B5F990FD3D" bold="true" box="[793,938,659,683]" pageId="4" pageNumber="51">Prosternum</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC138E8245B4FE3FFD3C" box="[958,1029,658,682]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
L) convex, with simple punctures; anterior collar bent antero-inferiorly and roundly margined with punctures clearly large and sub-ocellated, subequal in length to anterior apex of hypomera; posterior process (
|
||
<figureCitation id="131B9AAAFFACBC138EA045FCF9D9FD64" box="[924,995,730,754]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
L and 2M) with longitudinal shallow impression between procoxal cavities, slightly bent beyond procoxal cavities and then expanded as pentagonal shaped laterally, ventral surface narrowly and longitudinally convex in lateral view, lacking a longitudinal depression along each side. Procoxal cavities narrowly open posteriorly; pronotosternal sutures double, smooth madially, but becoming rugose anterioly, slightly sinuate, anterior portion distinctly furrowed and expanded ventrally (
|
||
<figureCitation id="131B9AAAFFACBC138C2F44A9FB67FC31" box="[275,349,911,935]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
N); hypomeron rather broad, with simple sub-elliptical punctures, anterior portion weakly elevated along inner margin, posterior margin nearly straight, but weakly and smoothly emarginate near procoxal cavities.
|
||
<emphasis id="B9545A3DFFACBC138C5244F0FBD2FC79" bold="true" box="[366,488,982,1007]" pageId="4" pageNumber="51">Scutellum</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC138CC744F1F878FC79" box="[507,578,983,1007]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
J and 2K) tongue shaped, 1.37 times (22: 16) as long as wide, widest at apical two thirds; anterior margin narrowest, truncate; lateral sides parallel madially and then broadly and roundly expanded laterally; posterior margin rounded; punctures large, coarse.
|
||
<emphasis id="B9545A3DFFACBC13896C4339FF02FBA1" bold="true" box="[1104,1336,1054,1079]" pageId="4" pageNumber="51">Mesocoxal cavities</emphasis>
|
||
broadly connected with mesoepimeron and narrowly with mesoepisternum. Metasternum convex; median longitudinal suture distinct; hind coxal plates (
|
||
<figureCitation id="131B9AAAFFACBC138F154341F855FBE9" box="[553,623,1127,1151]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
P) widest at insertion of trochanter, outer margin slightly convergent laterally.
|
||
<emphasis id="B9545A3DFFACBC138C3843ADFB07FB35" bold="true" box="[260,317,1163,1187]" pageId="4" pageNumber="51">Legs</emphasis>
|
||
slender; tarsomeres simple, claws (
|
||
<figureCitation id="131B9AAAFFACBC138FE343ACF91FFB34" box="[735,805,1162,1186]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
R) simple.
|
||
<emphasis id="B9545A3DFFACBC138E9443ADF9CCFB35" bold="true" box="[936,1014,1163,1187]" pageId="4" pageNumber="51">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC13893B43ACFE74FB34" box="[1031,1102,1162,1186]" captionStart="FIGURE 2" captionStartId="5.[151,250,1876,1899]" captionTargetBox="[170,1415,183,1849]" captionTargetId="figure@5.[162,1427,169,1855]" captionTargetPageId="5" captionText="FIGURE 2. SEM images of Cidnopus nigronitidus sp. nov. A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2 nd to 4 th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw." httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
|
||
O) sub-ovate, convex above, widest before middle, 2.33 times (77:33) as long as wide, slightly wider than pronotum (33:29), gradually and slightly broadening from base before middle and then gradually rounded posteriorly; sutural line weakly elevated; base weakly convex along anterior edge; striae punctate, distinct; strial intervals feebly convex, weakly rugose transversely. Abdomen with each lateral margin of visible sternites entirely and weakly crenulate; sternite 7 semi-oval, 2.33 times wider than long (42:18).
