141 lines
17 KiB
XML
141 lines
17 KiB
XML
<document id="D4B55EA774074E4BDF0CB1BDF4243BDE" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A0041FFC3FF182B6611033394" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus perpunctatus Perkins 2021, new species" docType="treatment" docVersion="6" lastPageNumber="471" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="470" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EF266766F91A5A84170BE4573BF9730E">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="884A72C6C7F62B7C6B577E89B5BA8D94">Perkins, Philip D.</mods:namePart>
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<mods:title id="9F033CB1B1B1C1906EABA701F50C7332">Zootaxa</mods:title>
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<mods:part id="8D4A0421CFA6B68EF61256FA22475547">
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<mods:date id="EEB7D34BB82C41CC477E1F027611D2A8">2021</mods:date>
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<mods:number id="9577F53CE3D428570F271D173430AA85">2021-05-21</mods:number>
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<mods:number id="B69195F9D78565DFB86886220CD2616B">4974</mods:number>
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<mods:identifier id="F36B2238AF009A1BF025D74836794F6B" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
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<treatment id="4772526A0041FFC3FF182B6611033394" ID-DOI="http://doi.org/10.5281/zenodo.4777895" ID-GBIF-Taxon="182642575" ID-Zenodo-Dep="4777895" LSID="urn:lsid:plazi:treatment:4772526A0041FFC3FF182B6611033394" httpUri="http://treatment.plazi.org/id/4772526A0041FFC3FF182B6611033394" lastPageId="12" lastPageNumber="471" pageId="11" pageNumber="470">
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<subSubSection id="87C1B0F70041FFC4FF182B66177C36A8" box="[151,609,1662,1689]" pageId="11" pageNumber="470" type="nomenclature">
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<paragraph id="CF64E37C0041FFC4FF182B66177C36A8" blockId="11.[151,609,1662,1725]" box="[151,609,1662,1689]" pageId="11" pageNumber="470">
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<heading id="942C54100041FFC4FF182B66177C36A8" bold="true" box="[151,609,1662,1689]" fontSize="11" level="1" pageId="11" pageNumber="470" reason="1">
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<emphasis id="FDAF3F6E0041FFC4FF182B66177C36A8" bold="true" box="[151,609,1662,1689]" pageId="11" pageNumber="470">
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<taxonomicName id="08DB98FF0041FFC4FF182B6614DF36A9" ID-CoL="8Y4QZ" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,450,1662,1689]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="470" phylum="Arthropoda" rank="species" species="perpunctatus" status="sp. nov.">
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<emphasis id="FDAF3F6E0041FFC4FF182B6614DF36A9" bold="true" box="[151,450,1662,1689]" italics="true" pageId="11" pageNumber="470">Hydrochus perpunctatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="E69C82150041FFC4FE5E2B67177C36A8" box="[465,609,1662,1689]" pageId="11" pageNumber="470" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF182BBB142B368D" box="[151,310,1699,1725]" pageId="11" pageNumber="470" type="description">
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<paragraph id="CF64E37C0041FFC4FF182BBB142B368D" blockId="11.[151,609,1662,1725]" box="[151,310,1699,1725]" pageId="11" pageNumber="470">
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<figureCitation id="57E0FFF90041FFC4FF182BBB15FE368D" box="[151,227,1699,1725]" captionStart="FIGURE 1" captionStartId="4.[151,250,1863,1887]" captionTargetBox="[186,1395,194,1808]" captionTargetId="figure-22@4.[151,1435,179,1837]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777843" httpUri="https://zenodo.org/record/4777843/files/figure.png" pageId="11" pageNumber="470">Figs. 1</figureCitation>
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,
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<figureCitation id="57E0FFF90041FFC4FF7E2BBB1413368D" box="[241,270,1699,1725]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">11</figureCitation>
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,
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<figureCitation id="57E0FFF90041FFC4FE952BBB142B368D" box="[282,310,1699,1725]" captionStart="FIGURES 28–30" captionStartId="35.[151,264,1944,1968]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-15@35.[158,1429,181,1920]" captionTargetPageId="35" captionText="FIGURES 28–30. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777904" httpUri="https://zenodo.org/record/4777904/files/figure.png" pageId="11" pageNumber="470">28</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="87C1B0F70041FFC4FF182BF311BE3735" box="[151,1187,1770,1798]" pageId="11" pageNumber="470" type="materials_examined">
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<paragraph id="CF64E37C0041FFC4FF182BF311BE3735" blockId="11.[151,1437,1770,2014]" box="[151,1187,1770,1798]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF182BF314583734" bold="true" box="[151,325,1770,1797]" pageId="11" pageNumber="470">Type material.</emphasis>
