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<document id="9F1A55742A10740348C22F07FC0B4CD4" ID-DOI="10.11646/zootaxa.4786.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3875378" ID-ZooBank="53FB7D7C-70DE-4FCD-BDE1-48C146DFB8DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591082920637" checkinUser="plazi" docAuthor="Park, Kyu-Tek, Koo, Jun-Mo, Agassiz, David J. L. &amp; Aarvik, Leif" docDate="2020" docId="A31187DAFF978F53FF29FEDAFAC14F34" docLanguage="en" docName="zootaxa.4786.2.1.pdf" docOrigin="Zootaxa 4786 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dragmatucha cochliana Park 2020, sp. nov." docType="treatment" docUuid="E2205E28-D3CE-460D-9DDD-17BC62ADE5A2" docUuidSource="ZooBank" docVersion="3" lastPageNumber="163" masterDocId="5F28FFA2FF9B8F5FFFBEFFABFFAA487B" masterDocTitle="A taxonomic review of the Afrotropical genus Dragmatucha Meyrick, 1908 (Lepidoptera, Gelechioidea, Lecithoceridae, Torodorinae), with descriptions of eleven new species" masterLastPageNumber="175" masterPageNumber="151" pageNumber="163" updateTime="1700276319097" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CB84004A7580851E090CD9B900538FCA">
<mods:title id="03C899C1731E66EC6412BF76E227A7E5">A taxonomic review of the Afrotropical genus Dragmatucha Meyrick, 1908 (Lepidoptera, Gelechioidea, Lecithoceridae, Torodorinae), with descriptions of eleven new species</mods:title>
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<mods:name id="B11323F64D9314563119EAB3214C4879" type="personal">
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<mods:namePart id="004165FF8242B5E2CD59381A96D9DAC0">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="90D0A11CB8ED04FA454783C2B09B519A">Bioresource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
</mods:name>
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<mods:namePart id="9BFA41EB8F35B0E09F6834E090BD7C1A">Koo, Jun-Mo</mods:namePart>
<mods:affiliation id="948FA795DA94CF14B8199312A54EED6C">Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea.</mods:affiliation>
</mods:name>
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<mods:namePart id="09C0C0CF7B3F372650BA87FA53C956FE">Agassiz, David J. L.</mods:namePart>
<mods:affiliation id="9F55CFBE03895D81B46FB9E6FE8FE96D">Dept. of Life Sciences, Insects Division, Natural History Museum, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="51EE37E3B5BF009752B91E5A08E33A1C">Aarvik, Leif</mods:namePart>
<mods:affiliation id="7131E6DFCEA160D2D11A25BC24018847">Natural History Museum, University of Oslo, P. O. Box 1172 Blindern, NO- 0318 Oslo, Norway.</mods:affiliation>
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<mods:title id="FB58D2E77540C6B184F8D1F53EB95A4D">Zootaxa</mods:title>
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<mods:part id="DB48953D6CE305406287FE8CFB34CD03">
<mods:date id="6832C7F312428134D2FAA7ED8DBE4A80">2020</mods:date>
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<mods:number id="16209829B02C52F7EBF29A7B5493765B">2020-06-02</mods:number>
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<mods:number id="35DC52895CB9A3EAF6F93E5AD7687C09">4786</mods:number>
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<mods:number id="7C478319BEA5C1215DF8BB251400CB22">2</mods:number>
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<mods:identifier id="963A320892DF83B108320AAE31D30742" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="63A26547FF978F53FF29FEDAFD2649F0" box="[151,652,369,396]" pageId="12" pageNumber="163" type="nomenclature">
<paragraph id="2B0736CCFF978F53FF29FEDAFD2649F0" blockId="12.[151,652,369,431]" box="[151,652,369,396]" pageId="12" pageNumber="163">
<heading id="704F81A0FF978F53FF29FEDAFD2649F0" bold="true" box="[151,652,369,396]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<emphasis id="19CCEADEFF978F53FF29FEDAFD2649F0" bold="true" box="[151,652,369,396]" pageId="12" pageNumber="163">
3.