|
||
<emphasis id="B9545A3DFFACBC138E004218F988FAC1" bold="true" box="[828,946,1342,1367]" pageId="4" pageNumber="51">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC138EF84219FE1AFAC1" box="[964,1056,1343,1367]" captionStart="FIGURE 3. A, B" captionStartId="6.[151,250,1200,1223]" captionTargetBox="[171,1424,577,1180]" captionTargetId="figure@6.[151,1436,563,1193]" captionTargetPageId="6" captionText="FIGURE 3. A, B: Cidnopus nigronitidus sp. nov. from Korea. C: Cidnopus koltzei from Is Sakhalin, Russia (DNA no. 2606). A: Aedeagus. B, C: Female reproductive organs. Acgl: Accessory gland; BC: Bursa copulatrix; CG; Colleterial gland; TE: Tubular extension." httpUri="https://zenodo.org/record/283184/files/figure.png" pageId="4" pageNumber="51">Fig. 3A</figureCitation>
|
||
) 2.75 times (22: 8.0) as long as wide; median lobe slender, sub-parallel-sided and then suddenly narrowed from anterior one fourth to apex, a little longer than parameres, 3.69 times (12.2: 3.3) longer than basal struts; parameres slightly narrow to apex, lateral margins very feebly sinuate, inner margins almost straight, with lateral subapical barb expanded strongly; lateral subapical barb round at anterior margin, 1.47 times (2.8: 1.9) as wide as long, with visible two setae sited on anteror-median portion; basal piece quadrate, widest just after middle, a little wider than long (1.1 times; 8.8: 8.0), 0.36 times (8.0: 22.0) total length of aedeagus.
|
||
</paragraph>
|
||
<paragraph id="8B9F862FFFACBC138DFB411DFB9FF8BC" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="51">
|
||
<emphasis id="B9545A3DFFACBC138DFB411DFB24F9C5" bold="true" box="[199,286,1595,1619]" pageId="4" pageNumber="51">Female</emphasis>
|
||
(
|
||
<figureCitation id="131B9AAAFFACBC138C0C411CFBB0F9C4" box="[304,394,1594,1618]" captionStart="FIGURE 1. A, B" captionStartId="3.[151,250,1414,1437]" captionTargetBox="[217,1326,242,1325]" captionTargetId="figure@3.[151,1436,199,1395]" captionTargetPageId="3" captionText="FIGURE 1. A, B: Cidnopus nigronitidus sp. nov. from Korea. C, D: Limonius scutellaris (Dolin, 2003) com. nov. E – H: Limoniscus kraazi. A (holotype), C, E, G: male; B (paratype), D, F, H: female. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/283182/files/figure.png" pageId="4" pageNumber="51">Fig. 1B</figureCitation>
|
||
). Length 6.6–7.0 mm, width
|
||
<quantity id="4CD82BCAFFACBC138FDD411CF957F9C4" box="[737,877,1594,1618]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="4" pageNumber="51" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.1–2.6 mm</quantity>
|
||
, larger, stouter than male; antennae shorter than male, approximately reaching to base of hind angles of pronotum, 4th antennomere distinctly wider than long (1.53–1.57 times); pronotum more convex and 1.05–1.06 times wider than long.
|
||
<emphasis id="B9545A3DFFACBC13891D41A4FF19F90D" bold="true" box="[1057,1315,1666,1691]" pageId="4" pageNumber="51">Reproductive organs:</emphasis>
|
||
ovipositor sub-triangular, with styli; vagina (
|
||
<figureCitation id="131B9AAAFFACBC138F284181F850F929" box="[532,618,1703,1727]" captionStart="FIGURE 3. A, B" captionStartId="6.[151,250,1200,1223]" captionTargetBox="[171,1424,577,1180]" captionTargetId="figure@6.[151,1436,563,1193]" captionTargetPageId="6" captionText="FIGURE 3. A, B: Cidnopus nigronitidus sp. nov. from Korea. C: Cidnopus koltzei from Is Sakhalin, Russia (DNA no. 2606). A: Aedeagus. B, C: Female reproductive organs. Acgl: Accessory gland; BC: Bursa copulatrix; CG; Colleterial gland; TE: Tubular extension." httpUri="https://zenodo.org/record/283184/files/figure.png" pageId="4" pageNumber="51">Fig. 3B</figureCitation>
|
||
) elongate; uterus enlarged at anterior, with ballon-like colleterial glands, lacking sclerotised structures; bursa copularix elongate, cylindrical, sub-equal in length with width from base to apical, coiled from apical half, with two accessory glands; basal accessory gland distinctly elongate, longer than second accessory gland.