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<materialsCitation id="7FB3E9210041FFC4FEC32BF311BE3735" ID-GBIF-Occurrence="3128194402" box="[332,1187,1770,1798]" collectingDate="1984-05-24" collectionCode="UNAM" collectorName="J. Pena" country="Mexico" location="Jalapa" pageId="11" pageNumber="470" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
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<typeStatus id="10605DDE0041FFC4FEC32BF314A93735" box="[332,436,1771,1797]" pageId="11" pageNumber="470" type="holotype">Holotype</typeStatus>
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(male): “
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<emphasis id="FDAF3F6E0041FFC4FD902BF2176B3734" bold="true" box="[543,630,1770,1796]" pageId="11" pageNumber="470">
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<collectingCountry id="B7CCA3EC0041FFC4FD902BF2176B3734" box="[543,630,1770,1796]" name="Mexico" pageId="11" pageNumber="470">Mexico</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="0D1F2D9E0041FFC4FD0A2BF317F13735" box="[645,748,1771,1797]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="11" pageNumber="470">Veracruz</collectingRegion>
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,
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<location id="CA04B5A70041FFC4FD792BF316223735" LSID="urn:lsid:plazi:treatment:4772526A0041FFC3FF182B6611033394:CA04B5A70041FFC4FD792BF316223735" box="[758,831,1771,1797]" country="Mexico" name="Jalapa" pageId="11" pageNumber="470" stateProvince="Veracruz de Ignacio de la Llave">Jalapa</location>
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,
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<date id="BB65C5BC0041FFC4FCC52BF316D83735" box="[842,965,1771,1797]" pageId="11" pageNumber="470" value="1984-05-24">
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<collectingDate id="AB213C540041FFC4FCC52BF316D83735" box="[842,965,1771,1797]" pageId="11" pageNumber="470" value="1984-05-24">24-V-1984</collectingDate>
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</date>
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,
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<collectorName id="622E86AA0041FFC4FC5F2BF3113F3735" box="[976,1058,1771,1797]" pageId="11" pageNumber="470">J. Pena</collectorName>
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” (
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<collectionCode id="A9CA7BB90041FFC4FBB42BF3118A3735" box="[1083,1175,1771,1797]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="11" pageNumber="470">UNAM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF482A1614D83789" pageId="11" pageNumber="470" type="diagnosis">
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<paragraph id="CF64E37C0041FFC4FF482A1614D83789" blockId="11.[151,1437,1770,2014]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF482A1614D03718" bold="true" box="[199,461,1806,1832]" pageId="11" pageNumber="470">Differential diagnosis.</emphasis>
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Among Central American species, recognized by the combination of moderate size (ca.
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<quantity id="08234E990041FFC4FF182A2B141D377D" box="[151,256,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="470" unit="mm" value="3.1">3.10 mm</quantity>
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), the densely coarsely punctate pronotum, the large and closely spaced strial punctures of the elytra, the three callosities of each elytron, and the male genitalia (
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<figureCitation id="57E0FFF90041FFC4FC9A2A4F16753741" box="[789,872,1879,1905]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">Fig. 11</figureCitation>
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; described below). The male genitalia can be differentiated from others by the aedeagus greatest width being near the base, and tapering toward the apex, plus the shape of the paramere tips.
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC3FF482ADA11EC33B1" lastPageId="12" lastPageNumber="471" pageId="11" pageNumber="470" type="description">
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<paragraph id="CF64E37C0041FFC3FF482ADA10713154" blockId="11.[151,1437,1770,2014]" lastBlockId="12.[151,1437,150,933]" lastPageId="12" lastPageNumber="471" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF482ADA144C37EC" bold="true" box="[199,337,1986,2012]" pageId="11" pageNumber="470">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="57E0FFF90041FFC4FDD52ADB17B737ED" box="[602,682,1987,2013]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">Fig. 11</figureCitation>
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). Size:
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<typeStatus id="10605DDE0041FFC4FD712ADB167D37ED" box="[766,864,1987,2013]" pageId="11" pageNumber="470" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.10/1.22; head width 0.76; pronotum l/w 0.67/0.76; PA 0.73; PB 0.61; elytra 2.04/1.22. Body size moderate (ca.