<taxonomicName id="ECB84D4FFF978F53FF0BFEDAFD8849F0" ID-CoL="9FXFT" authority="Park" authorityName="Park" authorityYear="2020" box="[181,546,369,396]" class="Insecta" family="Lecithoceridae" genus="Dragmatucha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="cochliana" status="sp. nov.">
<emphasis id="19CCEADEFF978F53FF0BFEDAFE7149F7" bold="true" box="[181,475,369,396]" italics="true" pageId="12" pageNumber="163">Dragmatucha cochliana</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="02FF57A5FF978F53FD91FEDAFD2649F0" box="[559,652,369,395]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF29FE3EFE8949D4" box="[151,291,405,431]" pageId="12" pageNumber="163" type="description">
<paragraph id="2B0736CCFF978F53FF29FE3EFE8949D4" blockId="12.[151,652,369,431]" box="[151,291,405,431]" pageId="12" pageNumber="163">
(
<figureCitation id="B3832A49FF978F53FF21FE3EFF4E49D4" box="[159,228,405,431]" captionStart="FIGURE 8" captionStartId="13.[151,250,1328,1353]" captionTargetBox="[216,1339,229,1269]" captionTargetId="figure@13.[185,1403,181,1303]" captionTargetPageId="13" captionText="FIGURE 8. Dragmatucha cochliana Park, sp. nov.: A, adult, holotype; B, label; C, male genitalia, holotype, gen. slide no. CIS- 7192; D, aedeagus, with close-up apical part; E, abdomen with spinous zones." figureDoi="http://doi.org/10.5281/zenodo.3875396" httpUri="https://zenodo.org/record/3875396/files/figure.png" pageId="12" pageNumber="163">Figs 8</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF29FE76FAEA4A60" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="2B0736CCFF978F53FF29FE76FAEA4A60" blockId="12.[151,1437,476,1871]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF29FE76FEED498C" bold="true" box="[151,327,476,503]" pageId="12" pageNumber="163">Type material.</emphasis>
<materialsCitation id="9BD03C91FF978F53FEEEFE76FAEA4A60" collectingDate="2014-11-19" collectingDateMax="2014-11-24" collectingDateMin="2014-11-19" collectionCode="NHMO" collectorName="Leif Aarvik &amp; Knud Larsen &amp; Barcode" country="Uganda" elevation="1500" location="Kasese Distr." pageId="12" pageNumber="163" specimenCount="1" stateProvince="Kasese" typeStatus="holotype">
<typeStatus id="F403886EFF978F53FEEEFE76FE17498C" box="[336,445,477,503]" pageId="12" pageNumber="163" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="19CCEADEFF978F53FDB1FE76FDC6498C" bold="true" box="[527,620,477,503]" pageId="12" pageNumber="163">
<collectingCountry id="53AF765CFF978F53FDB1FE76FDC6498C" box="[527,620,477,503]" name="Uganda" pageId="12" pageNumber="163">Uganda</collectingCountry>
</emphasis>
,
<collectingRegion id="E97CF82EFF978F53FDC3FE76FCBC498C" box="[637,790,477,503]" country="Uganda" name="Kasese" pageId="12" pageNumber="163">Kasese Distr.</collectingRegion>
,
<collectingRegion id="E97CF82EFF978F53FC9CFE76FCC7498C" box="[802,877,477,503]" country="Uganda" name="Kibale" pageId="12" pageNumber="163">Kibale</collectingRegion>
Nat. Park,
<date id="5F06100CFF978F53FC4CFE76FB3D498C" box="[1010,1175,477,503]" pageId="12" pageNumber="163" value="2014-11-19" valueMax="2014-11-24" valueMin="2014-11-19">
<collectingDate id="4F42E9E4FF978F53FC4CFE76FB3D498C" box="[1010,1175,477,503]" pageId="12" pageNumber="163" value="2014-11-19" valueMax="2014-11-24" valueMin="2014-11-19">1924 xi 2014</collectingDate>
</date>
[UTM:] 36N TF 0582 6208,
<quantity id="EC409B29FF978F53FF63FDABFE914A60" box="[221,315,512,539]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="1500.0">
<elevation id="A095D1FFFF978F53FF63FDABFE914A60" box="[221,315,512,539]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="1500.0">1,500 m</elevation>
</quantity>
,
<collectorName id="864D531AFF978F53FEF9FDAAFE634A60" box="[327,457,513,539]" pageId="12" pageNumber="163">Leif Aarvik</collectorName>
&amp;
<collectorName id="864D531AFF978F53FE53FDAAFD284A60" box="[493,642,513,539]" pageId="12" pageNumber="163">Knud Larsen</collectorName>
; gen. slide no. CIS-7192; COI
<collectorName id="864D531AFF978F53FC62FDAAFB934A60" box="[988,1081,513,539]" pageId="12" pageNumber="163">Barcode</collectorName>
CBNU083, in
<collectionCode id="4DA9AE09FF978F53FB5DFDAAFA964A60" box="[1251,1340,513,539]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="12" pageNumber="163" type="Museum">NHMO</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF79FD8FFE484A88" pageId="12" pageNumber="163" type="diagnosis">
<paragraph id="2B0736CCFF978F53FF79FD8FFE484A88" blockId="12.[151,1437,476,1871]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79FD8FFEE84A44" bold="true" box="[199,322,548,575]" pageId="12" pageNumber="163">Diagnosis.</emphasis>
This species is superficially similar to the preceding species,
<taxonomicName id="ECB84D4FFF978F53FC52FD8DFBC64A44" authority="Park &amp; Koo &amp; Agassiz &amp; Aarvik, 2020" authorityName="Park &amp; Koo &amp; Agassiz &amp; Aarvik" authorityYear="2020" box="[1004,1132,550,575]" class="Insecta" family="Lecithoceridae" genus="Dragmatucha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="goniotes" status="sp. nov.">
<emphasis id="19CCEADEFF978F53FC52FD8DFBC64A44" box="[1004,1132,550,575]" italics="true" pageId="12" pageNumber="163">D. goniotes</emphasis>
</taxonomicName>
<emphasis id="19CCEADEFF978F53FBCCFD8EFB614A44" bold="true" box="[1138,1227,549,575]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="02FF57A5FF978F53FBCCFD8EFB614A44" box="[1138,1227,549,575]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with similar wing pattern of both wings, but it can be distinguished by the larger size, the forewing more narrowly elongated with more oblique termen and the yellowish white postmedian band in the hind wing less sharply angled between veins R
<subScript id="B73C3489FF978F53FF14FD09FF6E4ACB" attach="left" box="[170,196,674,688]" fontSize="6" pageId="12" pageNumber="163">3+4</subScript>
and M
<subScript id="B73C3489FF978F53FEA9FD09FEB54ACB" attach="left" box="[279,287,674,688]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
. In addition, it can be distinguished from the latter by the male genitalia with cucullus more robust, half-moon-shaped, with more rounded outer margin; the juxta deeply concave on caudal margin, with triangularly produced latero-caudal lobes.
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF79FD57FE384E98" pageId="12" pageNumber="163" type="description">
<paragraph id="2B0736CCFF978F53FF79FD57FCC84D00" blockId="12.[151,1437,476,1871]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79FD57FEF34B6C" bold="true" box="[199,345,764,791]" pageId="12" pageNumber="163">Description.</emphasis>
Male (
<figureCitation id="B3832A49FF978F53FE17FD56FDAE4B6C" box="[425,516,765,791]" captionStart="FIGURE 8" captionStartId="13.[151,250,1328,1353]" captionTargetBox="[216,1339,229,1269]" captionTargetId="figure@13.[185,1403,181,1303]" captionTargetPageId="13" captionText="FIGURE 8. Dragmatucha cochliana Park, sp. nov.: A, adult, holotype; B, label; C, male genitalia, holotype, gen. slide no. CIS- 7192; D, aedeagus, with close-up apical part; E, abdomen with spinous zones." figureDoi="http://doi.org/10.5281/zenodo.3875396" httpUri="https://zenodo.org/record/3875396/files/figure.png" pageId="12" pageNumber="163">Fig. 8A</figureCitation>
). Wingspan
<quantity id="EC409B29FF978F53FD31FD57FD5D4B6C" box="[655,759,764,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="12" pageNumber="163" unit="mm" value="18.5">18.5 mm</quantity>
.
<emphasis id="19CCEADEFF978F53FCBCFD56FC954B6C" box="[770,831,765,791]" italics="true" pageId="12" pageNumber="163">Head</emphasis>
: Yellowish white, with yellowish brown scales posteriorly. Antenna slightly shorter than forewing; basal segment elongated, dark brown dorsally, yellowish white ventrally; flagellum yellowish white throughout, sharply acute apically. Second palpomere of labial palpus thickened, long, more than twice width of compound eye, arched, pale yellowish brown, speckled with dark-brown scales on outer surface; 3
<superScript id="DCCD9B84FF978F53FEFAFC27FEF84BE1" attach="left" box="[324,338,908,922]" fontSize="6" pageId="12" pageNumber="163">rd</superScript>
palpomere slender, strongly upturned, as long as 2
<superScript id="DCCD9B84FF978F53FC39FC27FC3D4BE1" attach="left" box="[903,919,908,922]" fontSize="6" pageId="12" pageNumber="163">nd</superScript>
palpomere, yellowish brown on outer surface.
<emphasis id="19CCEADEFF978F53FF29FC1AFF424BB0" box="[151,232,945,971]" italics="true" pageId="12" pageNumber="163">Thorax</emphasis>
: Tegula yellowish white, covered with yellowish brown scales anteriorly; thorax yellowish white with yellowish brown scales along anterior margin. Hind tibia yellowish white, with yellowish brown, rough scales medially. Forewing with dark brown sub-basal area; antemedian band broad, band-like, yellowish brown, slightly broadened toward costa and inner margin; area between antemedian band and postmedian band broad, about twice width of basal area; postmedian band linear, strongly angled inwardly on vein CuA
<subScript id="B73C3489FF978F53FBA8FBF9FBB44C1B" attach="left" box="[1046,1054,1106,1120]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
; apex produced; termen oblique; fringe yellowish white, with yellowish brown median band; venation with R
<subScript id="B73C3489FF978F53FC47FBDDFBAB4CFF" attach="left" box="[1017,1025,1142,1156]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
arising before middle of discal cell; distance between origins of R
<subScript id="B73C3489FF978F53FE5BFB31FE474CD3" attach="left" box="[485,493,1178,1192]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
and R
<subScript id="B73C3489FF978F53FD89FB31FD954CD3" attach="left" box="[567,575,1178,1192]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
about more than twice the length of R
<subScript id="B73C3489FF978F53FC4EFB31FC524CD3" attach="left" box="[1008,1016,1178,1192]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
and R
<subScript id="B73C3489FF978F53FBFFFB31FBE34CD3" attach="left" box="[1089,1097,1178,1192]" fontSize="6" pageId="12" pageNumber="163">3</subScript>
; R
<subScript id="B73C3489FF978F53FBD5FB31FBD94CD3" attach="left" box="[1131,1139,1178,1192]" fontSize="6" pageId="12" pageNumber="163">3</subScript>
stalked with R
<subScript id="B73C3489FF978F53FAA4FB31FA9E4CD3" attach="left" box="[1306,1332,1178,1192]" fontSize="6" pageId="12" pageNumber="163">4+5</subScript>
for more than basal 1/4; R
<subScript id="B73C3489FF978F53FEEEFB15FEF24CB7" attach="left" box="[336,344,1214,1228]" fontSize="6" pageId="12" pageNumber="163">4</subScript>
and R
<subScript id="B73C3489FF978F53FE21FB15FE0D4CB7" attach="left" box="[415,423,1214,1228]" fontSize="6" pageId="12" pageNumber="163">5</subScript>
stalked for about basal 3/5, R
<subScript id="B73C3489FF978F53FD51FB15FD5D4CB7" attach="left" box="[751,759,1214,1228]" fontSize="6" pageId="12" pageNumber="163">5</subScript>
reaching costa; M
<subScript id="B73C3489FF978F53FC7DFB15FC614CB7" attach="left" box="[963,971,1214,1228]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
remote from R
<subScript id="B73C3489FF978F53FBCFFB15FB214CB7" attach="left" box="[1137,1163,1214,1228]" fontSize="6" pageId="12" pageNumber="163">3+4</subScript>
at base; M
<subScript id="B73C3489FF978F53FABCFB15FAA04CB7" attach="left" box="[1282,1290,1214,1228]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
approximate to CuA
<subScript id="B73C3489FF978F53FF57FB49FF5B4C8B" attach="left" box="[233,241,1250,1264]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
; M
<subScript id="B73C3489FF978F53FEA4FB49FE884C8B" attach="left" box="[282,290,1250,1264]" fontSize="6" pageId="12" pageNumber="163">3</subScript>
merged to CuA
<subScript id="B73C3489FF978F53FE68FB49FE744C8B" attach="left" box="[470,478,1250,1264]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
; CuA
<subScript id="B73C3489FF978F53FD9DFB49FD814C8B" attach="left" box="[547,555,1250,1264]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
short-stalked with CuA
<subScript id="B73C3489FF978F53FC8BFB49FC974C8B" attach="left" box="[821,829,1250,1264]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
; 1A+2A not forked; cell closed. Hind wing yellowish brown, much paler in basal 1/3; postmedian band similar to that of forewing, slightly angled outwardly on M
<subScript id="B73C3489FF978F53FAF6FAADFAFA4D6F" attach="left" box="[1352,1360,1286,1300]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
; venation with M
<subScript id="B73C3489FF978F53FEA2FA81FE8E4D43" attach="left" box="[284,292,1322,1336]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
well-developed, arising from near lower corner of cell; M
<subScript id="B73C3489FF978F53FC0DFA81FC114D43" attach="left" box="[947,955,1322,1336]" fontSize="6" pageId="12" pageNumber="163">3</subScript>
merged to CuA
<subScript id="B73C3489FF978F53FBD1FA81FBDD4D43" attach="left" box="[1135,1143,1322,1336]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
; CuA
<subScript id="B73C3489FF978F53FB05FA81FB694D43" attach="left" box="[1211,1219,1322,1336]" fontSize="6" pageId="12" pageNumber="163">1</subScript>
connate with CuA
<subScript id="B73C3489FF978F53FA2AFA81FA364D43" attach="left" box="[1428,1436,1322,1336]" fontSize="6" pageId="12" pageNumber="163">2</subScript>
basally; CuP well-developed, arising from near middle of lower margin of discal cell.
<emphasis id="19CCEADEFF978F53FBE3FA96FB6D4D2C" box="[1117,1223,1341,1367]" italics="true" pageId="12" pageNumber="163">Abdomen</emphasis>
(
<figureCitation id="B3832A49FF978F53FB68FA96FA844D2C" box="[1238,1326,1341,1367]" captionStart="FIGURE 8" captionStartId="13.[151,250,1328,1353]" captionTargetBox="[216,1339,229,1269]" captionTargetId="figure@13.[185,1403,181,1303]" captionTargetPageId="13" captionText="FIGURE 8. Dragmatucha cochliana Park, sp. nov.: A, adult, holotype; B, label; C, male genitalia, holotype, gen. slide no. CIS- 7192; D, aedeagus, with close-up apical part; E, abdomen with spinous zones." figureDoi="http://doi.org/10.5281/zenodo.3875396" httpUri="https://zenodo.org/record/3875396/files/figure.png" pageId="12" pageNumber="163">Fig. 8E</figureCitation>
): Spinous zones on dorsal surface broadly occupied; segment VIII simple.
</paragraph>
<paragraph id="2B0736CCFF978F53FF79FA2EFC894EC4" blockId="12.[151,1437,476,1871]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79FA2EFEC54DE4" box="[199,367,1413,1439]" italics="true" pageId="12" pageNumber="163">Male genitalia</emphasis>
(
<figureCitation id="B3832A49FF978F53FE3EFA2EFDAC4DE4" box="[384,518,1413,1439]" captionStart="FIGURE 8" captionStartId="13.[151,250,1328,1353]" captionTargetBox="[216,1339,229,1269]" captionTargetId="figure@13.[185,1403,181,1303]" captionTargetPageId="13" captionText="FIGURE 8. Dragmatucha cochliana Park, sp. nov.: A, adult, holotype; B, label; C, male genitalia, holotype, gen. slide no. CIS- 7192; D, aedeagus, with close-up apical part; E, abdomen with spinous zones." figureDoi="http://doi.org/10.5281/zenodo.3875396" httpUri="https://zenodo.org/record/3875396/files/figure.png" pageId="12" pageNumber="163">Figs 8C, D</figureCitation>
): Uncus slender, heavily sclerotized, nearly parallel sided, with acute apex, exceeding apex of basal plate of gnathos. Basal plate of gnathos narrowed in apical part, with rounded apex; median process strongly bent beyond middle, with sharply pointed apex. Tegumen sclerotized, deeply concave on anterior margin. Valva broad basally, narrowed toward cucullus; costa convex basally, then slightly oblique; ventral margin arched; sacculus weakly sclerotized, broad at base, then gradually narrowed, terminated at lower corner of cucullus; cucullus short, with rounded outer margin, densely setose. Juxta narrowed distally, concave on caudal margin with triangular caudal lobes laterally; median ridge weakly developed.Aedeagus stout, as long as valva, bent at basal 1/3; dorsal surface terminated with ladle-like plate, upturned; cornuti consist of a sole-like plate and much smaller one, and a large patch bearing numerous spinules in distal part.
</paragraph>
<paragraph id="2B0736CCFF978F53FF79F962FE384E98" blockId="12.[151,1437,476,1871]" box="[199,402,1737,1763]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79F962FEB14E98" box="[199,283,1737,1763]" italics="true" pageId="12" pageNumber="163">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF79F947FD314F73" box="[199,667,1772,1800]" pageId="12" pageNumber="163" type="distribution">
<paragraph id="2B0736CCFF978F53FF79F947FD314F73" blockId="12.[151,1437,476,1871]" box="[199,667,1772,1800]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79F947FECB4F7C" bold="true" box="[199,353,1772,1799]" pageId="12" pageNumber="163">Distribution.</emphasis>
<collectingCountry id="53AF765CFF978F53FED9F946FE144F7C" box="[359,446,1773,1799]" name="Uganda" pageId="12" pageNumber="163">Uganda</collectingCountry>
(
<collectingRegion id="E97CF82EFF978F53FE73F946FDB34F7C" box="[461,537,1773,1799]" country="Uganda" name="Kibale" pageId="12" pageNumber="163">Kibale</collectingRegion>
Nat. Park).
</paragraph>
</subSubSection>
<subSubSection id="63A26547FF978F53FF79F8BAFAC14F34" pageId="12" pageNumber="163" type="etymology">
<paragraph id="2B0736CCFF978F53FF79F8BAFAC14F34" blockId="12.[151,1437,476,1871]" pageId="12" pageNumber="163">
<emphasis id="19CCEADEFF978F53FF79F8BAFEE44F50" bold="true" box="[199,334,1809,1835]" pageId="12" pageNumber="163">Etymology.</emphasis>
This species is derived from the Latin,
<emphasis id="19CCEADEFF978F53FCB5F8BAFCC74F50" box="[779,877,1809,1835]" italics="true" pageId="12" pageNumber="163">cochlear</emphasis>
(= ladle, spoon), referring to the ladle-like dorsoapical plate of the aedeagus in the male genitalia, with a Latin suffix, -
<emphasis id="19CCEADEFF978F53FC12F89DFC7C4F34" box="[940,982,1846,1871]" italics="true" pageId="12" pageNumber="163">ana</emphasis>
. It is a noun in the nominative case.
</paragraph>
</subSubSection>
</treatment>
</document>