|
||
</paragraph>
|
||
<paragraph id="8B9F862FFFACBC138DFB4010FBEDF82C" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="51">
|
||
<emphasis id="B9545A3DFFACBC138DFB4010FB92F8D9" bold="true" box="[199,424,1846,1871]" pageId="4" pageNumber="51">Reference species.</emphasis>
|
||
<taxonomicName id="4C20FDACFFACBC138C844011F905F8D9" authority="Reitter, 1895" authorityName="Reitter" authorityYear="1895" box="[440,831,1847,1871]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFACBC138C844011F846F8D9" box="[440,636,1847,1871]" italics="true" pageId="4" pageNumber="51">Cidnopus koltzei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFACBC138FA94011F90CF8D9" author="Reitter" box="[661,822,1847,1871]" pageId="4" pageNumber="51" refString="Reitter, E. (1895) Uebersicht der Elateriden- (Coleoptern-) Arten aus der Verwandtschaft des Limonius (Pheletes) aeneoniger Deg. Aus der Palaearctichen Fauna. Wiener Entomologische Zeitung, 14, 256." type="journal article" year="1895">Reitter, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D264DA6FFACBC138E6F4011F9F1F8D9" box="[851,971,1847,1871]" pageId="4" pageNumber="51" type="female">2 females</specimenCount>
|
||
,
|
||
<collectingCountry id="F337C6BFFFACBC138EDC4011FE17F8D9" box="[992,1069,1847,1871]" name="Russia" pageId="4" pageNumber="51">Russia</collectingCountry>
|
||
, Sakhalin, Yuzhno-Sakhalinsk, Vestochka.
|
||
<date id="FF9EA0EFFFACBC138C2B407CFB8BF8E4" box="[279,433,1882,1906]" pageId="4" pageNumber="51" value="2008-07-18">18. VII. 2008</date>
|
||
. J. W. Lee, identified by T. M. Han, DNA
|
||
<date id="FF9EA0EFFFACBC138EBB407CFE15F8E4" box="[903,1071,1882,1906]" pageId="4" pageNumber="51" value="2606" valueMax="2607">2606 and 2607</date>
|
||
;
|
||
<taxonomicName id="4C20FDACFFACBC138907407DFAE6F801" authority="Lewis, 1894" authorityName="Lewis" authorityYear="1894" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="marginicollis">
|
||
<emphasis id="B9545A3DFFACBC138907407DFF7AF8E5" box="[1083,1344,1883,1907]" italics="true" pageId="4" pageNumber="51">Cidnopus marginicollis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFACBC138873407CFAEEF801" author="Lewis" pageId="4" pageNumber="51" refString="Lewis, G. (1894) On the Elateridae of Japan. The Annals and Magazine of Natural History including Zoology, Botany, and Geology, 13, 311 - 320." type="journal article" year="1894">Lewis, 1894</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: male, female, Mt Gomandodan, Wakayama Pref, Honshu,
|
||
<collectingCountry id="F337C6BFFFACBC138E4A40A6F983F801" box="[886,953,1920,1943]" name="Japan" pageId="4" pageNumber="51">Japan</collectingCountry>
|
||
.