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<quantity id="08234E990046FFC3FF182DA215E230E5" box="[151,255,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="471" unit="mm" value="3.1">3.10 mm</quantity>
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). Dorsum grey to greenish grey, with moderately strong iridescence, elytra with large black spots. Punctation on elytra ca. 3–5x that of pronotum. Elytra interstriae ca. 1–1.5× strial puncture diameter. Interstria 9
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<superScript id="38AE4E340046FFC3FAC22DC6104730DC" attach="left" box="[1357,1370,222,236]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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more convex than others, overhanging 10
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<superScript id="38AE4E340046FFC3FDAA2C1A172F3120" attach="left" box="[549,562,258,272]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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interstria. Elytra with three calluses: usual area of callus on 5
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<superScript id="38AE4E340046FFC3FB522C1A11F73120" attach="left" box="[1245,1258,258,272]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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strongly raised, elongate, ca. 7 punctures in row on each side; additional callus of about same size on 5
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<superScript id="38AE4E340046FFC3FB0E2C3E11933104" attach="none" box="[1153,1166,294,308]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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, located slightly before midlength; third callus, smaller than others, on 7
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<superScript id="38AE4E340046FFC3FD382C5217D93168" attach="left" box="[695,708,330,344]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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interstria, ca. opposite gap between calluses on 5
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<superScript id="38AE4E340046FFC3FB622C5211E73168" attach="left" box="[1261,1274,330,344]" fontSize="6" pageId="12" pageNumber="471">th</superScript>
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interstria.
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</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482C77146F31E1" blockId="12.[151,1437,150,933]" pageId="12" pageNumber="471">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate, very minutely denticulate; anterior margin bisinuate; pronotum densely coarsely punctate; depressions large and deep; reliefs very narrow.</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482CC2102C3228" blockId="12.[151,1437,150,933]" pageId="12" pageNumber="471">Elytral punctures large and deep, spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with very slight angulation.</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482F3A117F3250" blockId="12.[151,1437,150,933]" pageId="12" pageNumber="471">Ventral characters: Mentum coarsely moderately densely punctate, with large median fovea. Submentum with two foveae. Legs testaceous, femora and tibiofemoral joints slightly darker than tibiae.</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482F7214543325" blockId="12.[151,1437,150,933]" pageId="12" pageNumber="471">
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Male genitalia (
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<figureCitation id="57E0FFF90046FFC3FEF32F7214D032B4" box="[380,461,618,644]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="12" pageNumber="471">Fig. 11</figureCitation>
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) general characters: paramere tips slender and pointed, narrower in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, plsa very slight; parameres distinctly sinuate in lateral view; aedeagus in dorsal/ventral views much wider in proximal ½ than distal ½, agw very near base; in lateral view narrower, without emargination where adbl and shaft meet; basal piece shorter than parameres.
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</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482E06117B336C" blockId="12.[151,1437,150,933]" pageId="12" pageNumber="471">Dorsal surface: pdmm separated slightly at adtl, then overlapping one another in ca. middle ½, then separating before adbl; adbl large; adtl at about distal 1/6; paramere tips without sclerotized ridge.</paragraph>
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<paragraph id="CF64E37C0046FFC3FF482E7E11EC33B1" blockId="12.[151,1437,150,933]" box="[199,1265,870,899]" pageId="12" pageNumber="471">Ventral surface: pvmm widely separated in proximal ½, but overlapping alm; avtl not apparent.</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70046FFC3FF482E9211033394" box="[199,1054,906,933]" pageId="12" pageNumber="471" type="etymology">
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<paragraph id="CF64E37C0046FFC3FF482E9211033394" blockId="12.[151,1437,150,933]" box="[199,1054,906,933]" pageId="12" pageNumber="471">
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<emphasis id="FDAF3F6E0046FFC3FF482E9214533394" bold="true" box="[199,334,906,932]" pageId="12" pageNumber="471">Etymology.</emphasis>
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Named in reference to the densely and deeply punctate dorsum.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |