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<document id="32514DE2D0B5A0F4555362903971BF4B" ID-DOI="10.11646/phytotaxa.213.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13638565" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725340364670" checkinUser="felipe" docAuthor="Chen, You-Sheng &amp; Yuan, Qian" docDate="2015" docId="9739CE7361170D19FF04FC69FF5AF074" docLanguage="en" docName="phytotaxa.213.3.1.pdf" docOrigin="Phytotaxa 213 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.213.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Saussurea jiulongensis Y. S. Chen 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="204" masterDocId="6B00B60B613A0D37FF8CFFADFF91F13D" masterDocTitle="Twenty-six new species of Saussurea (Asteraceae, Cardueae) from the Qinghai-Tibetan Plateau and adjacent regions" masterLastPageNumber="211" masterPageNumber="159" pageNumber="204" updateTime="1725346449099" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2F84EF51984B5872E21D0F1CD892FBB5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2EA2780C3E4048D0B9193A60BDE692A6">
<mods:title id="18F88B03E80D0708D9C0765D6E36FE9E">Twenty-six new species of Saussurea (Asteraceae, Cardueae) from the Qinghai-Tibetan Plateau and adjacent regions</mods:title>
</mods:titleInfo>
<mods:name id="90EFB17D4EDAE6B30A70D4E62DFA577F" type="personal">
<mods:role id="8518865A99DD0786780D33B10B2C870A">
<mods:roleTerm id="2F0450DB6D31D8AE27A9B41E430BDD96">Author</mods:roleTerm>
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<mods:namePart id="D4E3E6B4F10F1E7ABF569A41F2A92FFC">Chen, You-Sheng</mods:namePart>
<mods:affiliation id="C33D0E937D73E9FBED8003D92D90FEFF">State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, 20 Nanxincun, Xiangshan, Beijing 100093, China</mods:affiliation>
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</mods:role>
<mods:namePart id="84FD5CB12332641F75984D00577227E6">Yuan, Qian</mods:namePart>
<mods:affiliation id="3DB9E634A54CF8F3A7DA4E511E247692">State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, 20 Nanxincun, Xiangshan, Beijing 100093, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China.</mods:affiliation>
</mods:name>
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<mods:title id="6C494127FD2C1C287F0472552263F673">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="E7C58A0DF0A2596E7E94AB795DEE2125">
<mods:date id="47DBF64DF26DA8F59521680FCA7FCF42">2015</mods:date>
<mods:detail id="0EAFC33DAB08CA477FCB298ACD6998B1" type="pubDate">
<mods:number id="381E86E476F5514BDCAF26547BD851DE">2015-06-17</mods:number>
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<mods:detail id="E86AA0220F96A20FD0FB684C99A012AA" type="volume">
<mods:number id="9FD855CE0B892F7DCA4294259404F134">213</mods:number>
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<mods:number id="B9BCEEEE71338B95D887CF6DD529C458">3</mods:number>
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<mods:url id="613712D76BD6A433541CCAEE938EC1AC">http://dx.doi.org/10.11646/phytotaxa.213.3.1</mods:url>
</mods:location>
<mods:classification id="B2C5E1BB9157A84E2BECF0545EA21468">journal article</mods:classification>
<mods:identifier id="87CA64301C25762DEA09EC34325818BE" type="DOI">10.11646/phytotaxa.213.3.1</mods:identifier>