|
||
<date id="FF9EA0EFFFACBC138EF84059FE72F801" box="[964,1096,1919,1943]" pageId="4" pageNumber="51" value="1986-05-31">31. V. 1986</date>
|
||
. Leg. H. Hiramatsu, identified by Dr. H. Ôhira, DNA 3044.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AD5A4FFACBC138DFB40E1FE8EF849" box="[199,1204,1991,2015]" pageId="4" pageNumber="51" type="etymology">
|
||
<paragraph id="8B9F862FFFACBC138DFB40E1FE8EF849" blockId="4.[151,1437,151,2015]" box="[199,1204,1991,2015]" pageId="4" pageNumber="51">
|
||
<emphasis id="B9545A3DFFACBC138DFB40E1FB7CF849" bold="true" box="[199,326,1991,2015]" pageId="4" pageNumber="51">Etymology</emphasis>
|
||
. This new species was named
|
||
<taxonomicName id="4C20FDACFFACBC138FA240EEF97DF849" box="[670,839,1991,2015]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="nigronitidus">
|
||
<emphasis id="B9545A3DFFACBC138FA240EEF97DF849" box="[670,839,1991,2015]" italics="true" pageId="4" pageNumber="51">C. nigronitidus</emphasis>
|
||
</taxonomicName>
|
||
based on its black shining body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5FD6A7FFADBC128DAB4072F859F867" httpUri="https://zenodo.org/record/283183/files/figure.png" pageId="5" pageNumber="52" targetBox="[170,1415,183,1849]" targetPageId="5">
|
||
<paragraph id="8B9F862FFFADBC128DAB4072F859F867" blockId="5.[151,1436,1876,2034]" pageId="5" pageNumber="52">
|
||
<emphasis id="B9545A3DFFADBC128DAB4072FB2FF8FF" bold="true" box="[151,277,1876,1899]" pageId="5" pageNumber="52">FIGURE 2.</emphasis>
|
||
SEM images of
|
||
<taxonomicName id="4C20FDACFFADBC128CF54072F894F8FC" box="[457,686,1876,1898]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="nigronitidus" status="sp. nov.">
|
||
<emphasis id="B9545A3DFFADBC128CF54072F894F8FC" box="[457,686,1876,1898]" italics="true" pageId="5" pageNumber="52">Cidnopus nigronitidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A267E746FFADBC128F8A4073F93DF8FC" box="[694,775,1877,1898]" pageId="5" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A: Pronotum. B: Punctures of median portion of pronotum. C: Punctures of anterior portion of pronotum. D: Head. E: Punctures of median portion of head. F: Frontal view of the head. G: 2nd to 4th antennomeres, male. H: Hind angle of pronotum. I: Posterior margin of pronotum. J: Scutellum. K: Lateral view of scutellum. L: Prosternum. M: Lateral view of prosternal process. N: Anterior portion of hypomeron. O: Right elytron. P: Hind coxal plate. Q: Maxillary palpi. R: Fore claw.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33AD5A4FFAEBC118DFB47B1FBC0F91A" pageId="6" pageNumber="53" type="diagnosis">
|
||
<paragraph id="8B9F862FFFAEBC118DFB47B1FE0BFD81" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="53">
|
||
<emphasis id="B9545A3DFFAEBC118DFB47B1FB00FF26" bold="true" box="[199,314,151,176]" pageId="6" pageNumber="53">Diagnosis</emphasis>
|
||
. This new species is closely allied to
|
||
<taxonomicName id="4C20FDACFFAEBC118FDE47BEF9C0FF39" authority="Reitter, 1895" authorityName="Reitter" authorityYear="1895" box="[738,1018,151,176]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC118FDE47BEF976FF26" box="[738,844,152,176]" italics="true" pageId="6" pageNumber="53">C. koltzei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118E6747B1F9CBFF39" author="Reitter" box="[859,1009,151,175]" pageId="6" pageNumber="53" refString="Reitter, E. (1895) Uebersicht der Elateriden- (Coleoptern-) Arten aus der Verwandtschaft des Limonius (Pheletes) aeneoniger Deg. Aus der Palaearctichen Fauna. Wiener Entomologische Zeitung, 14, 256." type="journal article" year="1895">Reitter, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which occurs from Russian Far East to
|
||
<collectingCountry id="F337C6BFFFAEBC118D8F479AFB19FF42" box="[179,291,188,212]" name="Mongolia" pageId="6" pageNumber="53">Mongolia</collectingCountry>
|
||
. We examined two female specimens of
|
||
<taxonomicName id="4C20FDACFFAEBC118FD4479BF968FF42" box="[744,850,188,212]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC118FD4479BF968FF42" box="[744,850,188,212]" italics="true" pageId="6" pageNumber="53">C. koltzei</emphasis>
|
||
</taxonomicName>
|
||