<mods:identifier id="438BF0DD3A129BB75E48BDF7345A67E1" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="E70C480E760083FB086DEA07F15D682A" type="Zenodo-Dep">13638565</mods:identifier>
</mods:mods>
<treatment id="9739CE7361170D19FF04FC69FF5AF074" ID-DOI="http://doi.org/10.5281/zenodo.13638665" ID-Zenodo-Dep="13638665" LSID="urn:lsid:plazi:treatment:9739CE7361170D19FF04FC69FF5AF074" httpUri="http://treatment.plazi.org/id/9739CE7361170D19FF04FC69FF5AF074" lastPageId="46" lastPageNumber="204" pageId="45" pageNumber="204">
<subSubSection id="578A2CEE61170D1AFF04FC69FC9AF2E0" box="[136,779,962,989]" pageId="45" pageNumber="204" type="nomenclature">
<paragraph id="1F2F7F6561170D1AFF04FC69FC9AF2E0" blockId="45.[136,779,962,989]" box="[136,779,962,989]" pageId="45" pageNumber="204">
<heading id="4467C80961170D1AFF04FC69FC9AF2E0" box="[136,779,962,989]" fontSize="11" level="2" pageId="45" pageNumber="204" reason="4">
<emphasis id="2DE4A37761170D1AFF04FC69FF3AF2E0" bold="true" box="[136,171,964,989]" pageId="45" pageNumber="204">24.</emphasis>
<taxonomicName id="D89004E661170D1AFF3EFC6EFDACF2E0" ID-CoL="7YXWN" authority="Y. S. Chen" authorityName="Y. S. Chen" authorityYear="2015" box="[178,573,962,989]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="species" species="jiulongensis" status="sp. nov.">
<emphasis id="2DE4A37761170D1AFF3EFC6EFE2CF2E0" bold="true" box="[178,445,962,989]" italics="true" pageId="45" pageNumber="204">Saussurea jiulongensis</emphasis>
Y. S. Chen
</taxonomicName>
,
<emphasis id="2DE4A37761170D1AFDC4FC69FD33F2E0" bold="true" box="[584,674,964,989]" pageId="45" pageNumber="204">
<taxonomicNameLabel id="36D71E0C61170D1AFDC4FC69FD33F2E0" box="[584,674,964,989]" pageId="45" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="87AB63E061170D1AFD3DFC69FC93F2E0" box="[689,770,964,989]" captionStart="FIGURE 24" captionStartId="46.[136,229,1984,2005]" captionTargetBox="[226,1361,353,1958]" captionTargetId="figure-88@46.[226,1361,353,1958]" captionTargetPageId="46" captionText="FIGURE 24. Saussurea jiulongensis. A. Habit. B. Leaf section (adaxial surface).C. Leaf section (abaxial surface). D. Inner pappus bristle. E. Outer pappus bristle. F. Phyllaries (from left to right, outer to inner series). G. Anthers. H. Floret (with pappus removed). I. Floret. J. Style branches.All from Y. S. Chen 7551 (PE). Drawn by Mr. Y.X. Zhu." figureDoi="http://doi.org/10.5281/zenodo.13638639" httpUri="https://zenodo.org/record/13638639/files/figure.png" pageId="45" pageNumber="204">Fig. 24</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="578A2CEE61170D1AFF04FBA2FE60F790" pageId="45" pageNumber="204" type="materials_examined">
<paragraph id="1F2F7F6561170D1AFF04FBA2FECDF575" blockId="45.[136,1452,1039,1096]" pageId="45" pageNumber="204">
<materialsCitation id="AFF8753861170D1AFF04FBA2FEC9F575" collectingDate="2007-10-14" collectionCode="Y, S, PE" country="China" county="Jiulong County" elevation="2650" location="Yangfanggou" municipality="Type" pageId="45" pageNumber="204" specimenCount="2" stateProvince="Sichuan" typeStatus="holotype">
<collectingMunicipality id="FF4BE51F61170D1AFF04FBA2FF2BF519" box="[136,186,1039,1060]" pageId="45" pageNumber="204">Type</collectingMunicipality>
:—
<collectingCountry id="67873FF561170D1AFF5BFBA2FEB7F519" box="[215,294,1039,1060]" name="China" pageId="45" pageNumber="204">CHINA</collectingCountry>
.
<collectingRegion id="DD54B18761170D1AFEA3FBA2FE11F519" box="[303,384,1039,1060]" country="China" name="Sichuan" pageId="45" pageNumber="204">Sichuan</collectingRegion>
:
<collectingCounty id="F64E07E961170D1AFE06FBA2FDB1F519" box="[394,544,1039,1060]" pageId="45" pageNumber="204">Jiulong County</collectingCounty>
,
<location id="1A4F29BE61170D1AFDA5FBA2FD3DF519" LSID="urn:lsid:plazi:treatment:9739CE7361170D19FF04FC69FF5AF074:1A4F29BE61170D1AFDA5FBA2FD3DF519" box="[553,684,1039,1060]" country="China" county="Jiulong County" municipality="Type" name="Yangfanggou" pageId="45" pageNumber="204" stateProvince="Sichuan">Yangfanggou</location>
, in forest along the river,
<quantity id="D868D28061170D1AFC2BFBA2FC62F519" box="[935,1011,1039,1060]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="45" pageNumber="204" unit="m" value="2650.0">
<elevation id="94BD985661170D1AFC2BFBA2FC62F519" box="[935,1011,1039,1060]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="45" pageNumber="204" unit="m" value="2650.0">2650 m</elevation>
</quantity>
,
<date id="6B2E59A561170D1AFC72FBA2FB32F519" box="[1022,1187,1039,1060]" pageId="45" pageNumber="204" value="2007-10-14">
<collectingDate id="7B6AA04D61170D1AFC72FBA2FB32F519" box="[1022,1187,1039,1060]" pageId="45" pageNumber="204" value="2007-10-14">14 October 2007</collectingDate>
</date>
,
<emphasis id="2DE4A37761170D1AFB22FBA2FAD9F519" box="[1198,1352,1039,1060]" italics="true" pageId="45" pageNumber="204">
<collectionCode id="7981E7A061170D1AFB22FBA2FB2DF519" box="[1198,1212,1039,1060]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="45" pageNumber="204" type="Herbarium">Y</collectionCode>
.
<collectionCode id="7981E7A061170D1AFB4AFBA2FB44F519" box="[1222,1237,1039,1060]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="45" pageNumber="204" type="Herbarium">S</collectionCode>
. Chen 7551
</emphasis>
(
<typeStatus id="C02BC1C761170D1AFADAFBA2FA3DF519" box="[1366,1452,1039,1060]" pageId="45" pageNumber="204" type="holotype">holotype</typeStatus>
<collectionCode id="7981E7A061170D1AFF31FB9EFF4DF575" box="[189,220,1075,1096]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="45" pageNumber="204" type="Herbarium">PE</collectionCode>
;
<typeStatus id="C02BC1C761170D1AFF6AFB9EFEBAF575" box="[230,299,1075,1096]" pageId="45" pageNumber="204" type="isotype">isotype</typeStatus>
<collectionCode id="7981E7A061170D1AFEBDFB9EFEC0F575" box="[305,337,1075,1096]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="45" pageNumber="204" type="Herbarium">PE</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="1F2F7F6561170D1AFF04FBD5FE60F790" blockId="45.[136,1452,1144,2069]" pageId="45" pageNumber="204">
Herbs perennial, to
<quantity id="D868D28061170D1AFEE6FBD5FE21F5AC" box="[362,432,1144,1169]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="45" pageNumber="204" unit="cm" value="90.0">90 cm</quantity>
tall. Rhizomes ascending, unbranched, with many dark brown small roots. Stem solitary, erect, ca.
<quantity id="D868D28061170D1AFF74FB31FEAAF588" box="[248,315,1180,1205]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="45" pageNumber="204" unit="mm" value="4.0">4 mm</quantity>
in diameter, nearly glabrous, winged, longitudinally striate, apically paniculately branched. Leaves petiolate, papery; lower leaf blade elliptic, 8–16(–20) cm long, 3.5–5.5(–7.2) cm wide, both surfaces green, glabrous, but with sparse short glandular hairs, margin undulate dentate to shallowly pinnatifid, mucronate, apex acute to acuminate, base slightly cordate, decurrent into stems forming wings. Petioles winged,
<quantity id="D868D28061170D1AFBF6FAA5FB4CF41C" box="[1146,1245,1288,1313]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="45" pageNumber="204" unit="cm" value="8.0" valueMax="12.0" valueMin="4.0">4–12 cm</quantity>
long, base wider, auriculate and semiamplexicaul; wings entire. Upper stem leaves narrowly elliptic to lanceolate, gradually smaller apically on stem, base attenuate, petioles much shorter. Capitula numerous, paniculate, in sparse corymbs at the end of branchlets or stems. Involucre campanulate,
<quantity id="D868D28061170D1AFD14FAD9FD64F4B0" box="[664,757,1396,1421]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="45" pageNumber="204" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
in diameter. Phyllaries in ca. 5 rows, imbricate, margin entire, green to brown; outer phyllaries triangular-ovate, ca.
<quantity id="D868D28061170D1AFD64FA35FCBAF48C" box="[744,811,1432,1457]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="204" unit="mm" value="2.0">2 mm</quantity>
long, ca.
<quantity id="D868D28061170D1AFC11FA35FC4EF48C" box="[925,991,1432,1457]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="204" unit="mm" value="1.0">1 mm</quantity>
wide, leathery, pubescent, apex acute to obtuse; middle series of phyllaries longer, linear-oblong, pubescent, apex acute to acuminate; inner phyllaries linear-lanceolate, pubescent, apex acute to acuminate,
<quantity id="D868D28061170D1AFD21FA4DFCBFF4C4" box="[685,814,1504,1529]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="45" pageNumber="204" unit="mm" value="10.25" valueMax="11.0" valueMin="9.5">9.5–11 mm</quantity>
long,
<quantity id="D868D28061170D1AFCF4FA4DFC7DF4C4" box="[888,1004,1504,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="45" pageNumber="204" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
wide. Receptacle flat, densely covered with persistent white bristles, bristles spine-like,
<quantity id="D868D28061170D1AFD29F9A9FC90F720" box="[677,769,1540,1565]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="45" pageNumber="204" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long. Florets 15–20, corolla purple, ca.
<quantity id="D868D28061170D1AFB36F9A9FA9EF720" box="[1210,1295,1540,1565]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="45" pageNumber="204" unit="mm" value="9.5">9.5 mm</quantity>
long, tubular-funnelform, glabrous, tube ca.
<quantity id="D868D28061170D1AFE6EF985FDA9F77C" box="[482,568,1576,1601]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="45" pageNumber="204" unit="mm" value="3.5">3.5 mm</quantity>
long, limb ca.
<quantity id="D868D28061170D1AFD6DF985FCB3F77C" box="[737,802,1576,1601]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="45" pageNumber="204" unit="mm" value="6.0">6 mm</quantity>
long, lobes ca.
<quantity id="D868D28061170D1AFC5FF985FB85F77C" box="[979,1044,1576,1601]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="204" unit="mm" value="3.0">3 mm</quantity>
long. Anthers ca.
<quantity id="D868D28061170D1AFB6CF985FAB3F77C" box="[1248,1314,1576,1601]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="45" pageNumber="204" unit="mm" value="8.0">8 mm</quantity>
long, apical appendage acute, basal appendages obtuse, auricles with lacerate tails ca.
<quantity id="D868D28061170D1AFC58F9E1FBBDF758" box="[980,1068,1612,1637]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="45" pageNumber="204" unit="mm" value="1.8">1.8 mm</quantity>
long. Achenes cylindroid, ca.
<quantity id="D868D28061170D1AFA05F9E1FF25F7B4" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="45" pageNumber="204" unit="mm" value="2.5">2.5 mm</quantity>
long, ca.
<quantity id="D868D28061170D1AFEAEF9DDFEE9F7B4" box="[290,376,1648,1673]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="45" pageNumber="204" unit="mm" value="0.8">0.8 mm</quantity>
wide, glabrous, ribbed. Pappus light brown; outer bristles scabrid, ca.
<quantity id="D868D28061170D1AFB1DF9DDFB43F7B4" box="[1169,1234,1648,1673]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="204" unit="mm" value="2.0">2 mm</quantity>
long; inner bristles 14–16, plumose, ca.
<quantity id="D868D28061170D1AFEFCF939FE20F790" box="[368,433,1684,1709]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="45" pageNumber="204" unit="mm" value="9.0">9 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="578A2CEE61170D1AFF31F915FCE2F624" pageId="45" pageNumber="204" type="distribution">
<paragraph id="1F2F7F6561170D1AFF31F915FB5AF7C8" blockId="45.[136,1452,1144,2069]" pageId="45" pageNumber="204">
<emphasis id="2DE4A37761170D1AFF31F915FE72F7EC" bold="true" box="[189,483,1720,1745]" pageId="45" pageNumber="204">Distribution and habitat</emphasis>
:—
<taxonomicName id="D89004E661170D1AFD8AF915FC96F7EC" authorityName="Y. S. Chen" authorityYear="2015" box="[518,775,1720,1745]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="species" species="jiulongensis">
<emphasis id="2DE4A37761170D1AFD8AF915FC96F7EC" box="[518,775,1720,1745]" italics="true" pageId="45" pageNumber="204">Saussurea jiulongensis</emphasis>
</taxonomicName>
is currently known only from Jiulong County and Yanyuan County, southwestern
<collectingRegion id="DD54B18761170D1AFE0DF971FE4FF7C8" box="[385,478,1756,1781]" country="China" name="Sichuan" pageId="45" pageNumber="204">Sichuan</collectingRegion>
,
<collectingCountry id="67873FF561170D1AFE66F971FDA0F7C8" box="[490,561,1756,1781]" name="China" pageId="45" pageNumber="204">China</collectingCountry>
. It grows in forests or thickets at altitudes of
<quantity id="D868D28061170D1AFBA4F971FB57F7C8" box="[1064,1222,1756,1781]" metricMagnitude="3" metricUnit="m" metricValue="2.985" metricValueMax="3.32" metricValueMin="2.65" pageId="45" pageNumber="204" unit="m" value="2985.0" valueMax="3320.0" valueMin="2650.0">2650–3320 m</quantity>
.
</paragraph>
<paragraph id="1F2F7F6561170D1AFF31F8ADFCE2F624" blockId="45.[136,1452,1144,2069]" box="[189,883,1792,1817]" pageId="45" pageNumber="204">
<emphasis id="2DE4A37761170D1AFF31F8ADFEA8F624" bold="true" box="[189,313,1792,1817]" pageId="45" pageNumber="204">Phenology</emphasis>
:—Flowering and fruiting from August to October.
</paragraph>
</subSubSection>
<subSubSection id="578A2CEE61170D1AFF31F889FD5BF694" pageId="45" pageNumber="204" type="etymology">
<paragraph id="1F2F7F6561170D1AFF31F889FA38F600" blockId="45.[136,1452,1144,2069]" box="[189,1449,1826,1854]" pageId="45" pageNumber="204">
<emphasis id="2DE4A37761170D1AFF31F889FEAFF600" bold="true" box="[189,318,1828,1853]" pageId="45" pageNumber="204">Etymology</emphasis>
:—The specific epithet is derived from its
<typeStatus id="C02BC1C761170D1AFC96F889FCDBF600" box="[794,842,1828,1853]" pageId="45" pageNumber="204">type</typeStatus>
locality, Jiulong County. Chinese name:
<emphasis id="2DE4A37761170D1AFA9AF88FFA33F603" bold="true" box="[1302,1442,1826,1854]" pageId="45" pageNumber="204">AżṈLẄ</emphasis>
.
</paragraph>
<paragraph id="1F2F7F6561170D1AFF31F8E5FD5BF694" blockId="45.[136,1452,1144,2069]" pageId="45" pageNumber="204">
<materialsCitation id="AFF8753861170D1AFF31F8E5FCDDF6B8" collectingDate="2011-07-29" collectionCode="Y, S, C, PE" country="China" county="Yanyuan County" elevation="3320" location="Huolu Shan" municipality="Wodi" pageId="45" pageNumber="204" specimenCount="2" stateProvince="Sichuan">
<emphasis id="2DE4A37761170D1AFF31F8E5FD52F65C" bold="true" box="[189,707,1864,1889]" pageId="45" pageNumber="204">
<typeStatus id="C02BC1C761170D1AFF31F8E5FDA4F65C" box="[189,565,1864,1889]" isOtherTypeStatus="true" pageId="45" pageNumber="204">Additional specimens examined</typeStatus>
(
<typeStatus id="C02BC1C761170D1AFDCEF8E5FD2DF65C" box="[578,700,1864,1889]" pageId="45" pageNumber="204" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="67873FF561170D1AFD69F8E5FCD2F65C" box="[741,835,1864,1889]" name="China" pageId="45" pageNumber="204">CHINA</collectingCountry>
.