collected from Is. Sakhalin,
|
||
<collectingCountry id="F337C6BFFFAEBC1189AE479AFEDBFF42" box="[1170,1249,188,212]" name="Russia" pageId="6" pageNumber="53">Russia</collectingCountry>
|
||
. By comparative examination of the females of these two species, the new species can be easily distinguished from
|
||
<taxonomicName id="4C20FDACFFAEBC1189DA47C6FF6AFF6E" box="[1254,1360,224,248]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC1189DA47C6FF6AFF6E" box="[1254,1360,224,248]" italics="true" pageId="6" pageNumber="53">C. koltzei</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: In the former, the 2nd antennomere is subequal in length to the 3rd, the pronotal lateral margin is slightly arched in the middle, the prosternal process has a shallow impression between the procoxal cavities, and the lateral expansion of the prosternal process is rather short. In the latter, the 3rd antennomere is slightly longer (1.18−1.27 times) than the 2nd and almost parallel-sided from the anterior one-fourth posteriorly in the pronotal lateral margin, and the prosternal process is flat between the procoxal cavities and more or less elongate posterioly. In particular, the novel species is clearly distinguished from
|
||
<taxonomicName id="4C20FDACFFAEBC11892B469EFEBAFE46" box="[1047,1152,440,464]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC11892B469EFEBAFE46" box="[1047,1152,440,464]" italics="true" pageId="6" pageNumber="53">C. koltzei</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the bursa copulatrix (BC) of the female. The former has a cylindrical BC with a distinctly elongated basal accessory gland, but the latter has a more enlarged BC with a short basal accessory gland (
|
||
<figureCitation id="131B9AAAFFAEBC118EF046D9FE34FD81" box="[972,1038,511,535]" captionStart="FIGURE 3. A, B" captionStartId="6.[151,250,1200,1223]" captionTargetBox="[171,1424,577,1180]" captionTargetId="figure@6.[151,1436,563,1193]" captionTargetPageId="6" captionText="FIGURE 3. A, B: Cidnopus nigronitidus sp. nov. from Korea. C: Cidnopus koltzei from Is Sakhalin, Russia (DNA no. 2606). A: Aedeagus. B, C: Female reproductive organs. Acgl: Accessory gland; BC: Bursa copulatrix; CG; Colleterial gland; TE: Tubular extension." httpUri="https://zenodo.org/record/283184/files/figure.png" pageId="6" pageNumber="53">Fig. 3</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<caption id="DF5FD6A7FFAEBC118DAB4396FB6FFA95" httpUri="https://zenodo.org/record/283184/files/figure.png" pageId="6" pageNumber="53" targetBox="[171,1424,577,1180]" targetPageId="6">
|
||
<paragraph id="8B9F862FFFAEBC118DAB4396FB6FFA95" blockId="6.[151,1437,1200,1283]" pageId="6" pageNumber="53">
|
||
<emphasis id="B9545A3DFFAEBC118DAB4396FB29FB53" bold="true" box="[151,275,1200,1223]" pageId="6" pageNumber="53">FIGURE 3.</emphasis>
|
||
A, B:
|
||
<taxonomicName id="4C20FDACFFAEBC118C654396F806FB50" box="[345,572,1200,1222]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="nigronitidus" status="sp. nov.">
|
||
<emphasis id="B9545A3DFFAEBC118C654396F806FB50" box="[345,572,1200,1222]" italics="true" pageId="6" pageNumber="53">Cidnopus nigronitidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A267E746FFAEBC118F7F4397F8A8FB50" box="[579,658,1201,1222]" pageId="6" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Korea. C:
|
||
<taxonomicName id="4C20FDACFFAEBC118E074396F9DFFB50" box="[827,997,1200,1222]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC118E074396F9DFFB50" box="[827,997,1200,1222]" italics="true" pageId="6" pageNumber="53">Cidnopus koltzei</emphasis>
|
||
</taxonomicName>
|
||
from Is Sakhalin, Russia (DNA no. 2606). A: Aedeagus. B, C: Female reproductive organs. Acgl: Accessory gland; BC: Bursa copulatrix; CG; Colleterial gland; TE: Tubular extension.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F862FFFAEBC118DFB4216FBC0F91A" blockId="6.[151,1436,1328,1713]" pageId="6" pageNumber="53">
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118DFB4216FB8AFADE" author="Mertlik" box="[199,432,1328,1352]" pageId="6" pageNumber="53" refString="Mertlik, J. (1996) Cidnopus platiai sp. n. (Coleoptera: Elateridae) from Slovakia. Klapalekiana, 32, 203 - 208." type="journal article" year="1996">Mertlik (1996: 207)</bibRefCitation>
|
||
provided a key of six European species of
|
||
<taxonomicName id="4C20FDACFFAEBC118EFF4216FE14FADE" box="[963,1070,1328,1352]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9545A3DFFAEBC118EFF4216FE14FADE" box="[963,1070,1328,1352]" italics="true" pageId="6" pageNumber="53">Cidnopus</emphasis>
|
||
</taxonomicName>
|
||
. According to this scheme,
|
||
<taxonomicName id="4C20FDACFFAEBC1188BE4217FB25FAFA" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="nigronitidus" status="sp. nov.">
|
||
<emphasis id="B9545A3DFFAEBC1188BE4217FB25FAFA" italics="true" pageId="6" pageNumber="53">C. nigronitidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A267E746FFAEBC118C154273FBB8FAFA" box="[297,386,1365,1388]" pageId="6" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shares the same diagnostic characteristics, such as a slightly convex pronotum and visible lateral margin in the dorsal view, with
|
||
<taxonomicName id="4C20FDACFFAEBC118F75425FF9A8FA06" authority="Schwarz, 1900" authorityName="Schwarz" authorityYear="1900" box="[585,914,1400,1424]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B9545A3DFFAEBC118F75425FF8E8FA06" box="[585,722,1401,1424]" italics="true" pageId="6" pageNumber="53">C. crassipes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118FDD425EF9B0FA06" author="Schwarz" box="[737,906,1400,1424]" pageId="6" pageNumber="53" refString="Schwarz, O. (1900) Neue palaarktische Elateriden. Deutsche Entomologische Zeitschrift, 44, 98 - 111." type="journal article" year="1900">Schwarz, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F337C6BFFFAEBC118E8B425EFE32FA06" box="[951,1032,1400,1424]" name="Turkey" pageId="6" pageNumber="53">Turkey</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C20FDACFFAEBC11892F425FFF1DFA06" authority="Leske, 1785" authorityName="Leske" authorityYear="1785" box="[1043,1319,1400,1424]" class="Insecta" family="Elateridae" genus="Elater" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="pilosus">
|
||
<emphasis id="B9545A3DFFAEBC11892F425FFEB9FA06" box="[1043,1155,1400,1424]" italics="true" pageId="6" pageNumber="53">C. pilosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFAEBC1189AF425EFF25FA06" author="Leske" box="[1171,1311,1400,1424]" pageId="6" pageNumber="53" refString="Leske, N. G. (1785) Reise durch Sachsen in Ruksicht der Naturgeschichte und Okonomie unternommen und beschrieben. Leipzing, (Muller), pp. I - XXX, [1], 1 - 548, [1 - 2], Taf. A, Taf. 1 - 38." type="journal article" year="1785">Leske, 1785</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in Europe and
|
||
<collectingCountry id="F337C6BFFFAEBC118DFB42BAFB7DFA22" box="[199,327,1436,1460]" name="Turkey" pageId="6" pageNumber="53">Asia minor</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C20FDACFFAEBC118CBE42BBF8A2FA22" authority="Reitter, 1895" authorityName="Reitter" authorityYear="1895" box="[386,664,1436,1460]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="koltzei">
|
||
<emphasis id="B9545A3DFFAEBC118CBE42BBFBD6FA22" box="[386,492,1436,1460]" italics="true" pageId="6" pageNumber="53">C. koltzei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118CC042BAF8AAFA22" author="Reitter" box="[508,656,1436,1460]" pageId="6" pageNumber="53" refString="Reitter, E. (1895) Uebersicht der Elateriden- (Coleoptern-) Arten aus der Verwandtschaft des Limonius (Pheletes) aeneoniger Deg. Aus der Palaearctichen Fauna. Wiener Entomologische Zeitung, 14, 256." type="journal article" year="1895">Reitter, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in Far East
|
||
<collectingCountry id="F337C6BFFFAEBC118E1E42BAF94AFA22" box="[802,880,1436,1460]" name="Russia" pageId="6" pageNumber="53">Russia</collectingCountry>
|
||
. We did not examine a specimen of
|
||
<taxonomicName id="4C20FDACFFAEBC11883542BBFBB9FA4E" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9545A3DFFAEBC11883542BBFFA6FA22" box="[1289,1436,1436,1460]" italics="true" pageId="6" pageNumber="53">C. parallelus</emphasis>
|
||
(Motschulsky, 1860)
|
||
</taxonomicName>
|
||
in this study.