<collectingRegion id="DD54B18761170D1AFCC0F8E5FC3AF65C" box="[844,939,1864,1889]" country="China" name="Sichuan" pageId="45" pageNumber="204">Sichuan</collectingRegion>
:
<collectingCounty id="F64E07E961170D1AFC3FF8E5FBFEF65C" box="[947,1135,1864,1889]" pageId="45" pageNumber="204">Yanyuan County</collectingCounty>
,
<collectingMunicipality id="FF4BE51F61170D1AFBFBF8E5FB27F65C" box="[1143,1206,1864,1889]" pageId="45" pageNumber="204">Wodi</collectingMunicipality>
,
<location id="1A4F29BE61170D1AFB4CF8E5FAD6F65C" LSID="urn:lsid:plazi:treatment:9739CE7361170D19FF04FC69FF5AF074:1A4F29BE61170D1AFB4CF8E5FAD6F65C" box="[1216,1351,1864,1889]" country="China" county="Yanyuan County" municipality="Wodi" name="Huolu Shan" pageId="45" pageNumber="204" stateProvince="Sichuan">Huolu Shan</location>
, in
<taxonomicName id="D89004E661170D1AFAE0F8E5FA3DF65C" box="[1388,1452,1864,1889]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="genus">
<emphasis id="2DE4A37761170D1AFAE0F8E5FA3DF65C" box="[1388,1452,1864,1889]" italics="true" pageId="45" pageNumber="204">Pinus</emphasis>
</taxonomicName>
forests,
<quantity id="D868D28061170D1AFF53F8C1FEA4F6B8" box="[223,309,1900,1925]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="45" pageNumber="204" unit="m" value="3320.0">
<elevation id="94BD985661170D1AFF53F8C1FEA4F6B8" box="[223,309,1900,1925]" metricMagnitude="3" metricUnit="m" metricValue="3.32" pageId="45" pageNumber="204" unit="m" value="3320.0">3320 m</elevation>
</quantity>
,
<date id="6B2E59A561170D1AFECCF8C1FE41F6B8" box="[320,464,1900,1925]" pageId="45" pageNumber="204" value="2011-07-29">
<collectingDate id="7B6AA04D61170D1AFECCF8C1FE41F6B8" box="[320,464,1900,1925]" pageId="45" pageNumber="204" value="2011-07-29">29 July 2011</collectingDate>
</date>
,
<emphasis id="2DE4A37761170D1AFE56F8C1FC81F6B8" box="[474,784,1900,1925]" italics="true" pageId="45" pageNumber="204">
<collectionCode id="7981E7A061170D1AFE56F8C1FE7AF6B8" box="[474,491,1900,1925]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="45" pageNumber="204" type="Herbarium">Y</collectionCode>
.
<collectionCode id="7981E7A061170D1AFE78F8C1FD94F6B8" box="[500,517,1900,1925]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="45" pageNumber="204" type="Herbarium">S</collectionCode>
. Chen &amp;
<collectionCode id="7981E7A061170D1AFDE7F8C1FDEDF6B8" box="[619,636,1900,1925]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="45" pageNumber="204" type="Herbarium">Y</collectionCode>
.
<collectionCode id="7981E7A061170D1AFD09F8C1FD0AF6B8" box="[645,667,1900,1925]" country="Denmark" name="University of Copenhagen" pageId="45" pageNumber="204" type="Herbarium">C</collectionCode>
. Bi 11-073
</emphasis>
(
<collectionCode id="7981E7A061170D1AFC92F8C1FCD5F6B8" box="[798,836,1900,1925]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="45" pageNumber="204" type="Herbarium">PE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AFF8753861170D1AFCDAF8C1FD57F694" collectingDate="2011-07-28" collectionCode="Y, S, C, PE" country="China" county="Yanyuan County" elevation="2900" location="Huolu Shan" municipality="Wodi" pageId="45" pageNumber="204" specimenCount="1">
<collectingCounty id="F64E07E961170D1AFCDAF8C1FB83F6B8" box="[854,1042,1900,1925]" pageId="45" pageNumber="204">Yanyuan County</collectingCounty>
,
<collectingMunicipality id="FF4BE51F61170D1AFB97F8C1FBCBF6B8" box="[1051,1114,1900,1925]" pageId="45" pageNumber="204">Wodi</collectingMunicipality>
,
<location id="1A4F29BE61170D1AFBEFF8C1FB7BF6B8" LSID="urn:lsid:plazi:treatment:9739CE7361170D19FF04FC69FF5AF074:1A4F29BE61170D1AFBEFF8C1FB7BF6B8" box="[1123,1258,1900,1925]" country="China" county="Yanyuan County" municipality="Wodi" name="Huolu Shan" pageId="45" pageNumber="204">Huolu Shan</location>
, in thickets,
<quantity id="D868D28061170D1AFAFFF8C1FF31F694" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="45" pageNumber="204" unit="m" value="2900.0">
<elevation id="94BD985661170D1AFAFFF8C1FF31F694" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="45" pageNumber="204" unit="m" value="2900.0">2900 m</elevation>
</quantity>
,
<date id="6B2E59A561170D1AFF20F83DFEAEF694" box="[172,319,1936,1961]" pageId="45" pageNumber="204" value="2011-07-28">
<collectingDate id="7B6AA04D61170D1AFF20F83DFEAEF694" box="[172,319,1936,1961]" pageId="45" pageNumber="204" value="2011-07-28">28 July 2011</collectingDate>
</date>
,
<emphasis id="2DE4A37761170D1AFEC5F83DFD19F694" box="[329,648,1936,1961]" italics="true" pageId="45" pageNumber="204">
<collectionCode id="7981E7A061170D1AFEC5F83DFECBF694" box="[329,346,1936,1961]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="45" pageNumber="204" type="Herbarium">Y</collectionCode>
.
<collectionCode id="7981E7A061170D1AFEE8F83DFEE4F694" box="[356,373,1936,1961]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="45" pageNumber="204" type="Herbarium">S</collectionCode>
. Chen &amp;
<collectionCode id="7981E7A061170D1AFE53F83DFE61F694" box="[479,496,1936,1961]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="45" pageNumber="204" type="Herbarium">Y</collectionCode>
.
<collectionCode id="7981E7A061170D1AFE76F83DFD81F694" box="[506,528,1936,1961]" country="Denmark" name="University of Copenhagen" pageId="45" pageNumber="204" type="Herbarium">C</collectionCode>
. Bi 11-063
</emphasis>
(
<collectionCode id="7981E7A061170D1AFD14F83DFD2FF694" box="[664,702,1936,1961]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="45" pageNumber="204" type="Herbarium">PE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="578A2CEE61170D19FF31F819FF5AF074" lastPageId="46" lastPageNumber="205" pageId="45" pageNumber="204" type="discussion">
<paragraph id="1F2F7F6561170D19FF31F819FF5AF074" blockId="45.[136,1452,1144,2069]" lastBlockId="46.[136,1451,160,329]" lastPageId="46" lastPageNumber="205" pageId="45" pageNumber="204">
<emphasis id="2DE4A37761170D1AFF31F819FEAAF6F0" bold="true" box="[189,315,1972,1997]" pageId="45" pageNumber="204">Discussion</emphasis>
:—
<taxonomicName id="D89004E661170D1AFED3F819FDCDF6F0" authorityName="Y. S. Chen" authorityYear="2015" box="[351,604,1972,1997]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="species" species="jiulongensis">
<emphasis id="2DE4A37761170D1AFED3F819FDCDF6F0" box="[351,604,1972,1997]" italics="true" pageId="45" pageNumber="204">Saussurea jiulongensis</emphasis>
</taxonomicName>
belongs to
<emphasis id="2DE4A37761170D1AFD58F819FD7BF6F0" box="[724,746,1972,1997]" italics="true" pageId="45" pageNumber="204">S.</emphasis>
subgen.
<taxonomicName id="D89004E661170D1AFCCBF819FBF4F6F0" authority="sect. Saussurea" box="[839,1125,1972,1997]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="section" section="Saussurea">
<emphasis id="2DE4A37761170D1AFCCBF819FC28F6F0" box="[839,953,1972,1997]" italics="true" pageId="45" pageNumber="204">Saussurea</emphasis>
sect.
<emphasis id="2DE4A37761170D1AFC7FF819FBF4F6F0" box="[1011,1125,1972,1997]" italics="true" pageId="45" pageNumber="204">Saussurea</emphasis>
</taxonomicName>
because of its well-developed stems, roots not fibrously split, leaves evenly arranged along the stem, capitula numerous and in sparse corymbs, anther tails lacerate. In our molecular tree,
<taxonomicName id="D89004E661170D1AFDF1F851FCB2F928" authorityName="Y. S. Chen" authorityYear="2015" box="[637,803,2044,2069]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="species" species="jiulongensis">
<emphasis id="2DE4A37761170D1AFDF1F851FCB2F928" box="[637,803,2044,2069]" italics="true" pageId="45" pageNumber="204">S. jiulongensis</emphasis>
</taxonomicName>
is shown to be closely related to
<taxonomicName id="D89004E661170D1AFB23F851FA34F928" authorityName="Y. S. Chen" authorityYear="2015" box="[1199,1445,2044,2069]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="45" pageNumber="204" phylum="Tracheophyta" rank="species" species="pseudojiulongensis">
<emphasis id="2DE4A37761170D1AFB23F851FA34F928" box="[1199,1445,2044,2069]" italics="true" pageId="45" pageNumber="204">S. pseudojiulongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="D89004E661140D19FF04FF0DFD30F184" authority="Handel-Mazzetti (1936: 1149)" authorityName="Handel-Mazzetti (1936:" authorityPageNumber="1149" authorityYear="1149" box="[136,673,160,185]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="205" phylum="Tracheophyta" rank="species" species="pteridophylla">
<emphasis id="2DE4A37761140D19FF04FF0DFEAFF184" box="[136,318,160,185]" italics="true" pageId="46" pageNumber="205">S. pteridophylla</emphasis>
<bibRefCitation id="7B01029461140D19FECBFF0DFD30F184" author="Handel-Mazzetti, H." box="[327,673,160,185]" pageId="46" pageNumber="205" pagination="731 - 1450" refId="ref30968" refString="Handel-Mazzetti, H. (1936) Symbolae Sinicae, vol. 7. Julius Springer, Wien, pp. 731 - 1450." type="book chapter" year="1936">Handel-Mazzetti (1936: 1149)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D89004E661140D19FD50FF0DFB55F184" authority="Handel-Mazzetti (1936: 1147)" authorityName="Handel-Mazzetti" authorityPageNumber="1147" authorityYear="1936" box="[732,1220,160,185]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="205" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis id="2DE4A37761140D19FD50FF0DFCCEF184" box="[732,863,160,185]" italics="true" pageId="46" pageNumber="205">S. undulata</emphasis>
<bibRefCitation id="7B01029461140D19FCE5FF0DFB55F184" author="Handel-Mazzetti, H." box="[873,1220,160,185]" pageId="46" pageNumber="205" pagination="731 - 1450" refId="ref30968" refString="Handel-Mazzetti, H. (1936) Symbolae Sinicae, vol. 7. Julius Springer, Wien, pp. 731 - 1450." type="book chapter" year="1936">Handel-Mazzetti (1936: 1147)</bibRefCitation>
</taxonomicName>
, these species share characters in winged petioles and leaves base decurrent along stem to
<taxonomicName id="D89004E661140D19FC23FF69FBD0F1E0" box="[943,1089,196,221]" form="distinct" pageId="46" pageNumber="205" rank="form">form distinct</taxonomicName>
wings, capitula shape, size and arrangement. It is different from
<taxonomicName id="D89004E661140D19FE7DFF45FD32F03C" authorityName="Handel-Mazzetti (1936:" authorityYear="1149" box="[497,675,232,257]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="205" phylum="Tracheophyta" rank="species" species="pteridophylla">
<emphasis id="2DE4A37761140D19FE7DFF45FD32F03C" box="[497,675,232,257]" italics="true" pageId="46" pageNumber="205">S. pteridophylla</emphasis>
</taxonomicName>
by its leaf margin undulate-dentate to shallowly pinnatifid, mucronate (vs. pinnately lobed to pinnatisect, lateral lobes 6–10 pairs), petioles of lower leaves
<quantity id="D868D28061140D19FBA2FEA1FB1FF018" box="[1070,1166,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="46" pageNumber="205" unit="cm" value="8.0" valueMax="12.0" valueMin="4.0">4–12 cm</quantity>
long (vs. less than
<quantity id="D868D28061140D19FAEDFEA1FA3AF018" box="[1377,1451,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="205" unit="cm" value="2.5">2.5 cm</quantity>
long).
</paragraph>
</subSubSection>
</treatment>
</document>
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