|
||
<taxonomicName id="4C20FDACFFAEBC118F0D42E6F90AFA4E" box="[561,816,1472,1496]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="nigronitidus" status="sp. nov.">
|
||
<emphasis id="B9545A3DFFAEBC118F0D42E6F90AFA4E" box="[561,816,1472,1496]" italics="true" pageId="6" pageNumber="53">Cidnopus nigronitidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A267E746FFAEBC118E0042E7F9ADFA4E" box="[828,919,1473,1496]" pageId="6" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be easily separated from
|
||
<taxonomicName id="4C20FDACFFAEBC1189C242E7FFAFFA4E" box="[1278,1429,1472,1496]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9545A3DFFAEBC1189C242E7FFAFFA4E" box="[1278,1429,1472,1496]" italics="true" pageId="6" pageNumber="53">C. parallelus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118DAB42C2FB4BFA6A" author="Reitter" box="[151,369,1508,1532]" pageId="6" pageNumber="53" refString="Reitter, E. (1895) Uebersicht der Elateriden- (Coleoptern-) Arten aus der Verwandtschaft des Limonius (Pheletes) aeneoniger Deg. Aus der Palaearctichen Fauna. Wiener Entomologische Zeitung, 14, 256." type="journal article" year="1895">Reitter (1895: 256)</bibRefCitation>
|
||
characterized
|
||
<taxonomicName id="4C20FDACFFAEBC118F2B42C3F891FA6A" box="[535,683,1508,1532]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9545A3DFFAEBC118F2B42C3F891FA6A" box="[535,683,1508,1532]" italics="true" pageId="6" pageNumber="53">C. parallelus</emphasis>
|
||
</taxonomicName>
|
||
as having the carinate pronotal hind angle, which is invisibly and completely coincident with the lateral margin. Additionally,
|
||
<bibRefCitation id="EFB1FBDEFFAEBC118E42412EFED3F9B6" author="Gurjeva" box="[894,1257,1544,1568]" pageId="6" pageNumber="53" refString="Gurjeva, E. L. (1989) 40. Family Elateridae In: Ler, P. A. (Eds.), Key book of insect species of the Far East of Russia. Vol. 3. Coleoptera, or beetles. Pt. 1. Leningrad, Nauka, pp. 489 - 534." type="book chapter" year="1989">Gurjeva (1989: 494 and 501)</bibRefCitation>
|
||
illustrated the hypomeron, aedeagus, and the dorsal habitus of
|
||
<taxonomicName id="4C20FDACFFAEBC118FFC410BF96EF9D2" box="[704,852,1580,1604]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9545A3DFFAEBC118FFC410BF96EF9D2" box="[704,852,1580,1604]" italics="true" pageId="6" pageNumber="53">C. parallelus</emphasis>
|
||
</taxonomicName>
|
||
. Especially, the aedeagus of
|
||
<taxonomicName id="4C20FDACFFAEBC11899D410BFF0FF9D2" box="[1185,1333,1580,1604]" class="Insecta" family="Elateridae" genus="Cidnopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9545A3DFFAEBC11899D410BFF0FF9D2" box="[1185,1333,1580,1604]" italics="true" pageId="6" pageNumber="53">C. parallelus</emphasis>
|
||
</taxonomicName>
|
||
has subtriangular shape (not roundly expanded to apex) in the subapical barbs of the parameres and distinctly sinuate at the inner margins of the parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AD5A4FFAEBC118DFB41BEF804F926" box="[199,574,1688,1713]" pageId="6" pageNumber="53" type="distribution">
|
||
<paragraph id="8B9F862FFFAEBC118DFB41BEF804F926" blockId="6.[151,1436,1328,1713]" box="[199,574,1688,1713]" pageId="6" pageNumber="53">
|
||
<emphasis id="B9545A3DFFAEBC118DFB41BEFB63F927" bold="true" box="[199,345,1688,1713]" pageId="6" pageNumber="53">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F337C6BFFFAEBC118C5B41BEF803F926" box="[359,569,1688,1712]" name="South Korea" pageId="6" pageNumber="53">Republic of Korea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |